The Hornwort and Liverwort Atlas of Romania

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Lophocolea hookeriana Nees = Lophocolea bidentata (L.) Dumort. var. rivularis (Raddi) Schiffn.Lophocolea incisa Lindb. = Lophocolea heterophylla (Schrad.) Dumort.Lophocolea lateralis Dumort. = Lophocolea bidentata (L.) Dumort. var. rivularis (Raddi) Schiffn.Lophocolea latifolia Nees = Lophocolea bidentata (L.) Dumort. var. rivularis (Raddi) Schiffn.Lophocolea minor NeesLophocolea minor var. erosa (Nees) Gottsche et al. = Lophocolea minor NeesLophocolea profundus Nees = Lophocolea heterophylla (Schrad.) Dumort.Lophocolea reflexula Lindb. & Arnell = Lophocolea heterophylla (Schrad.) Dumort.Lophocolea setistipa Steph. = Lophocolea minor NeesLophozia acuta Dumort. = Leiocolea collaris (Nees) SchljakovLophozia acutiloba Schiffn. = Gymnocolea inflata subsp. acutiloba (Schiffn.) R.M. Schust. et Damsh. exSöderström et VáňaLophozia alpestris (Schleich. ex F. Web.) A. Evans = Leiocolea collaris (Nees) SchljakovLophozia alpestris auct. = Lophozia sudetica (Nees ex Huebener) GrolleLophozia alpestris var. amphigastriata (C.E.O. Jensen) Kaal. = Lophozia sudetica (Nees ex Huebener) GrolleLophozia alpestris var. gelida (Taylor) Macvicar = Lophozia sudetica (Nees ex Huebener) GrolleLophozia alpestris var. major (C.E.O. Jensen) Müll. Frib. = Lophozia excisa (Dicks.) Dumort.Lophozia ascendens (Warnst.) R.M. Schust.Lophozia attenuata (Mart.) Dumort. = Barbilophozia attenuata (Mart.) LoeskeLophozia badensis (Gottsche) Schiffn. = Leiocolea badensis (Gottsche) Jörg.Lophozia badensis var. obtusiloba (Bernet) Schiffn. = Leiocolea badensis (Gottsche) Jörg.Lophozia bantriensis (Hook.) Steph. = Leiocolea bantriensis (Hook.) Jörg.Lophozia barbata (Schmidel ex Schreb.) Dumort. = Barbilophozia barbata (Schmidel ex Schreb.) LoeskeLophozia baueriana (Schiffn.) Schiffn. = Barbilophozia hatcheri (A. Evans) LoeskeLophozia bicrenata (Schmidel ex Hoffm.) Dumort.Lophozia collaris (Nees) Dumort. = Leiocolea collaris (Nees) SchljakovLophozia confertifolia Schiffn. = Lophozia ventricosa (Dicks.) Dumort.Lophozia cylindracea (Dumort.) Dumort. = Lophozia excisa (Dicks.) Dumort.Lophozia ehrhartiana (F. Web.) Inoue & Steere = Lophozia ventricosa (Dicks.) Dumort.Lophozia ehrhartiana auct. = Lophozia sudetica (Nees ex Huebener) GrolleLophozia ehrhartiana F. Web. = Lophozia ventricosa (Dicks.) Dumort.← Lophozia elongata Steph.Lophozia excisa (Dicks.) Dumort.Lophozia excisa var. arenaria (Nees) Schiffn. = Lophozia excisa (Dicks.) Dumort.Lophozia excisa var. cylindracea (Dumort.) Müll. Frib. = Lophozia excisa (Dicks.) Dumort.Lophozia excisa var. jurensis (Meyl. ex Müll. Frib.) Müll. Frib. = Lophozia excisa (Dicks.) Dumort.Lophozia excisa var. limprichtii (Lindb. ex C. Massal. & Carestia) Schiffn. = Lophozia excisa (Dicks.) Dumort.Lophozia exsecta (Schmidel ex Schrad.) Dumort. = Tritomaria exsecta (Schmidel ex Schrad.) Schiffn. ex LoeskeLophozia exsectiformis (Breidl.) Jaap = Tritomaria exsectiformis (Breidl.) LoeskeLophozia fauriana Steph. = Lophozia ventricosa (Dicks.) Dumort.Lophozia floerkei (F. Web. & D. Mohr) Schiffn. = Barbilophozia floerkei (F. Web. & D. Mohr) LoeskeLophozia floerkei var. baueriana Schiffn. = Barbilophozia hatcheri (A. Evans) LoeskeLophozia floerkei var. naumanniana Schiffn. = Barbilophozia floerkei (F. Web. & D. Mohr) LoeskeLophozia fluitans (Nees) Boulay = Cladopodiella fluitans (Nees) H. BuchLophozia gelida (Taylor) Steph. = Lophozia sudetica (Nees ex Huebener) GrolleLophozia gillmanii (Aust.) R.M. Schust. = Leiocolea gillmanii (Aust.) A. EvansLophozia gracilis Schleich. ex Steph. = Barbilophozia attenuata (Mart.) LoeskeLophozia gracilis var. eflagellis Schiffn. = Barbilophozia attenuata (Mart.) LoeskeLophozia gracillima H. Buch = Lophozia ascendens (Warnst.) R.M. Schust.Lophozia groenlandica (Nees) Macoun = Lophozia wenzelii (Nees) Steph.Lophozia guttulata (Lindb. & Arnell) A. Evans = Lophozia longiflora (Nees) Schiffn.Lophozia gypsacea (Gottsche et al.) Schiffn. = Leiocolea badensis (Gottsche) Jörg.Lophozia hatcheri (A. Evans) Steph. = Barbilophozia hatcheri (A. Evans) LoeskeLophozia hatcheri var. ciliata Müll. Frib. = Barbilophozia lycopodioides (Wallr.) LoeskeLophozia hatcheri var. hageni Kaal. = Barbilophozia hatcheri (A. Evans) LoeskeLophozia helleriana (Nees ex Lindenb.) Macoun = Anastrophyllum hellerianum (Nees ex Lindenb.) R.M. Schust.Lophozia heterocolpos (Thed. ex C. Hartm.) M. Howe = Leiocolea heterocolpos (Thed. ex C. Hartm.) H. BuchLophozia hornschuchiana (Nees) Dumort. ex Schiffn. = Leiocolea bantriensis (Hook.) Jörg.Lophozia incisa (Schrad.) Dumort.Lophozia incisa subsp. opacifolia (Culm. ex Meyl.) R.M. Schust. et Damsh. = Lophozia opacifolia Culm. ex Meyl.427
Lophozia incisa var. inermis Müll. Frib. = Lophozia incisa (Schrad.) Dumort.Lophozia inflata (Huds.) M. Howe = Gymnocolea inflata (Huds.) Dumort.Lophozia inflata var. acutiloba Schiffn. = Gymnocolea inflata subsp. acutiloba (Schiffn.) R.M. Schust. et Damsh.ex Söderström et VáňaLophozia intermedia Dumort. = Lophozia excisa (Dicks.) Dumort.Lophozia jurensis Meyl. ex Müll. Frib. = Lophozia excisa (Dicks.) Dumort.Lophozia kaurinii (Limpr.) Steph. = Leiocolea gillmanii (Aust.) A. EvansLophozia kiaeri Jörg. = Lophozia excisa (Dicks.) Dumort.Lophozia kunzeana (Huebener) A. Evans = Barbilophozia kunzeana (Huebener) Müll. Frib.Lophozia kunzeana var. plicata (Hartm.) Lindb. = Barbilophozia kunzeana (Huebener) Müll. Frib.Lophozia limprichtii (Lindb. ex C. Massal. & Carestia) Steph. = Lophozia excisa (Dicks.) Dumort.Lophozia longidens (Lindb.) MacounLophozia longidens Schiffn. = Lophozia ascendens (Warnst.) R.M. Schust.Lophozia longiflora (Nees) Schiffn.Lophozia longiflora var. guttulata (Lindb. & Arnell) Schljakov = Lophozia longiflora (Nees) Schiffn.Lophozia lycopodioides (Wallr.) Cogn. = Barbilophozia lycopodioides (Wallr.) LoeskeLophozia lycopodioides var. obliqua Müll. Frib. = Barbilophozia lycopodioides (Wallr.) LoeskeLophozia lycopodioides var. parvifolia Schiffn. = Barbilophozia lycopodioides (Wallr.) LoeskeLophozia lyonii (Taylor) Steph. = Tritomaria quinquedentata (Huds.) H. BuchLophozia major (C.E.O. Jensen) Schljakov = Lophozia excisa (Dicks.) Dumort.Lophozia michauxii (F. Web.) Macoun = Anastrophyllum michauxii (F. Web.) H. BuchLophozia minuta (Schreb.) Schiffn. = Anastrophyllum minutum (Schreb.) R.M. Schust.Lophozia muelleri (Nees ex Lindenb.) Dumort. = Leiocolea collaris (Nees) SchljakovLophozia obtusa (Lindb.) A. EvansLophozia obtusa var. acutiloba Müll. Frib. = Lophozia obtusa (Lindb.) A. EvansLophozia opacifolia Culm. ex Meyl.Lophozia polita (Nees) Macoun = Tritomaria polita (Nees) Jörg.Lophozia porphyroleuca (C. Hartm.) Schiffn. = Lophozia longiflora (Nees) Schiffn.Lophozia porphyroleuca Müll. Frib. = Lophozia ascendens (Warnst.) R.M. Schust.Lophozia porphyroleuca var. guttulata (Lindb. & Arnell) Warnst. = Lophozia longiflora (Nees) Schiffn.Lophozia propagulifera (Gottsche) Steph. = Lophozia excisa (Dicks.) Dumort.Lophozia quadriloba (Lindb.) A. Evans = Barbilophozia quadriloba (Lindb.) LoeskeLophozia quadriloba var. glareosa (Jörg.) Bryhn = Barbilophozia quadriloba (Lindb.) LoeskeLophozia quinquedentata (Huds.) Cogn. = Tritomaria quinquedentata (Huds.) H. BuchLophozia quinquedentata var. tenera Arnell & C.E.O. Jensen = Tritomaria scitula (Taylor) Jörg.Lophozia saxicola (Schrad.) Schiffn. = Anastrophyllum saxicola (Schrad.) R.M. Schust.Lophozia scutata (F. Web. & D. Mohr) Dumort. = Harpanthus scutatus (F. Web. & D. Mohr) SpruceLophozia silvicola H. BuchLophozia silvicola var. grandiretis H. Buch & S.W. Arnell = Lophozia silvicola H. BuchLophozia ventricosa var. grandiretis (H. Buch & S.W. Arnell) R.M. Schust. & Damsh. = Lophozia silvicola H.BuchLophozia socia (Nees) Mig. = Lophozia excisa (Dicks.) Dumort.Lophozia sudetica (Nees ex Huebener) GrolleLophozia turbinata (Raddi) Steph. = Leiocolea turbinata (Raddi) H. BuchLophozia turbinata var. badensis (Gottsche ex Rabenh.) Hillier = Leiocolea badensis (Gottsche) Jörg.Lophozia ventricosa (Dicks.) Dumort.Lophozia ventricosa var. confertifolia (Schiffn.) Husn. = Lophozia ventricosa (Dicks.) Dumort.Lophozia ventricosa var. laxa (Lindenb.) Cogn. = Lophozia ventricosa (Dicks.) Dumort.Lophozia ventricosa var. longiflora (Nees) Macoun = Lophozia longiflora (Nees) Schiffn.Lophozia ventricosa var. longiflora auct., non (Nees) Macoun = Lophozia ventricosa (Dicks.) Dumort.Lophozia ventricosa var. porphyroleuca (C. Hartm.) Macoun = Lophozia longiflora (Nees) Schiffn.Lophozia ventricosa var. silvicola (H. Buch) E.W. Jones ex R.M. Schust. = Lophozia silvicola H. BuchLophozia ventricosa var. subarctica Schljakov = Lophozia ventricosa (Dicks.) Dumort.Lophozia ventricosa var. uliginosa Breidl. ex Schiffn. = Lophozia ventricosa (Dicks.) Dumort.Lophozia wenzelii (Nees) Steph.Lophoziella integerrima (Lindb.) Douin = Cephaloziella integerrima (Lindb.) Warnst.Lophoziella piriflora (Douin) Douin = Cephaloziella integerrima (Lindb.) Warnst.Lunularia alpina Nees = Sauteria alpina (Nees) NeesLunularia cruciata (L.) Lindb.Lunularia dillenii Le Jol. = Lunularia cruciata (L.) Lindb.428
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SORIN ŞTEFĂNUŢThe Hornwort and L
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CONTENTINTRODUCTION ...............
