The Hornwort and Liverwort Atlas of Romania

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Ptilidium pulcherrimum (Weber) VanioSyn.:– Blepharozia ciliaris var. pulcherrimum Warnst.– Blepharozia ciliaris var. wallrothianum Nees– Blepharozia pulcherrima (Weber) Lindb.– