Parâng Mountains, Curmătura Vidruţei, 1350 m alt., 26.07.1955, sub Haplozia (Ştefureac,Popescu & Lungu 1957a:68,69); Latoriţa Valley, Repedea Valley, 08-09.1954 (Ştefureac, Popescu &Lungu 1959:9,20, Tab.5); south to the peatbog from the edge <strong>of</strong> Câlcescu Lake (Ştefureac 1969a:104).Retezat Mountains, Râu de Mori (Schiffner 1914:308); Râul Mare Valley between Gura Zlata <strong>and</strong>Gura Apei (Páll 1964a:108); Retezat National Park, Leucodonto-Madothecietum platyphyllae,Anomodontetum attenuati (Barkm. 1939) Peciar 1965, Pterygyn<strong>and</strong>ro-Isothecietum myuri Plămadă(1976) 1978, Neckeretum crispae Ştefur. 1941 (Plămadă 1976a; Plămadă 1976b:20, Tab.2); betweenZlătuia Valley <strong>and</strong> Turcului Valley, 1100-1200 m alt., 7.09.1969, ass. Leucodonto-Madothecetumplatyphyllae Plămadă 1982, Zlătuia Valley, 790-800 m alt., 6-7.09.1969, Şesele Valley, 900-950 m alt.,9.09.1969, ass. Anomodontetum attenuati, Zlătuia Valley, 1000 m alt., 7.09.1969, Turcului Valley, 850 malt., 9.09.1969, Şesele Valley, 950 m alt., 14.07.1970, ass. Pterigyn<strong>and</strong>ro-Isothecietum myuri, Şesele -Pârgului Valley, 950 m alt., 9.09.1969, 1000 m alt., 14.07.1970, ass. Anomodonto attenuati-Neckeretumcrispae Plămadă 1982 (Plămadă 1982a:305-307,309, Tabs.2-5; Plămadă 1993:51, Tab.1); GemeneleValley, 08.2005, leg. Onete Marilena, det. Ştefănuţ S. [BUCA B3204].Mehedinţi County, Orşova (Hazslinszky 1885:41); Orşova, Cazane (Schiffner 1914:308); <strong>The</strong>Culm towards Poiana Salcuţei, Slătinic Valley, Gura Văii, 28.06.1945 (Ştefureac 1947a:138); †, s - Ada-Kaleh Isle, on Populus alba (Ştefureac & Mihai 1968a:298,305, Tab.1); Eşelniţa Valley, frequent, onQuercus <strong>and</strong> Acer (Plămadă 1970b:206); Danube Defile, between Orşova <strong>and</strong> Cerna Valley, in forest, onbark, frequent, 1966-1968 (Plămadă 1970c:393); Tricule, the first valley towards Canton, on stones,between Berzasca <strong>and</strong> Sviniţa, 1966, sub fo. propagulifera (Ştefureac & Mihai 1970:500).Domogled Mountain, Domogled (Degen 1901:14); ♀♂ - Crucea Albă, 500 m alt., 9.08.2004, leg.& det. Ştefănuţ S. [BUCA B2982].Caraş Severin County, Cerna Valley, Ciorici Peak (Boros, Vajda & Debreczy 1969:6,7).Poiana Ruscă Mountains, Luncani (Simonkai 1872:67; Fuss 1878:653); Poiana Ruscă, RuscaMontană, Pleşu, on hornbeam bark, 630 m alt. (Papp 1942a:51); Sinersig „Dumbrava” Forest, 29.07.1962(Ştefureac & Mihai 1967:14).Zar<strong>and</strong> Mountains, Highiş Peak, beech forest, Patru Gomile Valley, Slatina de Mureş, on bark(Csűrös M. 1978:77; Ardelean 2006:105).Codru-Moma Mountains, Monesei Valley, Osoiu Mare, Valea Lungă, Ursului Hill, BrihenilorValley, on beech <strong>and</strong> oak bark (Mohan 1996:49; Ardelean 2006:105).Arad County, Ciala Forest, Pecica, from Adea towards Sebiş along “Tőzmellék” (Simonkai1893:361; Fuss 1878:653); Ciala Forest, on oak, 3.09.2003, leg. & det. Ştefănuţ S. [BUCA B2332,B2340]; ♂♀ - Runcu-Groşi Reserve, 46°10'14"N / 22°07'34"E, 428 m alt., 20.08.2008, leg. & det.Ştefănuţ S. [BUCA B3926].Mureş County, Sighişoara, sub Jungermannia (Baumgarten 1846:229; Schur 1866:874; Fuss1878:653; Györffy 1917a:167); Lechinţa, oak forest, on bark, leg. Borza A., det. Papp C., incl. sub var.propagulifera (Papp 1944a:99); Ilva Valley, 9.08.1995 (Oroian 1997:113, Tab.1).Alba County, Piatra Ceţii, Trascău Mountains, 3.07.1868 (Barth 1868:140,141); Galda de Sus, leg.& det. Barth J. (Fuss 1878:653); Valea Lungă, on bark, 1866, leg. & det. Barth J. [SIB HF 1556, 1557](Fuss 1878:653; Borza & Lupşa 1964:151; Drăgulescu 1983:59); fer - Vidra, „Piatra Struţu”<strong>and</strong>Scărişoara, Piatra Runcului (Schiffner 1914:308; Goia 2001:82); Feneş Gorges, sub fo. propagulifera(Papp 1944b:236; Papp 1945a:162); the region between Geoagiu-Mureş-Ampoi-Feneş <strong>and</strong> Detunata Peakfrom Trascău Mountain, frequent (Plămadă 1986a:58); Pojar Hill, on hornbeam, 580 m alt., 25.08.1994,ass. Isothecietum myuri Hilitzer 1925, ass. Madotheco-Leskeeletum nervosae, Drăghiţa Valley, on fallenbeech tree, 750 m alt., 20.08.94, ass. Anomodonto-Leucodontetum Wisn. 1930 (Goia 1998:256-258,Tabs.6-8); Măgura Valley, 1300 m alt., ass. Pterygyn<strong>and</strong>retum filiformis Hilitzer 1925, Drăghiţa Valley,750 m alt., 28.08.1994, ass. Anomodonto-Leucodontetum (Goia 2000:120, Tab.8); Arieşul Mic Basin(Goia & Schumacker 2000:92); Popeştilor Brook, on willow, below Lucia Cave, on willow, Pojar Hill, onhornbeam, Morii Valley, on common spruce, beech <strong>and</strong> rotten wood, Ursoaia Saddle, common spruce,Cobleş Valley, Poiana Vadului, on ash tree, Divaia Valley, Popeştilor Valley, Vidrişoara Valley, ArieşulMic towards springs, Măgura Valley, Gârda – Scărişoara interfluve, Ordâncuşa Valley, Scărişoara Cave –aven, on beech tree, Galbena Valley, Drăghiţa Valley, Pătroieştilor Valley, on beech forest <strong>and</strong> on rottenwood, Vârciorog Valley <strong>and</strong> Ţoha Mică Valley, on rotten wood (Goia 2001:83); Popeştilor Valley,15.08.1994, Drăghiţa Valley, 20.08.1994, ass. Pylaisietum polyanthae Felf. 1941, Pojar Hill, 25.08.1994,Vidrişoara Valley, 20.08.1992, Cheia Valley, 31.07.1994, Morii Valley, 13.08.1994, Arieşul Mic Valley,243
16,20.08.1994, ass. Pterigyn<strong>and</strong>retum filiformis Hilitzer 1925, Drăghiţa Valley, 20.08.1994, MoriiValley, 7.08.1994, ass. Homalothecium sericei-Porelletum platyphyllae Størmer 1938, PătroieştilorValley, 22.08.1992, Morii Valley, 13.08.1994, Divaia Valley, 21.08.1994, ass. Anomodontetum attenuatiCain et Sharp 1938, Arieşul Mic Valley, 20.08.1994, Drăghiţa Valley, 20.08.1994, Pojar Hill, 25.08.1994,ass. Isothecietum myuri Hilitzer 1925, Pojar Valley, 25.08.1994, Drăghiţa Valley, 20.08.1996, ass.Anomodonto viticulosi-Leucodontetum sciuroidis Wisn. 1930, Pojar Hill, 25.08.1994, PătroieştilorValley, 22.08.1992, ass. Orthodicrano montani-Hypnetum filiformis Wisn. 1930 (Goia & Schumacker2003:59,60,62-64, Tabs.1-5); Arieşul Mare Basin, on rotten wood (Goia & Schumacker 2002:87);Vramiţă Valley, towards Avram Iancu, 27.05.1998 (Smets 2003:60); Colbeş Valley, 860 m alt.,10.08.1996, ass. Pylaisietum polyantae Felf. 1941 orthotrichetosum obtusifoliae Marst. 