alt., leg. & det. Boros Á, Ciceu (“Csicsói”) Mountain, 1900 m alt., leg. & det. Pócs T. (Boros & Vajda1967:223,224; Bálint & Orbán 2006:470); Greu Mountain, 1100 m alt., 11.07.2000, Minei Valley, 1100-1200 m alt., 11.07.2000, Vermet Valley, 1050-1250 m alt., 2.07.2000, Asău Valley, 1000-1100 m alt.,22.10.2000, Mic Valley, 1050-1150 m alt., 16.08.2001, Pietros Valley, 900 m alt., 28.08.2001, UlieşulMic, 1150 m alt., 17.04.2001, Luci Meleche Forest, 1050 m alt., 13.08.2001, leg. Bálint L. & Csízi K.(Bálint & Orbán 2003:14; Bálint & Orbán 2006:470); Mohoş, sub S. dentata (Bálint & Orbán 2006:470).Harghita County, Piatra Piscului, leg. & det. Demeter K. [CL](Igmándy 1943:56); Harghita,26.07.1901 (Barth 1905:13); Chirui Spa from Lueta, 14.08.1869, leg. & det. Barth J. [SIB HF 1554](Fuss1878:657; Drăgulescu 1983:58).Neamţ County, Cernegura Mountain, Sterghinosu Brook Valley, 400-550 m alt. (Lungu 1963a:242).Bacău County, Şurei Hill, in forest, near Slănic-Moldova Spa, on right side <strong>of</strong> Slănic Brook, 500m alt., sub „S. penata” (Papp 1969a:181); Slănic Brook, on rocks, 2.05-8.11.1971 (Ştefureac & BarabaşV. 1972:78); Ruginii Brook, incl. sub var. dentata (Eftimie 1972:151); without locality, 450 m alt.(Barabaş V. 1982b:70, Tab.20).Ciucaş Mountains, Berii Brook Valley, Alb Brook Valley, Sterp Brook Valley (Mohan 1990b:43);Gârbova Massif, Cazacului Valley, Azuga’s affluent, 950 m alt., det. Ştefureac T. (Badea &Cretzoiu 1941:285).Prahova County, Teleajen Valley, (Mohan 1990a:24).Făgăraş Mountains, Arpaş Valley, incl. sub var. rivularis, common spruce forest (Heufler1853:48-49; Fuss 1854:20; Schur 1866:875; Fuss 1878:657; Hazslinszky 1885:67; Drăgulescu 2003:16);Cârţişoara Mountains, leg. & det. Barth J. (Fuss 1878:657); Bâlea (Schur 1866:875; Fuss 1878:657;Hazslinszky 1885:67; Drăgulescu 2003:16); Negoiu, 2000 m alt., leg. & det. Loitlesberger K., sub var.purpurea [Krypt. Exs. 473] (Loitlesberger 1898:193,194; Zahlbruckner 1900:188; Drăgulescu 2003:16);Topolog Valley, 1600 m alt. (Herzog 1919:296); Arpăşel Valley, 1000 m alt., sub S. dentata var.heterophylla (Pócs 1958:108); Făgăraş Mountains, det. Vajda L., sub subsp. dentata (Pócs 1958:109);Arpaş Valley, leg. & det. Vajda L., Călţun Lake, 2350 m alt., leg. & det. Pócs T. (Boros & Vajda1967:223,224); Sărata Glacial Ring, right side <strong>of</strong> Sărata Valley, peatbog, 1750 m alt., det. Ştefureac T.(Diaconeasa 1969:317; Drăgulescu 2003:16).Iezer-Păpuşa Mountains, Păpău Sheepfold, 1600 m alt., incl. sub S. dentata (Loitlesberger1898:193,194); Iezer Valley, Bătrâna Valley, Râuşor Valley (Mohan 1990a:24); below Iezer Refuge,7.08.1993, Iezer Valley, 19.08.95, near the springs <strong>and</strong> alpine brooks, ass. Chrysosplenio-Saxifragetumstellaris Pawl. & Walas 1949 (Alexiu 1995:40, Tab.2; Alexiu 1996:184).Cozia Mountain, north-eastern side <strong>of</strong> Jgheabul Bulzului, on soil, 1975-1976 (Dihoru G. 1990:53);Slavei Brook, on soil, sub var. dentata (Dihoru G. 1990:53).Cindrel Mountains, between Rozdeşti <strong>and</strong> Onceşti (Herzog 1919:296); Beşineu, incl. sub fo.elongata, Păltiniş, Râul Mare Valley, leg. & det. Vajda L., sub var. dentata (Boros & Vajda 1967:223,224;Gündisch 