The Hornwort and Liverwort Atlas of Romania

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Jungermannia plicata var. kunzeana (Huebener) Hartm. = Barbilophozia kunzeana (Huebener) Müll. Frib.Jungermannia polaris Lindb.Jungermannia polita Nees 1836 = Tritomaria polita (Nees) Jörg.Jungermannia polita var. acuta (Lindb. & Arnell) Meyl. = Tritomaria scitula (Taylor) Jörg.Jungermannia polyanthos L. 1753 = Chiloscyphus polyanthos (L.) CordaJungermannia polyanthos var. pallescens (Ehrh. ex Hoffm.) Lindenb. = Chiloscyphus pallescens (Ehrh. ex Hoffm.)Dumort. var. pallescensJungermannia polyanthos var. rivularis (Schrad.) Lindb. = Chiloscyphus polyanthos var. rivularis (Schrad.)Gottsche et al.Jungermannia porelloides Torr. ex Nees 1833 = Plagiochila porelloides (Torr. ex Nees) Lindenb.Jungermannia porphyroleuca Nees = Lophozia longiflora (Nees) Schiffn.Jungermannia porphyroleuca var. attenuata Nees = Lophozia longidens (Lindb.) MacounJungermannia porphyroleuca var. guttulata (Lindb. & Arnell) Warnst. = Lophozia longiflora (Nees) Schiffn.Jungermannia potamophila Müll. Arg. ex Moug. et al. = Jungermannia atrovirens Dumort.Jungermannia propagulifera Gottsche = Lophozia excisa (Dicks.) Dumort.Jungermannia pubescens Schrank 1792 = Apometzgeria pubescens (Schrank) Kuwah.Jungermannia pulcherrima Weber 1778 = Ptilidium pulcherrimum (Weber) Vain.Jungermannia pulvinata Raddi = Marsupella emarginata (Ehrh.) Dumort.Jungermannia pumila Dicks.Jungermannia pumila Limpr., non With. = Jungermannia atrovirens Dumort.Jungermannia pumila subsp. polaris (Lindb.) Berggr. = Jungermannia polaris Lindb.Jungermannia pumila var. notha Gottsche = Jungermannia atrovirens Dumort.Jungermannia pumila var. alpestris Lindb. = Jungermannia pumila Dicks.Jungermannia pumila var. polaris (Lindb.) Berggr. = Jungermannia polaris Lindb.Jungermannia pumila var. rivularis (Schiffn.) Frye & L. Clark = Jungermannia pumila Dicks.Jungermannia pumila var. sauteriana Gottsche = Jungermannia atrovirens Dumort.Jungermannia pumila var. sphaerocarpoidea (De Not.) C. Massal. = Jungermannia atrovirens Dumort.Jungermannia punctata Gottsche = Jungermannia caespiticia Lindenb.Jungermannia pusilla L., non (C.E.O. Jensen) H. Buch = Fossombronia pusilla (L.) NeesJungermannia pusilla var. capitata Nees = Fossombronia wondraczekii (Corda) Lindb.Jungermannia quadriloba Lindb. 1883 = Barbilophozia quadriloba (Lindb.) LoeskeJungermannia quadriloba var. glareosa Jörg. = Barbilophozia quadriloba (Lindb.) LoeskeJungermannia quinquedentata Huds. 1762 = Tritomaria quinquedentata (Huds.) H. BuchJungermannia quinquedentata var. attenuata Mart. = Barbilophozia attenuata (Mart.) LoeskeJungermannia raddiana C. Massal. = Cephaloziella rubella (Nees) Warnst.Jungermannia radicans L. = Bazzania trilobata (L.) GrayJungermannia rauana Steph. = Jamesoniella autumnalis (DC.) Steph.Jungermannia reclusa Taylor = Cephalozia catenulata (Huebener) Lindb.Jungermannia renifolia Lehm. = Scapania uliginosa (Sw. ex Lindenb.) Dumort.Jungermannia repens Dietrich = Jungermannia pumila Dicks.Jungermannia reptans L. 1753 = Lepidozia reptans (L.) Dumort.Jungermannia resupinata Hook. = Scapania compacta (Roth) Dumort.Jungermannia resupinata L., non Hook. = Scapania undulata (L.) Dumort.Jungermannia rigida (Lindb.) Lindb. = Anastrophyllum minutum (Schreb.) R.M. Schust.Jungermannia rigida Wallr. = Aneura pinguis (L.) Dumort.Jungermannia rigidula Huebener = Cephalozia bicuspidata (L.) Dumort.Jungermannia riparia Taylor = Jungermannia atrovirens Dumort.Jungermannia riparia var. bactrocalyx C. Massal. = Jungermannia atrovirens Dumort.Jungermannia riparia var. rivularis (Bernet) Meyl. = Jungermannia atrovirens Dumort.Jungermannia rosacea Corda = Scapania curta (Mart.) Dumort.Jungermannia rostellata Huebener = Jungermannia pumila Dicks.Jungermannia rotaeana (De Not.) Bertol. = Nardia scalaris GrayJungermannia rubella Nees 1834 = Cephaloziella rubella (Nees) Warnst.Jungermannia rupestris Schleich. = Scapania aequiloba (Schwägr.) Dumort.Jungermannia saccatula Lindb. = Anastrophyllum minutum (Schreb.) R.M. Schust.Jungermannia saxicolus Schrad. 1796 = Anastrophyllum saxicola (Schrad.) R.M. Schust.Jungermannia scalariformis Nees = Jungermannia confertissima NeesJungermannia scalariformis var. scandica C.E.O. Jensen ex S.W. Arnell = Jungermannia confertissima NeesJungermannia scalaris (Gray) Hook., non Schmidel = Nardia scalaris GrayJungermannia scalaris var. colorata Nees = Jungermannia gracillima Sm.423
Jungermannia scalaris var. denudata Mart. 1817 = Odontoschisma denudatum (Mart.) Dumort.Jungermannia scalaris var. minor Nees = Nardia geoscyphus (De Not.) Lindb.Jungermannia scalaris var. repanda Huebener = Nardia geoscyphus (De Not.) Lindb.Jungermannia schiffneri (Loitl.) A. Evans, non (Loitl.) Steph. = Jungermannia polaris Lindb.Jungermannia schmidelianum Huebener = Jungermannia hyalina LyellJungermannia schraderi Nees = Jamesoniella autumnalis (DC.) Steph.Jungermannia schreberi (Nees) Nees = Barbilophozia barbata (Schmidel ex Schreb.) LoeskeJungermannia schultzii Spreng. = Kurzia pauciflora (Dicks.) GrolleJungermannia scitula Taylor 1846 = Tritomaria scitula (Taylor) Jörg.Jungermannia scutata F. Web. & D. Mohr = Harpanthus scutatus (F. Web. & D. Mohr) SpruceJungermannia serpentina Nees = Leiocolea collaris (Nees) SchljakovJungermannia setacea auct., non Weber = Kurzia pauciflora (Dicks.) GrolleJungermannia sicca Nees = Lophozia sudetica (Nees ex Huebener) GrolleJungermannia sinuata Dicks. = Pallavicinia lyellii (Hook.) Carruth.Jungermannia sinuata Dicks. = Riccardia chamaedryfolia (With.) GrolleJungermannia socia Nees = Lophozia excisa (Dicks.) Dumort.Jungermannia sphacelata Gieseke ex Lindenb. 1829 = Marsupella sphacelata (Gieseke ex Lindenb.) Dumort.Jungermannia sphaerocarpa auct., non Hook. = Jungermannia confertissima NeesJungermannia sphaerocarpa Hook.Jungermannia sphaerocarpa var. amplexicaulis (Dumort.) Frye & L. Clark = Jungermannia sphaerocarpa Hook.Jungermannia sphaerocarpa var. elongata Huebener = Jungermannia sphaerocarpa Hook.Jungermannia sphaerocarpa var. gracilescens Nees = Jungermannia gracillima Sm.Jungermannia sphaerocarpa var. lurida Pearson = Jungermannia sphaerocarpa Hook.Jungermannia sphaerocarpa var. nana (Nees ex Breidl.) Frye & L. Clark = Jungermannia sphaerocarpa Hook.Jungermannia sphaerocarpoidea De Not. = Jungermannia atrovirens Dumort.Jungermannia sphagni Dicks. = Odontoschisma sphagni (Dicks.) Dumort.Jungermannia sphagni var. vegetior Nees = Odontoschisma sphagni (Dicks.) Dumort.Jungermannia sprengelii Nees = Calypogeia fissa (L.) RaddiJungermannia starkei Funck ex Nees = Cephaloziella divaricata (Sm.) Schiffn.Jungermannia stipulacea Hook. = Harpanthus scutatus (F. Web. & D. Mohr) SpruceJungermannia subalpina Nees ex Lindenb. 