1985, GârdaSeacă Valley, 1000 m alt., 8.08.1996, Scărişoara cave, 1070 m alt., 11.08.1996, Ordâncuşa Valley, 1180m alt., 11.08.1996, Galbena Valley, 980 m alt., 12.09.1996, between Măgura <strong>and</strong> Obârşia Valley, 980 malt., 27.09.1996, between Gârda Valley <strong>and</strong> Scărişoara cave, 1000 m alt., 11.08.1996, ass.Pterigyn<strong>and</strong>retum filiformis, Scărişoara cave, 1080 m alt., 11.08.1996, ass. Homalothecio sericei-Porelletum platyphyllae, Ordâncuşa Valley, 1060 m alt., 11.08.1996, between Gârda Valley <strong>and</strong>Scărişoara Cave, 1000 m alt., 11.08.1996, ass. Isothecietum myuri, between Măgura Valley <strong>and</strong> ObârşiaValley, 700 m alt., 27.09.1996, ass. Anomodonto viticulosi-Leucodontetum sciuroides, between Mătişeşti<strong>and</strong> Şaua Ursoaia, 730 m alt., ass. Orthodicrano montani-Hypnetum filiformis (Goia & Schumacker 2004:107,109-111, Tabs.1-4).Cluj County, Southern Ciucea on Surduc Valley (Boros 1942a:34); Turda, Băile Sărate (Todor1947:41); Făget, Greben Valley, Mănăştur (Soó 1951:40; Boros 1958:8); Cluj-Napoca Botanical Garden,sub fo. propagulifera (Boros 1958:5,8; Plămadă 1963:179); Hoia, Pleşca Valley, Cluj-Napoca, LombForest (Boros 1958:8); Hoia Forest, north-west to Cluj, on bark, 1961-1963 (Plămadă 1966:60); Cluj-Napoca Botanical Garden, on bark, 1961-1966, incl. sub fo. propagulifera (Plămadă 1967:470; Ştefureac1982c:197,200, Tab.2); Mera Forest Basin, on bark, 1963-1965, sub fo. propagulifera (Plămadă1968:25); Cluj-Napoca Botanical Garden, cemetery, Sporting Park, on bark (Plămadă 1969:170); HuzaForest on Filii Mountain, Muntele Mare, near the Hut, on bark, 700-750 m alt., in Leucodontea (Plămadă& Coldea 1989b:121); Turda’s Gorges, on Crataegus (Plămadă & Goia 1994:87); Sălicea, Făget-St. Ion,Valea Popii, Cluj-Napoca, Central Park (Goia 2002:115, Tab.1).Bihor County, Felix Spa (Simonkai 1876; Borbas 1879:262); Drăgan Valley (Boros 1942a:35);Rădvani Forest, on oak bark, det. Ştefureac T. (Pop I. 1968:70).Bihor Mountains, without locality (Péterfi M. 1908:278; Goia 2001:83); Someşul Cald Gorges(Goia & Mătase 2001:17,18); ♀♂ - Cetăţile Ponorului, AB, 46°33'55"N / 22°42'15"E, 1050 m alt.,3.09.2007, leg. & det. Ştefănuţ S. [BUCA B3840].Radula lindenbergiana Gottsche et C. Hartm.Syn.:– Radula complanata subsp. lindenbergiana (Gottsche ex C. Hartm.) R.M. Schust.– Radula bornmuelleri Schiffn.– Radula commutata Gottsche ex Husn.– Radula complanata var. propagulifera (Nees) Gottsche et al.– Radula constricta Steph.– Radula germana J.B. Jack– Radula lindbergiana Gottsche ex J.B. Jack– Radula lindenbergiana var. germana (J.B. Jack) C. Massal.– Radula ovata J.B. Jack ex Steph.First citation for <strong>Romania</strong>: Loitlesberger 1898.<strong>Romania</strong> (Schuster 1980:609).Moldova (Papp 1943:666; Ştefureac & al. 1973:149).Muntenia (Mohan 1988b:110).Transylvania, sub R. lindenbergii (Pax 1908:130).Ecology: epilithic, acidophytic, xero-mesophytic, sciophytic; Eurasia; mountain-alpine; Hch.Herbaria <strong>and</strong> Exsiccate: BUCADistribution in Europe: Is, Fo, No, Se, Fi, Ie, Br, Fr, De, Po, Ch, At, Cz, Sk, Hu, Pt, Es, Bl, Co, Sa, Sc, It, Si, Hr, Me, Ba, Ro,Bg, Al, Gr, Ct, Tu, Lv, Lt, Ua, Rn, Ca, Az, Ma, Cn.Distribution in <strong>Romania</strong>:Maramureş County, Covatar Waterfall from Săpânţa, 900 m alt., 15-21.06.1997, leg. & det. JakabG. (Jakab 1999:53,55).244
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SORIN ŞTEFĂNUŢThe Hornwort and L
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CONTENTINTRODUCTION ...............