1977:51); Crinţ, 900 m alt., Păltiniş, 1500 m alt., Cânaia, 1700 m alt., Săroi, 1200 m alt.,Şerbănei, 1800 m alt., Bătrina, 1870 m alt., Beşineu, 1900 m alt., Jujbea, 2100 m alt. (Gündisch 1977:51;Drăgulescu 2003:16); Cindrel, 2200 m alt. (Gündisch 1977:51; Drăgulescu 1992:55; Drăgulescu 2003:16);Frumoasa - Platoul Diavolului [SIB HG](Drăgulescu 1992:55; Drăgulescu 2003:16).Sebeşului Mountains, Sebeş Valley, between Oaşa <strong>and</strong> Ciban, in common spruce forest, 800-1200m alt., sub S. dentata (Borza 1959:69,283, Tab.38; Drăgulescu 2003:16).Lotrului Mountains, the peatbog from Sărăcinul Mic, on left side <strong>of</strong> Lotru Valley, near PuruForest Hut, Voineasa Commune, 1956 (Ştefureac, Popescu & Lungu 1957b:872).Căpăţânei Mountains, Lotru Valley, Bucureasa, Malaia, 9-15.09.1952, sub S. dentata (Ştefureac,Popescu & Lungu 1955:527).Parâng Mountains, the springs <strong>of</strong> Mânileasa Brook, 1200 m alt., 26.07.1955 (Ştefureac, Popescu& Lungu 1956b:1314); the edge <strong>of</strong> the peatbog near Câlcescu Lake, 11.08.1955 (Ştefureac 1959:757);Mândra Mountain, Jieţ Valley, 1810 m alt., leg. & det. Pócs T. (Boros & Vajda 1967:223); south to thepeatbog from edge <strong>of</strong> Câlcescu Lake, incl. sub var. aequatiformis (Ştefureac 1969a:103,104); east to thepeatbog on Câlcescu Lake border (Ştefureac 1969a:105).Retezat Mountains, Zănoaga (Simonkai 1872:67; Fuss 1878:657; Hazslinszky 1885:67); JudelePeak, 2200 m alt., 22.07.1914, leg. Péterfi M., sub S. dentata var. speciosa Nees [BRHE 9](Györffy &Péterfi M. 1916:37); Mlaştina de Sus on Zănoaga Valley, Zănoaga-Slăveiul marked path, 1870 m alt.,Mlaştina I between Slăvei <strong>and</strong> Muchia Ascuţită, 1990 m alt., incl. sub S. dentata, Mlaştina II between279
Slăvei <strong>and</strong> Muchia Ascuţită, 2000 m alt., 1971, ass. Philonotido-Saxifragetum stellaris Horv. 1949,Mlaştina Mică at west to Zănoaga Lake below Zănoaga Peak, 2000 m alt., Mlaştina III from JudeleGlacial Valley, 2050 m alt., 1971, ass. Calthaetum laetae Krajina 1933 (Boşcaiu, Plămadă & Péterfi Ş.1972:177-179, Tabs.1,2; S<strong>and</strong>a & Popescu A. 1992:172, Tab.1); Judele Brook, on wet stones, 2080 m alt.(Plămadă 1973b:101); Judele Glacial Valley, in water, on stones, the vales between Muchia Ascuţită <strong>and</strong>Slăvei Culm – Judele 2 Valley, 1990-2100 m alt., Judele 2 Glacial Valley, in peatbogs, 1990-2090 m alt.,incl. sub S. dentata (Plămadă 1973b:102); Rovine Brook, 2000 m alt., 5.09.1967, Cârligului Valley at“Casa alpinet”, 1900 m alt., 9.07.1968, Ştirbului Brook, 1950-2000 m alt., 12.07.1971, Judele Brook,2070-2090 m alt., 9.07.1971, ass. Philonotido-Hygrohypnetum dilatati Plămadă 1974, Stânişoara Valley,1700 m alt., Judele Valley between Slăvei <strong>and</strong> Muchia Ascuţită, 1900 m alt., ass. Scapanietum undulataeSchwick. 1944, Zlătuia Brook Valley, 1600 m alt., ass. Brachythecietum rivulare Herzog 1943, BârleiBrook, 1850 m alt., 4.09.1967, Şesele Brook, 1050 m alt., 21.06.1968, Judele Brook, 2070-2100 m alt.,9.07.1971, ass. Fontinaletum gracilis Plămadă 1974, near Lia Lake, 1950-2000 m alt., 6.06.1969, nearGemenea Lake, 2100 m alt., 7.07.1969, upstream Tăul Cârligului, 2000 m alt., 8.07.1969, StânişoaraValley below Lolaia Peak, 1860 m alt., 7.07.1970, incl. sub S. dentata, ass. Philonotido-Saxifragetumstellaris Horv. 1949 (Plămadă 1974b:98-103, Tabs.3,4,5; Plămadă 1976a; Plămadă 1976b:16,22; Plămadă1993:55, Tabs.5,6); Retezat, 800-1200 m alt., ass. Chrysosplenio-Cardaminetum amarae (Tx. 1937)Mass 1959, 1750-2050 m alt., ass. Philonotido-Saxifragetum stellaris Horv. 1949, 1750-2050 m alt.,Calthaetum laetae Krajina 1933 (Coldea 1993:40, Tab.7).Ţarcu Mountains, Muntele Mic, 1000-1800 m alt., 03-08.1943, leg. Boşcaiu N. (Ştefureac1948a:220).Caraş-Severin County, „Şapte Izvoare”, Herculane Spa, 200 m alt., 22.06.1945, sub S. dentata(Ştefureac 1947a:138,150).Poiana Ruscă Mountains, Luncani, Luncanilor Valley (Hazslinszky 1885:67).Arad County, Dulcele on Mare Valley (Simonkai 1893:361).Mureş County, Răchitiş Peak „Poiana”, 25.07.1995, ass. Chrysosplenio-Cardaminetum amaraeMass. 1959 (Oroian 1996:247, Tab.22).Alba County, Galda de Sus, Rimetea (Barth 1880:121); Cărpiniş (Péterfi M. 1908:277); GăinaMountain, 1000 m alt., Galbena Valley, 600 m alt., sub S. dentata (Péterfi M. 1908:277; Goia &Schumacker 2000:92; Goia 2001:74); below Cucurbăta Mare, 1600-1700 m alt. (Coldea 1996:4, Tab.3;Goia 2001:74); Cucurbăta Mare Peak, 4.08.1994, 1700-1720 m alt., ass. Fontinalietum antipyreticaeGretzer (Goia 1998:251, Tab.2); Cobleş Valley, on rotten wood (Goia 2001:74); Arieşul Mare Basin, onrotten wood (Goia & Schumacker 2002:87).Cluj County, Turzii Gorges (Barth 1880:121); Dobrin Mountain (Péterfi M. 1918:61); DobrinMountain, 14.04.1916, leg. & det. Győrffy I. & Péterfi M. (Györffy 1924b:38); Făget, leg. & det. PéterfiM. (Boros 1958:4,8); the peatbogs from spring <strong>and</strong> upper course <strong>of</strong> Someşul Rece, incl. Muntele Mare<strong>and</strong> Dobrin (Pop E. 1960:397); Someşul Rece Valley, leg. & det. Vajda L. (Boros & Vajda 1967:224);Gilău Mountains, common spruce forest (Plămadă 1985b:40).Bihor County, Bulz, Pietros, 500 m alt., Cristior, 400 m alt., Stâna de Vale, Buteasa, 1300 m alt.,incl. sub S. dentata (Péterfi M. 1908:277); Iadului Valley between Remetea <strong>and</strong> Bulz, 26.05.1915, leg. &det. Győrffy I. & Péterfi M., sub S. dentata (Györffy 1924b:38; Boros 1942a:33; Boros & Vajda1967:225); Apuseni Mountains (Ştefureac 1975b:313).Bihor Mountains, Padiş, 1250 m alt., 08.1999, sub fo. dentata (Jakab 2000b:70).Scapania verrucosa HeegSyn.:– Scapania parva Steph.– Scapania verrucosa fo. isoloba Müll. Frib.– Scapania verrucosa var. schiffneriana Müll. Frib.– Scapania verrufifera C. Massal.First citation for <strong>Romania</strong>: Loitlesberger 1898.Transylvania (Pax 1908:130; Frey & al. 2006:101).<strong>Romania</strong> (Stephani 1910:134; Szweykowski 1969:59,61, Figs.1,2; Ştefănuţ 2003b:239, Fig.7).Ecology: epilithic, acidophytic, hygrophytic, sciophytic; circumpolar; suboceanic-subalpine; Hch.Herbaria <strong>and</strong> Exsiccatae: H.A.T.R.Distribution in Europe: Fr, Ch, At, Es, It, Si, Hr, Ba, Ro, Bg, Ua, Ca.280
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SORIN ŞTEFĂNUŢThe Hornwort and L
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CONTENTINTRODUCTION ...............