1829 = Scapania subalpina (Nees ex Lindenb.) Dumort.Jungermannia subapicalis Huebener, non Nees = Nardia scalaris GrayJungermannia subapicalis Nees, non Huebener = Jamesoniella autumnalis (DC.) Steph.Jungermannia subdichotoma Lindb. = Gymnocolea inflata (Huds.) Dumort.Jungermannia subelliptica (Lindb. ex Kaal.) LevierJungermannia subulata A. EvansJungermannia sudetica Nees ex Huebener 1834 = Lophozia sudetica (Nees ex Huebener) GrolleJungermannia sullivantii Austin = Cephaloziella rubella (Nees) Warnst.Jungermannia supina Taylor = Lophozia incisa (Schrad.) Dumort.Jungermannia tamarisci L. 1753 = Frullania tamarisci (L.) Dumort.Jungermannia tamariscifolia L. = Frullania tamarisci (L.) Dumort.Jungermannia taxifolia Wahlenb. 1812 = Diplophyllum taxifolium (Wahlenb.) Dumort.Jungermannia taylorii Hook. 1816 = Mylia taylorii (Hook.) GrayJungermannia taylorii var. anomalus (Hook.) Nees = Mylia anomala (Hook.) GrayJungermannia tenerrima Raddi, non Steph. = Jungermannia gracillima Sm.Jungermannia tersa Nees = Jungermannia sphaerocarpa Hook.Jungermannia tersa var. attenuata Nees = Jungermannia obovata NeesJungermannia tersa var. densa Nees = Jungermannia obovata NeesJungermannia tersa var. explanata Nees = Jungermannia obovata NeesJungermannia tersa var. rivularis Nees = Jungermannia obovata NeesJungermannia tomentella Ehrh. 1785 = Trichocolea tomentella (Ehrh.) Dumort.Jungermannia towadensis (S. Okamura) S. Hatt. = Jungermannia atrovirens Dumort.Jungermannia treviranii Huebener = Anastrophyllum minutum (Schreb.) R.M. Schust.Jungermannia triangularis Schleich. ex Steud. = Bazzania tricrenata (Wahlenb.) Lindb.Jungermannia trichomanis (L.) Dicks. = Calypogeia azurea Stotler & CrotzJungermannia trichomanis (L.) Scop. = Calypogeia fissa (L.) RaddiJungermannia tricophylla L. 1753 = Blepharostoma trichophyllum (L.) Dumort.Jungermannia tricrenata Wahlenb. 1814 = Bazzania tricrenata (Wahlenb.) Lindb.Jungermannia tridenticulata Michx. = Bazzania trilobata (L.) GrayJungermannia trilobata L. 1753 = Bazzania trilobata (L.) Gray424
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SORIN ŞTEFĂNUŢThe Hornwort and L
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CONTENTINTRODUCTION ...............
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The beginning of the 20 th century
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asin of the Arieş river. Bryotaxon
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Abbreviations used in the text† -
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The presentation of the methodology
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Fig. 3. The GEOCOD mapping system.
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THE HORNWORT AND LIVERWORT SPECIES
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THE HORNWORT AND LIVERWORT CLASSIFI
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Suborder Lepidoziineae R.M. Schust.
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46. Cephalozia pleniceps (Austin) L
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147. Pallavicinia lyellii (Hook.) C
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13 Corsinia coriandrina (Spreng.) L
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11. Porella obtusata (Taylor) Trevi
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34Phaeoceros carolinianus (Michx.)