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The beginning of the 20 th century
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asin of the Arieş river. Bryotaxon
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Abbreviations used in the text† -
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The presentation of the methodology
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Fig. 3. The GEOCOD mapping system.
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THE HORNWORT AND LIVERWORT SPECIES
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THE HORNWORT AND LIVERWORT CLASSIFI
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Suborder Lepidoziineae R.M. Schust.
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46. Cephalozia pleniceps (Austin) L
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147. Pallavicinia lyellii (Hook.) C
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13 Corsinia coriandrina (Spreng.) L
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11. Porella obtusata (Taylor) Trevi
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34Phaeoceros carolinianus (Michx.)
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Făgăraş Mountains, Cârţişoara
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Cluj County, Someşul Rece Valley a
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Parâng Mountains, Repedea Valley,
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Argeş County, Rucăr (Loitlesberge
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Cindrel Mountains, Cindrel, 2200 m
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Parâng Mountains, Latoriţa Valley
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Athalamia hyalina (Sommerf.) S. Hat
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Barbilophozia barbata (Schmidel ex
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Bârgău Mountains, Cârţibavu Mar
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First citation for Romania: Schur 1
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- Pleuroschisma triangulare (Schlei
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and Băiuţ, common spruce forest,
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Distribution in Europe: Sj, Is, Fo,
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Poiana Stampei Peatbog, SV, 47°20'
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alt., 8.08.2003, leg. Pop O., det.
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leg. & det. Ştefureac T., ass. Buc
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Rodna Mountains, Puzdra Mountain, 1
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Alba County, Vârciorog Valley, 780
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Bârgău Mountains, Cucureasa Reser
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Suceava County, Tinovul Mare de la
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Suceava County, Rodna Nouă, on Per
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2001:74,75); Arieşul Mare Basin, o
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Ţibleş Mountains, Mesteacănul Va
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Cindrel Mountains, Găuşoara, leg.
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Parâng Mountains, the edge of Câl
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& det. Ştefănuţ S. [BUCA B3684];
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Alba County, at 20,5 km to Zlatna,
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Cephaloziella spinigera (Lindb.) Wa
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First citation for Romania: Péterf
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Iezer-Păpuşa Mountains, Iezer Val
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- Cladopodiella fluitans var. gigan
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Caraş-Severin County, Cerna Valley
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(Mititelu, Barabaş N. & al. 1968:1
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1885:30; Ştefureac 1949a:287,288;
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104Conocephalum salebrosum Szweyk.,
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Retezat Mountains, Retezat, sub Jun
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Cindrel Mountains, Cânaia, 1750 m
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Iaşi County, Sineşti Forest (Papp
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[BUCA B2793], on rotten beech trunk
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Orthodicrano montani-Hypnetum filif
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Distribution in Romania:Maramureş
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- Gymnocolea inflata var. laxa (Nee
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Ciucaş Mountains, Caprei Brook Val
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Giumalău Mountain, Giumalău Secul
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- Haplozia riparia var. rivularis B
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- Jungermannia karl-muelleri auct.,
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Jungermannia gracillima Sm.Syn.:- A
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Mountain, PH, 45°24'40"N / 25°28'
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Retezat Mountains, Rovine Brook, on
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Romania, alt. 600-2280 m (Váňa 19
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- Solenostoma subellipticum (Lindb.
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Distribution in Romania:Bârgău Mo
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13.07.1946, sub Lophozia alpestris
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Distribution in Romania:Maramureş
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Rodna Mountains, Lala Valley, 1200
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Ştefănuţ S. [BUCA B2574, B2580,
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Dealul Colbului and Pietrele Doamne
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150var. rivularis (Raddi) Warnst.Sy
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Penteleu Massif, fer - Roşu Lake,
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Satu Mare County, Noroieni, Ghiarma
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Lophozia ascendens (Warnst.) R.M. S
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- Jungermannia intermedia Lindenb.-
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Căliman Mountains, on humus and ro
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Hăşmaş Mountains, Hăşmaş, sub
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Herbaria and Exsiccate: BUCADistrib
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peatbog on the border of Câlcescu
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? - Iezer-Păpuşa Mountains, Iezer
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Făgăraş Mountains, s - above Avr
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Nemira Mountains, on edge of Nemira
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2003:7); Măgura Cisnădiei [SIB HL
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Fuss 1854:20; Ştefureac 1949a:288;
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- Nardia brevissima (Dumort.) Lindb
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1919:295); Ciortea Mountain, on wet
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- Sarcoscyphus sphacelatus var. med
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Siriu Mountain, Poiana din Fundul M
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Iaşi County, below Repedea, 7.10.1
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Valley, Arieşul Mic - upstream min
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Alba County, Valea Lungă, 10.02.18
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Fig. 17. Distribution of Anthelia j
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Fig. 21. Distribution of Asterella
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Fig. 25. Distribution of Barbilopho
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Fig. 29. Distribution of Barbilopho
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Fig. 33. Distribution of Bazzania t
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Fig. 37. Distribution of Calypogeia
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Fig. 41. Distribution of Calypogeia
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Fig. 45. Distribution of Cephalozia
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Fig. 49. Distribution of Cephalozia
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Fig. 53. Distribution of Cephalozie
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Fig. 57. Distribution of Cephalozie
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Fig. 61. Distribution of Cephalozie
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Fig. 65. Distribution of Cololejeun
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Fig. 69. Distribution of Corsinia c
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Fig. 73. Distribution of Fossombron
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Fig. 77. Distribution of Frullania
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Fig. 81. Distribution of Gymnocolea
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Fig. 85. Distribution of Harpanthus
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Fig. 89. Distribution of Jungermann
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Fig. 93. Distribution of Jungermann
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Fig. 97. Distribution of Jungermann
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Fig. 101. Distribution of Jungerman
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Fig. 105. Distribution of Leiocolea
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Fig. 109. Distribution of Lejeunea
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Fig. 113. Distribution of Lophocole
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Fig. 117. Distribution of Lophozia