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The beginning of the 20 th century
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asin of the Arieş river. Bryotaxon
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Abbreviations used in the text† -
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The presentation of the methodology
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Fig. 3. The GEOCOD mapping system.
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THE HORNWORT AND LIVERWORT SPECIES
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THE HORNWORT AND LIVERWORT CLASSIFI
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Suborder Lepidoziineae R.M. Schust.
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46. Cephalozia pleniceps (Austin) L
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147. Pallavicinia lyellii (Hook.) C
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13 Corsinia coriandrina (Spreng.) L
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11. Porella obtusata (Taylor) Trevi
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34Phaeoceros carolinianus (Michx.)
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Făgăraş Mountains, Cârţişoara
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Cluj County, Someşul Rece Valley a
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Parâng Mountains, Repedea Valley,
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Argeş County, Rucăr (Loitlesberge
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Cindrel Mountains, Cindrel, 2200 m
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Parâng Mountains, Latoriţa Valley
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Athalamia hyalina (Sommerf.) S. Hat
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Barbilophozia barbata (Schmidel ex
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Bârgău Mountains, Cârţibavu Mar
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First citation for Romania: Schur 1
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- Pleuroschisma triangulare (Schlei
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and Băiuţ, common spruce forest,
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Distribution in Europe: Sj, Is, Fo,
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Poiana Stampei Peatbog, SV, 47°20'
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alt., 8.08.2003, leg. Pop O., det.
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leg. & det. Ştefureac T., ass. Buc
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Rodna Mountains, Puzdra Mountain, 1
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Alba County, Vârciorog Valley, 780
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Bârgău Mountains, Cucureasa Reser
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Suceava County, Tinovul Mare de la
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Suceava County, Rodna Nouă, on Per
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2001:74,75); Arieşul Mare Basin, o
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Ţibleş Mountains, Mesteacănul Va
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Cindrel Mountains, Găuşoara, leg.
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Parâng Mountains, the edge of Câl
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& det. Ştefănuţ S. [BUCA B3684];
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Alba County, at 20,5 km to Zlatna,
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Cephaloziella spinigera (Lindb.) Wa
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First citation for Romania: Péterf
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Iezer-Păpuşa Mountains, Iezer Val
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- Cladopodiella fluitans var. gigan
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Caraş-Severin County, Cerna Valley
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(Mititelu, Barabaş N. & al. 1968:1
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1885:30; Ştefureac 1949a:287,288;
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104Conocephalum salebrosum Szweyk.,
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Retezat Mountains, Retezat, sub Jun
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Cindrel Mountains, Cânaia, 1750 m
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Iaşi County, Sineşti Forest (Papp
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[BUCA B2793], on rotten beech trunk
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Orthodicrano montani-Hypnetum filif
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Distribution in Romania:Maramureş
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- Gymnocolea inflata var. laxa (Nee
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Ciucaş Mountains, Caprei Brook Val
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Giumalău Mountain, Giumalău Secul
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- Haplozia riparia var. rivularis B
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- Jungermannia karl-muelleri auct.,
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Jungermannia gracillima Sm.Syn.:- A
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Mountain, PH, 45°24'40"N / 25°28'
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Retezat Mountains, Rovine Brook, on
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Romania, alt. 600-2280 m (Váňa 19
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- Solenostoma subellipticum (Lindb.
