Nemira Mountains, Nemira Mountain, on soil, in common spruce forest, on Fagus sylvatica <strong>and</strong>Acer pseudoplatanus, 900-1400 m alt. (Barabaş V. 1974:26,29,30); Farcu Mare Peak, on Fagus sylvatica<strong>and</strong> Alnus incana trunks <strong>and</strong> wet rocks, 1970-1977 (Mihai & Barabaş V. 1978:117); Nemira Mountain,800-1600 m alt., Slănic <strong>and</strong> Uz Brooks Vales, 500 m alt., Bălătău Lake, 900 m alt. (Barabaş V.1982b:58,59,63,67,74,75, Tabs.7,11,17,25,26).Bacău County, Pufu Hill, in forest, 800-900 m alt., Porcului Hill, in forest, Ciungetul BrookValley, sub fo propagulifera (Papp 1957:285; Papp 1958b:396); Berzunţ Culm, 990 m alt., SavaHermitage, in forest, sub var. propagulifera (Papp & Bîrcă 1960:316,324); Tajbuga Culm, on bark, incl.sub fo. propagulifera (Papp 1966:219-221); Seaca Forest, Tocila (Mititelu, Barabaş N. & al.1968:147,168; Ştefureac & al. 1973:149; Costică 1995:252); Uz Valley, near the dam, on rotten stumps,1966-1967 (Mihai & Barabaş V. 1969:71,75); Podu Turcului, on Quercus robur (Mihai & Barabaş V.1970:67,70,71); s - Măgura Hill, north-west to Târgu Ocna, on Quercus petraea rotten stumps <strong>and</strong> rocks,680 m alt., rare, incl. sub fo. propagulifera, on stones (Mihai & Barabaş V. 1971:676,686); s, fer -Sohodol Forest, Măgura Commune, at base <strong>of</strong> Carpinus betulus tree, Traian Forest, on Quercus robur,Petricica Hill, Măgura Commune, on rotten beech trees, 1970-1971 (Mihai & Barabaş V. 1972:87,92;Costică 1995:252); Cheşcheş Culm, on hornbeam bark, 2.05-8.11.1971 (Ştefureac & Barabaş V.1972:78,82); Ghioina Forest, incl. sub var. propagulifera, Lada Forest, Slabului Hill, the forest way fromÎnţărcătoare towards Zboina Verde, on Populus tremula, Carpinus betulus, Fagus sylvatica <strong>and</strong> Acercampestre (Eftimie 1972:151); Codrul Secular Runc-Racova, on bark (Barabaş V. 1982a:49); MăguraHill – Târgu Ocna, 400 m alt. (Barabaş V. 1982b:77,78, Tab.30).Bodoc Mountains, on wood (Ştefureac & Kovács 1981:117).Vrancea County, Ploscuţeni Forest, 303 m alt., at base <strong>of</strong> beech trees (Mihai & Barabaş V.1970:67,71-73); Tişiţei Gorges, on bark, 1977 (Mihai & Pascal 1986:46).Brăila County, Brăila, on willow <strong>and</strong> poplar barks, 29.05.1969, leg. Moşneagă M. & det. MohanG. [PHHM 6207727](Mohan 1971a:9).Tulcea County, Dealul Mare, Niculiţel Commune, on bark (Papp 1938:289,293,295; Mihai, Pascal& Viţalaru 1962:285; Ştefureac, Mihai & Pascal 1970:123); Letea Forest, north to C.A. RosettiCommune, on bark (Mihai & Pascal 1963:264; Ştefureac, Mihai & Pascal 1970:123); Codru Forest,Babadag (Mihai, Pascal & Viţalaru 1962:285); Babadag Forest, 220 m alt., Ciucurova Topolog Forest,350 m alt., Niculiţel Forest, on bark, 20-23.07.1968 (Ştefureac & Mohan 1969:140,144; Ştefureac, Mihai& Pascal 1970:123); ♀♂ - Canaraua Fetii, on Ulmus bark, 6.06.2007, leg. & det. Ştefănuţ S. [BUCAB3993], s - [BUCA B3994]; Topolog, 44°51'55"N / 28°22'45"E, 13.06.2008, leg. & det. Ştefănuţ S.