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The beginning of the 20 th century
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asin of the Arieş river. Bryotaxon
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Abbreviations used in the text† -
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The presentation of the methodology
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Fig. 3. The GEOCOD mapping system.
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THE HORNWORT AND LIVERWORT SPECIES
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THE HORNWORT AND LIVERWORT CLASSIFI
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Suborder Lepidoziineae R.M. Schust.
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46. Cephalozia pleniceps (Austin) L
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147. Pallavicinia lyellii (Hook.) C
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13 Corsinia coriandrina (Spreng.) L
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11. Porella obtusata (Taylor) Trevi
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34Phaeoceros carolinianus (Michx.)
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Făgăraş Mountains, Cârţişoara
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Cluj County, Someşul Rece Valley a
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Parâng Mountains, Repedea Valley,
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Argeş County, Rucăr (Loitlesberge
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Cindrel Mountains, Cindrel, 2200 m
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Parâng Mountains, Latoriţa Valley
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Athalamia hyalina (Sommerf.) S. Hat
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Barbilophozia barbata (Schmidel ex
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Bârgău Mountains, Cârţibavu Mar
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First citation for Romania: Schur 1
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- Pleuroschisma triangulare (Schlei
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and Băiuţ, common spruce forest,
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Distribution in Europe: Sj, Is, Fo,
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Poiana Stampei Peatbog, SV, 47°20'
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alt., 8.08.2003, leg. Pop O., det.
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leg. & det. Ştefureac T., ass. Buc
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Rodna Mountains, Puzdra Mountain, 1
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Alba County, Vârciorog Valley, 780
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Bârgău Mountains, Cucureasa Reser
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Suceava County, Tinovul Mare de la
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Suceava County, Rodna Nouă, on Per
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2001:74,75); Arieşul Mare Basin, o
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Ţibleş Mountains, Mesteacănul Va
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Cindrel Mountains, Găuşoara, leg.
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Parâng Mountains, the edge of Câl
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& det. Ştefănuţ S. [BUCA B3684];
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Alba County, at 20,5 km to Zlatna,
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Cephaloziella spinigera (Lindb.) Wa
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First citation for Romania: Péterf
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Iezer-Păpuşa Mountains, Iezer Val
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- Cladopodiella fluitans var. gigan
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Caraş-Severin County, Cerna Valley
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(Mititelu, Barabaş N. & al. 1968:1
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1885:30; Ştefureac 1949a:287,288;
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104Conocephalum salebrosum Szweyk.,
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Retezat Mountains, Retezat, sub Jun
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Cindrel Mountains, Cânaia, 1750 m
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Iaşi County, Sineşti Forest (Papp
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[BUCA B2793], on rotten beech trunk
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Orthodicrano montani-Hypnetum filif
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Distribution in Romania:Maramureş
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- Gymnocolea inflata var. laxa (Nee
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Ciucaş Mountains, Caprei Brook Val
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Giumalău Mountain, Giumalău Secul
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- Haplozia riparia var. rivularis B
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- Jungermannia karl-muelleri auct.,
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Jungermannia gracillima Sm.Syn.:- A
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Mountain, PH, 45°24'40"N / 25°28'
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Retezat Mountains, Rovine Brook, on
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Romania, alt. 600-2280 m (Váňa 19
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- Solenostoma subellipticum (Lindb.