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Distribution in Romania:Bârgău Mo
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13.07.1946, sub Lophozia alpestris
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Distribution in Romania:Maramureş
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Rodna Mountains, Lala Valley, 1200
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Ştefănuţ S. [BUCA B2574, B2580,
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Dealul Colbului and Pietrele Doamne
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150var. rivularis (Raddi) Warnst.Sy
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Penteleu Massif, fer - Roşu Lake,
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Satu Mare County, Noroieni, Ghiarma
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Lophozia ascendens (Warnst.) R.M. S
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- Jungermannia intermedia Lindenb.-
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Căliman Mountains, on humus and ro
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Hăşmaş Mountains, Hăşmaş, sub
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Herbaria and Exsiccate: BUCADistrib
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peatbog on the border of Câlcescu
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? - Iezer-Păpuşa Mountains, Iezer
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Făgăraş Mountains, s - above Avr
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Nemira Mountains, on edge of Nemira
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2003:7); Măgura Cisnădiei [SIB HL
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Fuss 1854:20; Ştefureac 1949a:288;
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- Nardia brevissima (Dumort.) Lindb
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1919:295); Ciortea Mountain, on wet
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- Sarcoscyphus sphacelatus var. med
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Siriu Mountain, Poiana din Fundul M
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Iaşi County, below Repedea, 7.10.1
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Valley, Arieşul Mic - upstream min
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Alba County, Valea Lungă, 10.02.18
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Rarău Massif, between Dealul Colbu
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Harghita County, Harghita Spa, on r
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Căliman Mountains, in alpine zone,
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Stânişoara Mountains, Suha Mare B
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Ecology: humicolous, acidophytic, h
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Bacău County, Slănic Brook (Papp
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Rarău Mountain, Codrul Secular Sl
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1915:197,198); Cândeşti, 07.1891,
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Ţibleş Mountains, Piciorul Ţigle
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Ceahlău Mountain, Durău Valley, i
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major (Ştefureac 1962b:69); Benea-
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Ciucaş Mountains, Berii Brook Vall
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Zlătuia Valley, 1100 m alt., 7.09.
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- Plagiochila dillenii (Taylor) Tay
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Wallfisch R. (Dihoru G. & al. 2003:
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Ştefănuţ M. 2003:79); Iezerul Pi
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utabuli-Mnietum cuspidati (Felföld
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Iaşi County, Mârzeşti Forest, Bu
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3.04.2004, leg. Atanasiu Paulina, d
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Arad County, Săvârşin, sub Madot
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Preissia quadrata (Scop.) NeesSyn.:
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Diana, BV, 1100 m alt., 22.08.2001,
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Rodna Mountains (Pax 1908:130); Pie
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Descut, 15.08.2001, Podul de Piatr
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Bârgău Mountains, Cucureasa Reser
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B3521, B3522] (Ştefănuţ 2006b:34
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16,20.08.1994, ass. Pterigynandretu
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- Marchantia crinita Michx.- Marcha
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Giumalău Mountain, Giumalău Secul
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Riccardia multifida (L.) GraySyn.:-
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Penteleu Massif, the left side of C
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Herbaria and Exsiccate: SIBDistribu
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558,560; Papp 1939a:358; Papp & Eft
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Riccia glauca L. var. glaucaSyn.:-
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Distribution in Europe: Is, Fo, No,
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Bihor County, Tărian, sub Riccia (
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1960:1113); upper side of Mălăie
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Scapania brevicaulis TaylorSyn.:- S
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Suceava County, Cristişor-Neagra B
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Piatra Mare Mountain, 1700 m alt.,
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- Scapania amurensis Warnst.- Scapa