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Fig. 121. Distribution of Lophozia
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Fig. 125. Distribution of Lunularia
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Fig. 129. Distribution of Marchanti
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Fig. 133. Distribution of Marsupell
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Fig. 137. Distribution of Marsupell
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Fig. 141. Distribution of Moerckia
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Fig. 145. Distribution of Nardia ge
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Fig. 149. Distribution of Odontosch
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Fig. 153. Distribution of Pedinophy
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Fig. 157. Distribution of Peltolepi
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Fig. 161. Distribution of Porella a
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Fig. 165. Distribution of Preissia
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Fig. 169. Distribution of Radula li
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Fig. 173. Distribution of Riccardia
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Fig. 177. Distribution of Riccia ca
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Fig. 181. Distribution of Riccia cr
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Fig. 185. Distribution of Riccia go
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Fig. 189. Distribution of Ricciocar
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Fig. 193. Distribution of Scapania
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Fig. 197. Distribution of Scapania
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Fig. 201. Distribution of Scapania
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Fig. 205. Distribution of Scapania
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Fig. 209. Distribution of Scapania
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Fig. 213. Distribution of Scapania
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Fig. 217. Distribution of Tritomari
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Evaluating the distribution dataFol
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Fig. 224. Hornworts and Liverworts
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Signs used:Bolded names - species i
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Marchantiaceae50. VU - Bucegia roma
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Fig. 228. The distribution of zones
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Anthoceros communis Steph. = Phaeoc
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Blasia funckii Corda = Blasia pusil
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Cephalozia crassiflora Spruce = Cep
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Cephaloziella spinigera (Lindb.) Wa
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Diplomitrium lyellii (Hook.) Corda
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Gymnocolea inflata subsp. acutiloba
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Jungermannia acuta var. gypsacea Go
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Jungermannia dentata Raddi = Cephal
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Jungermannia limprichtii Lindb. = L
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Jungermannia scalaris var. denudata
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Leiocolea heterocolpos (Thed. ex C.
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Lophozia incisa var. inermis Müll.
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Marchantia polymorpha var. dilatata
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Mastigobryum deflexum (Mart.) Ångs
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Odontoschisma sphagni var. denudatu
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Porella platyphylla (L.) Pfeiff.Por
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Riccia maior Roth = Riccia glauca L
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Scapania heterophylla M. Howe = Sca
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Sphagnocetis communis var. macrior
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Mittheilungen des siebenbürgischen
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91. Borza A., 1922a. Schedae ad "Fl
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142. Boşcaiu N., Täuber F., Colde
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199. Coldea G., 1975. Étude phytos
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256. Csűrös Ş., Moldovan I., Cs
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317. Dihoru G., 1987b. Contribuţie
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367. Domin K., 1923. Grundzüge der
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487. Györffy I., 1906c. Polytrichu
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540. Györffy I., 1943b. Erdélyi P
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598. Ionescu Venera, Cristurean I.,
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658. Lungu Lucia, 1967a. Contribuţ
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712. Mihai G., 1971e. Aspecte ale b
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824. Mohan G., 1988c. Die Gattung T
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941. Papp C., 1931b. O schiţă a v
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999. Papp C., Eftimie Elena, 1963.
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1059. Péterfi L.Ş., 1963. Flagela
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1119. Plămadă, Viţalariu, 1971.
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1178. Pop I., Hodişan I., 1958. Co
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1231. Prunescu-Arion Elena, Toniuc
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1285. Raţiu O., 1970. Consideraţi
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1342. Rüegsegger F., 1986. Frullan
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1396. Schiffner V., 1913b. Über ei
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1455. Söderström L., Urmi E., Vá
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1515. Ştefănuţ S., 2004d. The Ho
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1567. Ştefureac T., 1960b. Cercet
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1624. Ştefureac T., 1976c. Rolul b
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1675. Ştefureac T., Cristurean I.,
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1727. Tarnavschi I., Nedelcu G., 19
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1785. Wallfisch R., 2008. Contribut
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var. fragilis (A. Roth) Müll. Frib
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Riccardia incurvata Lindb. 28, 248,