[BUCA B3886].Constanţa County, ? – near Valul lui Traian Commune, in forest. Obs. Quated according Mihai,Pascal & Viţalaru 1962, but missing in this paper (Ştefureac, Mihai & Pascal 1970:123); Făurei Forest,south-east to Băneasa Commune, on Acer tataricum <strong>and</strong> Quercus cerris (Mihai & Pascal 1963:264;Ştefureac, Mihai & Pascal 1970:123); Danube Bank between Galiţa <strong>and</strong> Ostrov, Canaraua Fetii Forest,1963 (Mihai 1965:160); Cărpiniş Forest, Băneasa Commune (Mihai 1965; Ştefureac, Mihai & Pascal1970:123); Negureni Forest, on Quercus cerris (Mihai 1981b:10); Canaraua Fetii <strong>and</strong> Esechioi ForestReserves (Mititelu, Parincu & Gheorghiţă 1993:33); Neptun railway station, 2.07.2005, leg. & det.Ştefănuţ S. [BUCA B3194]; ♀♂ - Comarova Forest near Neptun railway station, 2.07.2006, leg. & det.Ştefănuţ S. [BUCA B3537].Penteleu Massif, Tisa Reserve, on bark, 1000 m alt. (Dihoru G. 1964:399, Tab.3; Dihoru G.1987b:94); Şapte Izvoare Valley, above forest hut, common spruce forest, on rotten common sprucewood, 11.10.1983 (Dihoru G. 1987b:94).Buzău County, ♀♂ - Sărata-Monteoru, 07.2005, leg. Ivănescu Cristina, det. Ştefănuţ S. [BUCAB3187]; ♂♀ - Spătaru Forest, on Fraxinus pallisiae bark, 45°05'22"N / 26°47'03"E, 11.11.2008, leg. &det. Ştefănuţ S. [BUCA B3982].Ciucaş Mountains, Berii Brook Valley, Tigăile Mici Culm, Roşu Mountain, Zăgan Mountain,Bratocea Culm, Stâncoasă Culm (Mohan 1990b:44).Braşov County, Braşov (Schur 1866:874; Fuss 1878:653); Predeal (Loitlesberger 1898:196);Cheia Brook Valley, Râşnov Gorges, on fir bark, 850 m alt., 22.09.1940, leg. Cretzoiu P., det. Papp C.[FRE 2388; PHHM 6207961, 6207806, 6207807, 6207405] (Borza 1943:23); Lempeş Hill, on Quercusrobur, 05.1961 (Mihai 1962:278); ♀♂ - Tâmpa, 18.06.2006, leg. & det. Ştefănuţ S. [BUCA B3508,241
B3521, B3522] (Ştefănuţ 2006b:34; Ştefănuţ 2008b:29); ♀♂ - Făgăraş, Galaţi Hill, 24.11.2002, leg. &det. Ştefănuţ S. [BUCA B2059]; ♀♂ - Predeal, 6.04.2006, leg. & det. Ştefănuţ S. [BUCA B3396].Prahova County, Româneşti Forest, 16.04.1969, leg. Moşneagă M., det. Mohan G. [PHHM5936/3345](Mohan 1971a:9); Păuleşti-Ploieşti Forest, 22.09.1969, leg. Moşneagă M., det. Mohan G.[PHHM 6207655](Mohan 1971a:9); D<strong>of</strong>tana Valley, Secăria Valley, Păltinoasa Valley, on beech bark(Mohan 1971a:9; Mohan 1971b:48); Crângul lui Bot Forest, fer - Predeşti, Stejaru, fer - Tinosu (DihoruA. 1994:4); ♀♂ - Prahova Valley, 45°09'16"N / 25°42'56"E, 459 m alt., 27.04.2006, leg. & det. ŞtefănuţS. [BUCA B3404]; ♀♂ - D<strong>of</strong>tana Valley, in Quercus petraea forest, 45°10'11,8"N / 25°45'04,6"E,26.04.2006, leg. & det. Ştefănuţ S. [BUCA B3411, B3420] (Vasiliu-Oromulu 2008:87); ♀♂ - D<strong>of</strong>tanaValley, in mixed Quercus petraea with Fagus sylvatica forest, 45°10'01,6"N / 25°45'09,2"E, 27.04.2006,leg. & det. Ştefănuţ S. [BUCA B3428, B3423] (Vasiliu-Oromulu 2008:87).Gârbova Massif, Piscul Câinelui (Loitlesberger 1898:196); Rea Valley, 1000-1300 m alt.,Cumpătul Peak, 1651 m alt. (Mickiewicz 1968:30-34); Azuga Valley, Tufa Valley, Rea Valley, Valea luiBogdan, on Fagus sylvatica, 1967-1970 (Lungu & Ştefureac 1970:5); Cumpătu Mountain, on beech bark,830 m alt., 6.10.1985, leg. & det. Lungu L. [CSH].Bucegi Massif, fer - Bucşoiu, Mălăieşti Valley, 08-09.1911 (Degen 1914:217); Vârful cu Dor,1891 m alt., Urlătoarea Valley, 900-1000 m alt. (Mickiewicz 1968:30-34); Urlătoarea Valley,25.05.2000, leg. & det. Constantinescu A. [PHHM].Piatra Craiului National Park, Gura Râului, 28.09.2001, leg. Pop O., det. Ştefănuţ S. (Dihoru G. &al. 2003:70; Ştefănuţ & Pop O.G. 2006:82); ♀♂ - Great Gorges <strong>of</strong> Dâmboviţa River, AG, on Pinussylvestris, 45°23'18"N / 25°11'44"E, 800 m alt., 10.07.2003, leg. & det. Ştefănuţ S. [BUCA B2591, B2598,B2599], ♀♂ - [BUCA B2594] (Ştefănuţ & Pop O.G. 2006:82); Gura Râului, BV, 45°33'07"N / 25°17'50"E,900 m alt., 7.07.2003, leg. & det. Ştefănuţ S. [BUCA B2731] (Ştefănuţ & Pop O.G. 2006:82); Gar<strong>of</strong>iţaPietrei Craiului Hut, BV, 45°30'47"N / 25°10'26"E, 1080 m alt., 9.07.2003, leg. & det. Ştefănuţ S. [BUCAB2792] (Ştefănuţ & Pop O.G. 2006:82); Crăpătura Pietrei Craiului, BV, 45°33'44"N / 25°15'51"E, 1100 malt., 21.08.2003, leg. & det. Ştefănuţ S. [BUCA B2764] (Ştefănuţ & Pop O.G. 2006:82).Ialomiţa County, „Bărăganu” Experimental Forest Station (Dihoru G. & Mărgăritescu 1994:95,Tab.1).Ilfov County, Cocioc, Piscu-Hereasca <strong>and</strong> Ţigăneşti Forest, 22,23.04.1899, Cernica-Pusnicu,18.06.1900, sub var. propagulifera, fer - Cocioc, Vlăsia <strong>and</strong> Snagov Forest, 22,24.04.1899, Căldăruşani,5.06.1900 (Radian 1901b:41); Brăneşti-Pusnicul Forest, on bark, 22.12.1935, leg. Cretzoiu P., ClementO. & Beldie A. (Papp 1937a:160,163); Mogoşoaia Forest, 29.12.1959, leg. Racoviţă A., det. Ştefureac T.(Racoviţă A. 1960:1112,1113); Mogoşoaia Forest, 15.11.1959 (Racoviţă A. 1963:188).Bucharest, Băneasa Forest (Racoviţă A. 1941:154, Pl.I, Figs.4-8).Giurgiu County, s - Comana, frequent, on Quercus cerris <strong>and</strong> Q. frainetto, 1968-1969, sub fo.propagulifera (Ştefureac & Mohan 1970a:353); Tătarului Forest, on Quercus cerris, 28.10.1968, sub var.propagulifera (Ştefureac & Mohan 1970b:151,153,155, Tab.1); ♀♂ - Călugăreni Forest, 3.05.2004, leg.& det. Ştefănuţ S. [BUCA B2930]; Căscioarele Forest, 10.05.2005, leg. Ivănescu Cristina, det. Ştefănuţ S.[BUCA B3181]; Comana Forest, 31.05.2006, leg. & det. Ştefănuţ S. [BUCA B3534].Teleorman County, ♀♂ - Bujoreni, 21.04.2004, leg. & det. Ştefănuţ S. [BUCA B2936].Făgăraş Mountains, Arpaş Mountain (Schur 1866:874; Fuss 1878:653); Arpaşul Mare Valley,SB, on Quercus robur, 590 m alt., 8.10.2003, leg. & det. Ştefănuţ S. [BUCA B2925].Sibiu County, Sibiu (Schur 1866:874; Fuss 1878:653); Măgura Cisnădiei, Ursului Brook, leg. &det. Lerchenfeld J., sub Jungermannia [SIB HL 33712](Fuss 1878:653; Drăgulescu 1983:59; Gündisch1977:52); Muma, Prejba [SIB SCHUR 4608](Schur 1866:874; Fuss 1878:653; Gündisch 1977:52);Tălmaciu [SIB HF 1915](Fuss 1878:653; Gündisch 1977:52); Bradu-Halberweg, on bark, 3.04.1872, leg.& det. Fuss M. [SIB HF 1563](Fuss 1878:653; Drăgulescu 1983:59); Caşolţ, Şura Mare, Guşteriţa (Fuss1878:653); Guşteriţa-Formenthal, on srubs, 04.1865, leg. Fuss M. & det. Juratzka J. [SIB HF 1558](Fuss1865:28; Drăgulescu 1983:59); Bradu, 10.04.1865, 3.05.1875, leg. Fuss M., det. & rev. Drăgulescu C.[SIB HF 1559-1562,1564,1568] (Drăgulescu 1983:59).Cindrel Mountains, Crinţ, 900 m alt., Păltiniş, 1400 m alt., Ursului Hill, 1200 m alt. (Gündisch1977:52).Căpăţânei Mountains, Lotru Valley, Bucureasa, Malaia, 9-15.09.1952 (Ştefureac, Popescu &Lungu 1955:527).242
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SORIN ŞTEFĂNUŢThe Hornwort and L
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CONTENTINTRODUCTION ...............
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The beginning of the 20 th century
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asin of the Arieş river. Bryotaxon
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Abbreviations used in the text† -
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The presentation of the methodology
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Fig. 3. The GEOCOD mapping system.
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THE HORNWORT AND LIVERWORT SPECIES
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THE HORNWORT AND LIVERWORT CLASSIFI
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Suborder Lepidoziineae R.M. Schust.
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46. Cephalozia pleniceps (Austin) L
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147. Pallavicinia lyellii (Hook.) C
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13 Corsinia coriandrina (Spreng.) L
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11. Porella obtusata (Taylor) Trevi
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34Phaeoceros carolinianus (Michx.)
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Făgăraş Mountains, Cârţişoara
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Cluj County, Someşul Rece Valley a
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Parâng Mountains, Repedea Valley,
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Argeş County, Rucăr (Loitlesberge
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Cindrel Mountains, Cindrel, 2200 m
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Parâng Mountains, Latoriţa Valley
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Athalamia hyalina (Sommerf.) S. Hat
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Barbilophozia barbata (Schmidel ex
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Bârgău Mountains, Cârţibavu Mar
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First citation for Romania: Schur 1
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- Pleuroschisma triangulare (Schlei
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and Băiuţ, common spruce forest,
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Distribution in Europe: Sj, Is, Fo,
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Poiana Stampei Peatbog, SV, 47°20'
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alt., 8.08.2003, leg. Pop O., det.
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leg. & det. Ştefureac T., ass. Buc
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Rodna Mountains, Puzdra Mountain, 1
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Alba County, Vârciorog Valley, 780
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Bârgău Mountains, Cucureasa Reser
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Suceava County, Tinovul Mare de la
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Suceava County, Rodna Nouă, on Per
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2001:74,75); Arieşul Mare Basin, o
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Ţibleş Mountains, Mesteacănul Va
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Cindrel Mountains, Găuşoara, leg.