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Distribution in Romania:Bârgău Mo
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13.07.1946, sub Lophozia alpestris
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Distribution in Romania:Maramureş
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Rodna Mountains, Lala Valley, 1200
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Ştefănuţ S. [BUCA B2574, B2580,
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Dealul Colbului and Pietrele Doamne
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150var. rivularis (Raddi) Warnst.Sy
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Penteleu Massif, fer - Roşu Lake,
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Satu Mare County, Noroieni, Ghiarma
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Lophozia ascendens (Warnst.) R.M. S
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- Jungermannia intermedia Lindenb.-
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Căliman Mountains, on humus and ro
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Hăşmaş Mountains, Hăşmaş, sub
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Herbaria and Exsiccate: BUCADistrib
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peatbog on the border of Câlcescu
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? - Iezer-Păpuşa Mountains, Iezer
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Făgăraş Mountains, s - above Avr
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Nemira Mountains, on edge of Nemira
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2003:7); Măgura Cisnădiei [SIB HL
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Fuss 1854:20; Ştefureac 1949a:288;
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- Nardia brevissima (Dumort.) Lindb
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1919:295); Ciortea Mountain, on wet
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- Sarcoscyphus sphacelatus var. med
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Siriu Mountain, Poiana din Fundul M
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Iaşi County, below Repedea, 7.10.1
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Valley, Arieşul Mic - upstream min
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Alba County, Valea Lungă, 10.02.18
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Rarău Massif, between Dealul Colbu
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Harghita County, Harghita Spa, on r
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Căliman Mountains, in alpine zone,
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Stânişoara Mountains, Suha Mare B
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Ecology: humicolous, acidophytic, h
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Bacău County, Slănic Brook (Papp
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Rarău Mountain, Codrul Secular Sl
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1915:197,198); Cândeşti, 07.1891,
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Ţibleş Mountains, Piciorul Ţigle
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Ceahlău Mountain, Durău Valley, i
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major (Ştefureac 1962b:69); Benea-
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Ciucaş Mountains, Berii Brook Vall
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Zlătuia Valley, 1100 m alt., 7.09.
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- Plagiochila dillenii (Taylor) Tay
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Wallfisch R. (Dihoru G. & al. 2003:
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Ştefănuţ M. 2003:79); Iezerul Pi
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utabuli-Mnietum cuspidati (Felföld
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Iaşi County, Mârzeşti Forest, Bu
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- Page 237 and 238: Rodna Mountains (Pax 1908:130); Pie
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- Page 241 and 242: Bârgău Mountains, Cucureasa Reser
- Page 243 and 244: B3521, B3522] (Ştefănuţ 2006b:34
- Page 245 and 246: 16,20.08.1994, ass. Pterigynandretu
- Page 247 and 248: - Marchantia crinita Michx.- Marcha
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- Page 265 and 266: 1960:1113); upper side of Mălăie
- Page 267 and 268: Scapania brevicaulis TaylorSyn.:- S
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- Page 273 and 274: - Scapania amurensis Warnst.- Scapa
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- Page 279: - Scapania heterophylla M. Howe- Sc
- Page 283 and 284: Distribution in Europe: Ie, Br, Fr,
- Page 285 and 286: Nemira Mountains, Nemira, on humus,
- Page 287 and 288: - Sphenolobus medelpadicus (Arnell)
- Page 289 and 290: Parâng Mountains, Latoriţa Valley
- Page 291 and 292: Fig. 9. Distribution of Phaeoceros
- Page 293 and 294: Fig. 13. Distribution of Anastrophy
- Page 295 and 296: Fig. 17. Distribution of Anthelia j
- Page 297 and 298: Fig. 21. Distribution of Asterella
- Page 299 and 300: Fig. 25. Distribution of Barbilopho
- Page 301 and 302: Fig. 29. Distribution of Barbilopho
- Page 303 and 304: Fig. 33. Distribution of Bazzania t
- Page 305 and 306: Fig. 37. Distribution of Calypogeia
- Page 307 and 308: Fig. 41. Distribution of Calypogeia
- Page 309 and 310: Fig. 45. Distribution of Cephalozia
- Page 311 and 312: Fig. 49. Distribution of Cephalozia
- Page 313 and 314: Fig. 53. Distribution of Cephalozie
- Page 315 and 316: Fig. 57. Distribution of Cephalozie
- Page 317 and 318: Fig. 61. Distribution of Cephalozie
- Page 319 and 320: Fig. 65. Distribution of Cololejeun
- Page 321 and 322: Fig. 69. Distribution of Corsinia c
- Page 323 and 324: Fig. 73. Distribution of Fossombron