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Făgăraş Mountains, Cârţişoara
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Cluj County, Someşul Rece Valley a
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Parâng Mountains, Repedea Valley,
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Argeş County, Rucăr (Loitlesberge
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Cindrel Mountains, Cindrel, 2200 m
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Parâng Mountains, Latoriţa Valley
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Athalamia hyalina (Sommerf.) S. Hat
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Barbilophozia barbata (Schmidel ex
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Bârgău Mountains, Cârţibavu Mar
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First citation for Romania: Schur 1
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- Pleuroschisma triangulare (Schlei
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and Băiuţ, common spruce forest,
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Distribution in Europe: Sj, Is, Fo,
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Poiana Stampei Peatbog, SV, 47°20'
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alt., 8.08.2003, leg. Pop O., det.
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leg. & det. Ştefureac T., ass. Buc
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Rodna Mountains, Puzdra Mountain, 1
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Alba County, Vârciorog Valley, 780
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Bârgău Mountains, Cucureasa Reser
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Suceava County, Tinovul Mare de la
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Suceava County, Rodna Nouă, on Per
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2001:74,75); Arieşul Mare Basin, o
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Ţibleş Mountains, Mesteacănul Va
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Cindrel Mountains, Găuşoara, leg.
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Parâng Mountains, the edge of Câl
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& det. Ştefănuţ S. [BUCA B3684];
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Alba County, at 20,5 km to Zlatna,
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Cephaloziella spinigera (Lindb.) Wa
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First citation for Romania: Péterf
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Iezer-Păpuşa Mountains, Iezer Val
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- Cladopodiella fluitans var. gigan
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Caraş-Severin County, Cerna Valley
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(Mititelu, Barabaş N. & al. 1968:1
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1885:30; Ştefureac 1949a:287,288;
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104Conocephalum salebrosum Szweyk.,
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Retezat Mountains, Retezat, sub Jun
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Cindrel Mountains, Cânaia, 1750 m
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Iaşi County, Sineşti Forest (Papp
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[BUCA B2793], on rotten beech trunk
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Orthodicrano montani-Hypnetum filif
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Distribution in Romania:Maramureş
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- Gymnocolea inflata var. laxa (Nee
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Ciucaş Mountains, Caprei Brook Val
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Giumalău Mountain, Giumalău Secul
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- Haplozia riparia var. rivularis B
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- Jungermannia karl-muelleri auct.,
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Jungermannia gracillima Sm.Syn.:- A
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Mountain, PH, 45°24'40"N / 25°28'
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Retezat Mountains, Rovine Brook, on
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Romania, alt. 600-2280 m (Váňa 19
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- Solenostoma subellipticum (Lindb.
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Distribution in Romania:Bârgău Mo
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13.07.1946, sub Lophozia alpestris
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Distribution in Romania:Maramureş
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Rodna Mountains, Lala Valley, 1200
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Ştefănuţ S. [BUCA B2574, B2580,
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Dealul Colbului and Pietrele Doamne
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150var. rivularis (Raddi) Warnst.Sy
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Penteleu Massif, fer - Roşu Lake,
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Satu Mare County, Noroieni, Ghiarma
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Lophozia ascendens (Warnst.) R.M. S
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- Jungermannia intermedia Lindenb.-
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Căliman Mountains, on humus and ro
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Hăşmaş Mountains, Hăşmaş, sub
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Herbaria and Exsiccate: BUCADistrib
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peatbog on the border of Câlcescu
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? - Iezer-Păpuşa Mountains, Iezer
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Făgăraş Mountains, s - above Avr
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Nemira Mountains, on edge of Nemira
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2003:7); Măgura Cisnădiei [SIB HL
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Fuss 1854:20; Ştefureac 1949a:288;
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- Nardia brevissima (Dumort.) Lindb
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1919:295); Ciortea Mountain, on wet
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- Sarcoscyphus sphacelatus var. med
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Siriu Mountain, Poiana din Fundul M
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Iaşi County, below Repedea, 7.10.1
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Valley, Arieşul Mic - upstream min
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Alba County, Valea Lungă, 10.02.18
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Rarău Massif, between Dealul Colbu
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Harghita County, Harghita Spa, on r
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Căliman Mountains, in alpine zone,
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Stânişoara Mountains, Suha Mare B
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Ecology: humicolous, acidophytic, h
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Bacău County, Slănic Brook (Papp
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Rarău Mountain, Codrul Secular Sl
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1915:197,198); Cândeşti, 07.1891,
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Ţibleş Mountains, Piciorul Ţigle
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Ceahlău Mountain, Durău Valley, i
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major (Ştefureac 1962b:69); Benea-
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Ciucaş Mountains, Berii Brook Vall
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Zlătuia Valley, 1100 m alt., 7.09.