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Parâng Mountains, the edge of Câl
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& det. Ştefănuţ S. [BUCA B3684];
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Alba County, at 20,5 km to Zlatna,
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Cephaloziella spinigera (Lindb.) Wa
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First citation for Romania: Péterf
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Iezer-Păpuşa Mountains, Iezer Val
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- Cladopodiella fluitans var. gigan
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Caraş-Severin County, Cerna Valley
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(Mititelu, Barabaş N. & al. 1968:1
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1885:30; Ştefureac 1949a:287,288;
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104Conocephalum salebrosum Szweyk.,
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Retezat Mountains, Retezat, sub Jun
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Cindrel Mountains, Cânaia, 1750 m
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Iaşi County, Sineşti Forest (Papp
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[BUCA B2793], on rotten beech trunk
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Orthodicrano montani-Hypnetum filif
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Distribution in Romania:Maramureş
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- Gymnocolea inflata var. laxa (Nee
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Ciucaş Mountains, Caprei Brook Val
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Giumalău Mountain, Giumalău Secul
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- Haplozia riparia var. rivularis B
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- Jungermannia karl-muelleri auct.,
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Jungermannia gracillima Sm.Syn.:- A
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Mountain, PH, 45°24'40"N / 25°28'
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Retezat Mountains, Rovine Brook, on
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Romania, alt. 600-2280 m (Váňa 19
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- Solenostoma subellipticum (Lindb.
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Distribution in Romania:Bârgău Mo
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13.07.1946, sub Lophozia alpestris
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Distribution in Romania:Maramureş
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Rodna Mountains, Lala Valley, 1200
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Ştefănuţ S. [BUCA B2574, B2580,
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Dealul Colbului and Pietrele Doamne
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150var. rivularis (Raddi) Warnst.Sy
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Penteleu Massif, fer - Roşu Lake,
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Satu Mare County, Noroieni, Ghiarma
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Lophozia ascendens (Warnst.) R.M. S
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- Jungermannia intermedia Lindenb.-
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Căliman Mountains, on humus and ro
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Hăşmaş Mountains, Hăşmaş, sub
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Herbaria and Exsiccate: BUCADistrib
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peatbog on the border of Câlcescu
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? - Iezer-Păpuşa Mountains, Iezer
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Făgăraş Mountains, s - above Avr
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Nemira Mountains, on edge of Nemira
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2003:7); Măgura Cisnădiei [SIB HL
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Fuss 1854:20; Ştefureac 1949a:288;
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- Nardia brevissima (Dumort.) Lindb
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1919:295); Ciortea Mountain, on wet
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- Sarcoscyphus sphacelatus var. med
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Siriu Mountain, Poiana din Fundul M
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Iaşi County, below Repedea, 7.10.1
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Valley, Arieşul Mic - upstream min
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Fig. 13. Distribution of Anastrophy
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Fig. 17. Distribution of Anthelia j
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Fig. 21. Distribution of Asterella
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Fig. 25. Distribution of Barbilopho
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Fig. 29. Distribution of Barbilopho
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Fig. 33. Distribution of Bazzania t
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Fig. 37. Distribution of Calypogeia
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Fig. 41. Distribution of Calypogeia
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Fig. 45. Distribution of Cephalozia
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Fig. 49. Distribution of Cephalozia
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Fig. 53. Distribution of Cephalozie
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Fig. 57. Distribution of Cephalozie
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Fig. 61. Distribution of Cephalozie
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Fig. 65. Distribution of Cololejeun
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Fig. 69. Distribution of Corsinia c
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Fig. 73. Distribution of Fossombron
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Fig. 77. Distribution of Frullania
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Fig. 81. Distribution of Gymnocolea
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Fig. 85. Distribution of Harpanthus
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Fig. 89. Distribution of Jungermann
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Fig. 93. Distribution of Jungermann
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Fig. 97. Distribution of Jungermann
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Fig. 101. Distribution of Jungerman
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Fig. 105. Distribution of Leiocolea
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Fig. 109. Distribution of Lejeunea
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Fig. 113. Distribution of Lophocole
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Fig. 117. Distribution of Lophozia
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Fig. 121. Distribution of Lophozia
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Fig. 125. Distribution of Lunularia