- Page 325 and 326: Fig. 77. Distribution of Frullania
- Page 327 and 328: Fig. 81. Distribution of Gymnocolea
- Page 329 and 330: Fig. 85. Distribution of Harpanthus
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Fig. 89. Distribution of Jungermann
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Fig. 93. Distribution of Jungermann
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Fig. 97. Distribution of Jungermann
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Fig. 101. Distribution of Jungerman
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Fig. 105. Distribution of Leiocolea
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Fig. 109. Distribution of Lejeunea
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Fig. 113. Distribution of Lophocole
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Fig. 117. Distribution of Lophozia
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Fig. 121. Distribution of Lophozia
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Fig. 125. Distribution of Lunularia
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Fig. 129. Distribution of Marchanti
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Fig. 133. Distribution of Marsupell
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Fig. 137. Distribution of Marsupell
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Fig. 141. Distribution of Moerckia
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Fig. 145. Distribution of Nardia ge
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Fig. 149. Distribution of Odontosch
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Fig. 153. Distribution of Pedinophy
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Fig. 157. Distribution of Peltolepi
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Fig. 161. Distribution of Porella a
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Fig. 165. Distribution of Preissia
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Fig. 169. Distribution of Radula li
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Fig. 173. Distribution of Riccardia
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Fig. 177. Distribution of Riccia ca
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Fig. 181. Distribution of Riccia cr
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Fig. 185. Distribution of Riccia go
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Fig. 189. Distribution of Ricciocar
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Fig. 193. Distribution of Scapania
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Fig. 197. Distribution of Scapania
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Fig. 201. Distribution of Scapania
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Fig. 205. Distribution of Scapania
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Fig. 209. Distribution of Scapania
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Fig. 213. Distribution of Scapania
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Fig. 217. Distribution of Tritomari
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Evaluating the distribution dataFol
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Fig. 224. Hornworts and Liverworts
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Signs used:Bolded names - species i
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Marchantiaceae50. VU - Bucegia roma
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Fig. 228. The distribution of zones
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Anthoceros communis Steph. = Phaeoc
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Blasia funckii Corda = Blasia pusil
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Cephalozia crassiflora Spruce = Cep
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Cephaloziella spinigera (Lindb.) Wa
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Diplomitrium lyellii (Hook.) Corda
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Gymnocolea inflata subsp. acutiloba
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Jungermannia acuta var. gypsacea Go
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Jungermannia dentata Raddi = Cephal
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Jungermannia limprichtii Lindb. = L
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Jungermannia scalaris var. denudata
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Leiocolea heterocolpos (Thed. ex C.
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Lophozia incisa var. inermis Müll.
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Marchantia polymorpha var. dilatata
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Mastigobryum deflexum (Mart.) Ångs
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Odontoschisma sphagni var. denudatu
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Porella platyphylla (L.) Pfeiff.Por
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Riccia maior Roth = Riccia glauca L
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Scapania heterophylla M. Howe = Sca
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Sphagnocetis communis var. macrior
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Mittheilungen des siebenbürgischen
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1059. Péterfi L.Ş., 1963. Flagela
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1178. Pop I., Hodişan I., 1958. Co
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1231. Prunescu-Arion Elena, Toniuc
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1342. Rüegsegger F., 1986. Frullan
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1396. Schiffner V., 1913b. Über ei
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1567. Ştefureac T., 1960b. Cercet
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