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- Plagiochila dillenii (Taylor) Tay
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Wallfisch R. (Dihoru G. & al. 2003:
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Ştefănuţ M. 2003:79); Iezerul Pi
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utabuli-Mnietum cuspidati (Felföld
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Iaşi County, Mârzeşti Forest, Bu
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3.04.2004, leg. Atanasiu Paulina, d
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Arad County, Săvârşin, sub Madot
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Preissia quadrata (Scop.) NeesSyn.:
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Diana, BV, 1100 m alt., 22.08.2001,
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Rodna Mountains (Pax 1908:130); Pie
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Descut, 15.08.2001, Podul de Piatr
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Bârgău Mountains, Cucureasa Reser
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B3521, B3522] (Ştefănuţ 2006b:34
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16,20.08.1994, ass. Pterigynandretu
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- Marchantia crinita Michx.- Marcha
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Giumalău Mountain, Giumalău Secul
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Riccardia multifida (L.) GraySyn.:-
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Penteleu Massif, the left side of C
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Herbaria and Exsiccate: SIBDistribu
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558,560; Papp 1939a:358; Papp & Eft
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Riccia glauca L. var. glaucaSyn.:-
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Distribution in Europe: Is, Fo, No,
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Bihor County, Tărian, sub Riccia (
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1960:1113); upper side of Mălăie
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Scapania brevicaulis TaylorSyn.:- S
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Suceava County, Cristişor-Neagra B
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Piatra Mare Mountain, 1700 m alt.,
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- Scapania amurensis Warnst.- Scapa
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Mureş County, Sighişoara - Fuchsl
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Căliman Mountains, the peatbogs fr
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- Scapania heterophylla M. Howe- Sc
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Slăvei and Muchia Ascuţită, 2000
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Distribution in Europe: Ie, Br, Fr,
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Nemira Mountains, Nemira, on humus,
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- Sphenolobus medelpadicus (Arnell)
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Parâng Mountains, Latoriţa Valley
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Fig. 9. Distribution of Phaeoceros
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Fig. 13. Distribution of Anastrophy
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Fig. 17. Distribution of Anthelia j
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Fig. 21. Distribution of Asterella
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Fig. 25. Distribution of Barbilopho
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Fig. 29. Distribution of Barbilopho
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Fig. 33. Distribution of Bazzania t
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Fig. 37. Distribution of Calypogeia
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Fig. 41. Distribution of Calypogeia
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Fig. 45. Distribution of Cephalozia
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Fig. 49. Distribution of Cephalozia
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Fig. 53. Distribution of Cephalozie
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Page 317 and 318:
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Fig. 65. Distribution of Cololejeun
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Fig. 69. Distribution of Corsinia c
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Fig. 73. Distribution of Fossombron
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Fig. 77. Distribution of Frullania
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Fig. 81. Distribution of Gymnocolea
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Fig. 85. Distribution of Harpanthus
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Fig. 89. Distribution of Jungermann
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Page 335 and 336:
Page 337 and 338:
Fig. 101. Distribution of Jungerman
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Fig. 105. Distribution of Leiocolea
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Fig. 109. Distribution of Lejeunea
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Fig. 113. Distribution of Lophocole
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Fig. 117. Distribution of Lophozia
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Fig. 121. Distribution of Lophozia
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Fig. 125. Distribution of Lunularia
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Fig. 129. Distribution of Marchanti
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Fig. 133. Distribution of Marsupell
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Fig. 137. Distribution of Marsupell
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Fig. 141. Distribution of Moerckia
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Fig. 145. Distribution of Nardia ge
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Fig. 149. Distribution of Odontosch
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Fig. 153. Distribution of Pedinophy
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Fig. 157. Distribution of Peltolepi
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Fig. 161. Distribution of Porella a
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Fig. 165. Distribution of Preissia
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Fig. 169. Distribution of Radula li
Page 373 and 374: Fig. 173. Distribution of Riccardia
Page 375 and 376: Fig. 177. Distribution of Riccia ca
Page 377 and 378: Fig. 181. Distribution of Riccia cr
Page 379 and 380: Fig. 185. Distribution of Riccia go
Page 381 and 382: Fig. 189. Distribution of Ricciocar
Page 383 and 384: Fig. 193. Distribution of Scapania
Page 395 and 396: Fig. 217. Distribution of Tritomari
Page 397 and 398: Evaluating the distribution dataFol
Page 399 and 400: Fig. 224. Hornworts and Liverworts
Page 401 and 402: Signs used:Bolded names - species i
Page 403 and 404: Marchantiaceae50. VU - Bucegia roma
Page 405 and 406: Fig. 228. The distribution of zones
Page 407 and 408: Anthoceros communis Steph. = Phaeoc
Page 409 and 410: Blasia funckii Corda = Blasia pusil
Page 411 and 412: Cephalozia crassiflora Spruce = Cep
Page 413 and 414: Cephaloziella spinigera (Lindb.) Wa
Page 415 and 416: Diplomitrium lyellii (Hook.) Corda
Page 417 and 418: Gymnocolea inflata subsp. acutiloba
Page 419 and 420: Jungermannia acuta var. gypsacea Go
Page 421 and 422: Jungermannia dentata Raddi = Cephal
Page 423: Jungermannia limprichtii Lindb. = L
Page 427 and 428: Leiocolea heterocolpos (Thed. ex C.
Page 429 and 430: Lophozia incisa var. inermis Müll.
Page 431 and 432: Marchantia polymorpha var. dilatata
Page 433 and 434: Mastigobryum deflexum (Mart.) Ångs
Page 435 and 436: Odontoschisma sphagni var. denudatu
Page 437 and 438: Porella platyphylla (L.) Pfeiff.Por
Page 439 and 440: Riccia maior Roth = Riccia glauca L
Page 441 and 442: Scapania heterophylla M. Howe = Sca
Page 443 and 444: Sphagnocetis communis var. macrior
Page 445 and 446: Mittheilungen des siebenbürgischen
Page 447 and 448: 91. Borza A., 1922a. Schedae ad "Fl
Page 449 and 450: 142. Boşcaiu N., Täuber F., Colde
Page 451 and 452: 199. Coldea G., 1975. Étude phytos
Page 453 and 454: 256. Csűrös Ş., Moldovan I., Cs
Page 455 and 456: 317. Dihoru G., 1987b. Contribuţie
Page 457 and 458: 367. Domin K., 1923. Grundzüge der
Page 459 and 460: 426. Gergely I., 1969. Pajişti mez
Page 461 and 462: 487. Györffy I., 1906c. Polytrichu
Page 463 and 464: 540. Györffy I., 1943b. Erdélyi P
Page 465 and 466: 598. Ionescu Venera, Cristurean I.,
Page 467 and 468: 658. Lungu Lucia, 1967a. Contribuţ
Page 469 and 470: 712. Mihai G., 1971e. Aspecte ale b
Page 471 and 472: 767. Mihăilescu Simona, Ştefănu
Page 473 and 474: 824. Mohan G., 1988c. Die Gattung T
Page 475 and 476:
879. Niţu I.T, 1973. Solurile cu e
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941. Papp C., 1931b. O schiţă a v
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999. Papp C., Eftimie Elena, 1963.
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1059. Péterfi L.Ş., 1963. Flagela
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1119. Plămadă, Viţalariu, 1971.
Page 485 and 486:
1178. Pop I., Hodişan I., 1958. Co
Page 487 and 488:
1231. Prunescu-Arion Elena, Toniuc
Page 489 and 490:
1285. Raţiu O., 1970. Consideraţi
Page 491 and 492:
1342. Rüegsegger F., 1986. Frullan
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1396. Schiffner V., 1913b. Über ei
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var. fragilis (A. Roth) Müll. Frib
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Riccardia incurvata Lindb. 28, 248,
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The Hornwort and Liverwort Atlas of Romania

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