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Mureş County, Sighişoara - Fuchsl
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Căliman Mountains, the peatbogs fr
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- Scapania heterophylla M. Howe- Sc
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Slăvei and Muchia Ascuţită, 2000
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Distribution in Europe: Ie, Br, Fr,
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Nemira Mountains, Nemira, on humus,
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- Sphenolobus medelpadicus (Arnell)
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Parâng Mountains, Latoriţa Valley
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Fig. 9. Distribution of Phaeoceros
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Fig. 13. Distribution of Anastrophy
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Fig. 17. Distribution of Anthelia j
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Fig. 21. Distribution of Asterella
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Fig. 25. Distribution of Barbilopho
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Fig. 29. Distribution of Barbilopho
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Fig. 33. Distribution of Bazzania t
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Fig. 37. Distribution of Calypogeia
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Fig. 41. Distribution of Calypogeia
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Fig. 45. Distribution of Cephalozia
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Fig. 49. Distribution of Cephalozia
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Fig. 53. Distribution of Cephalozie
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Fig. 65. Distribution of Cololejeun
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Fig. 69. Distribution of Corsinia c
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Fig. 73. Distribution of Fossombron
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Fig. 77. Distribution of Frullania
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Fig. 81. Distribution of Gymnocolea
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Fig. 85. Distribution of Harpanthus
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Fig. 89. Distribution of Jungermann
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Fig. 101. Distribution of Jungerman
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Fig. 105. Distribution of Leiocolea
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Fig. 109. Distribution of Lejeunea
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Fig. 113. Distribution of Lophocole
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Fig. 117. Distribution of Lophozia
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Fig. 121. Distribution of Lophozia
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Fig. 125. Distribution of Lunularia
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Fig. 129. Distribution of Marchanti
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Fig. 133. Distribution of Marsupell
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Fig. 137. Distribution of Marsupell
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Fig. 141. Distribution of Moerckia
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Fig. 145. Distribution of Nardia ge
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Fig. 149. Distribution of Odontosch
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Fig. 153. Distribution of Pedinophy
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Fig. 157. Distribution of Peltolepi
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Fig. 161. Distribution of Porella a
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Fig. 165. Distribution of Preissia
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Fig. 169. Distribution of Radula li
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Fig. 173. Distribution of Riccardia
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Fig. 177. Distribution of Riccia ca
Page 377 and 378: Fig. 181. Distribution of Riccia cr
Page 379 and 380: Fig. 185. Distribution of Riccia go
Page 381 and 382: Fig. 189. Distribution of Ricciocar
Page 383 and 384: Fig. 193. Distribution of Scapania
Page 395 and 396: Fig. 217. Distribution of Tritomari
Page 397 and 398: Evaluating the distribution dataFol
Page 399 and 400: Fig. 224. Hornworts and Liverworts
Page 401 and 402: Signs used:Bolded names - species i
Page 403 and 404: Marchantiaceae50. VU - Bucegia roma
Page 405 and 406: Fig. 228. The distribution of zones
Page 407 and 408: Anthoceros communis Steph. = Phaeoc
Page 409 and 410: Blasia funckii Corda = Blasia pusil
Page 411 and 412: Cephalozia crassiflora Spruce = Cep
Page 413 and 414: Cephaloziella spinigera (Lindb.) Wa
Page 415 and 416: Diplomitrium lyellii (Hook.) Corda
Page 417 and 418: Gymnocolea inflata subsp. acutiloba
Page 419 and 420: Jungermannia acuta var. gypsacea Go
Page 421 and 422: Jungermannia dentata Raddi = Cephal
Page 423 and 424: Jungermannia limprichtii Lindb. = L
Page 425 and 426: Jungermannia scalaris var. denudata
Page 427: Leiocolea heterocolpos (Thed. ex C.
Page 431 and 432: Marchantia polymorpha var. dilatata
Page 433 and 434: Mastigobryum deflexum (Mart.) Ångs
Page 435 and 436: Odontoschisma sphagni var. denudatu
Page 437 and 438: Porella platyphylla (L.) Pfeiff.Por
Page 439 and 440: Riccia maior Roth = Riccia glauca L
Page 441 and 442: Scapania heterophylla M. Howe = Sca
Page 443 and 444: Sphagnocetis communis var. macrior
Page 445 and 446: Mittheilungen des siebenbürgischen
Page 447 and 448: 91. Borza A., 1922a. Schedae ad "Fl
Page 449 and 450: 142. Boşcaiu N., Täuber F., Colde
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The Hornwort and Liverwort Atlas of Romania

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