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Fig. 129. Distribution of Marchanti
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Fig. 133. Distribution of Marsupell
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Fig. 137. Distribution of Marsupell
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Fig. 141. Distribution of Moerckia
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Fig. 145. Distribution of Nardia ge
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Fig. 149. Distribution of Odontosch
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Fig. 153. Distribution of Pedinophy
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Fig. 157. Distribution of Peltolepi
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Fig. 161. Distribution of Porella a
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Fig. 165. Distribution of Preissia
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Fig. 169. Distribution of Radula li
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Fig. 173. Distribution of Riccardia
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Fig. 177. Distribution of Riccia ca
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Fig. 181. Distribution of Riccia cr
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Fig. 185. Distribution of Riccia go
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Fig. 189. Distribution of Ricciocar
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Fig. 193. Distribution of Scapania
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Fig. 197. Distribution of Scapania
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Fig. 201. Distribution of Scapania
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Fig. 205. Distribution of Scapania
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Fig. 209. Distribution of Scapania
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Fig. 213. Distribution of Scapania
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Fig. 217. Distribution of Tritomari
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Evaluating the distribution dataFol
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Fig. 224. Hornworts and Liverworts
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Signs used:Bolded names - species i
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Marchantiaceae50. VU - Bucegia roma
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Fig. 228. The distribution of zones
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Anthoceros communis Steph. = Phaeoc
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Blasia funckii Corda = Blasia pusil
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Cephalozia crassiflora Spruce = Cep
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Cephaloziella spinigera (Lindb.) Wa
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Diplomitrium lyellii (Hook.) Corda
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Gymnocolea inflata subsp. acutiloba
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Jungermannia acuta var. gypsacea Go
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Jungermannia dentata Raddi = Cephal
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Jungermannia limprichtii Lindb. = L
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Jungermannia scalaris var. denudata
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Leiocolea heterocolpos (Thed. ex C.
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Lophozia incisa var. inermis Müll.
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Marchantia polymorpha var. dilatata
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Mastigobryum deflexum (Mart.) Ångs
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Odontoschisma sphagni var. denudatu
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Porella platyphylla (L.) Pfeiff.Por
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Riccia maior Roth = Riccia glauca L
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Scapania heterophylla M. Howe = Sca
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Sphagnocetis communis var. macrior
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Mittheilungen des siebenbürgischen
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91. Borza A., 1922a. Schedae ad "Fl
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142. Boşcaiu N., Täuber F., Colde
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199. Coldea G., 1975. Étude phytos
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256. Csűrös Ş., Moldovan I., Cs
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317. Dihoru G., 1987b. Contribuţie
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367. Domin K., 1923. Grundzüge der
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426. Gergely I., 1969. Pajişti mez
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487. Györffy I., 1906c. Polytrichu
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540. Györffy I., 1943b. Erdélyi P
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598. Ionescu Venera, Cristurean I.,
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658. Lungu Lucia, 1967a. Contribuţ
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712. Mihai G., 1971e. Aspecte ale b
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767. Mihăilescu Simona, Ştefănu
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824. Mohan G., 1988c. Die Gattung T
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879. Niţu I.T, 1973. Solurile cu e
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941. Papp C., 1931b. O schiţă a v
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999. Papp C., Eftimie Elena, 1963.
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1059. Péterfi L.Ş., 1963. Flagela
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1119. Plămadă, Viţalariu, 1971.
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1178. Pop I., Hodişan I., 1958. Co
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1231. Prunescu-Arion Elena, Toniuc
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1285. Raţiu O., 1970. Consideraţi
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1342. Rüegsegger F., 1986. Frullan
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1396. Schiffner V., 1913b. Über ei
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1455. Söderström L., Urmi E., Vá
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1515. Ştefănuţ S., 2004d. The Ho
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1567. Ştefureac T., 1960b. Cercet
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1624. Ştefureac T., 1976c. Rolul b
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1675. Ştefureac T., Cristurean I.,
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1727. Tarnavschi I., Nedelcu G., 19
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1785. Wallfisch R., 2008. Contribut
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var. fragilis (A. Roth) Müll. Frib
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Riccardia incurvata Lindb. 28, 248,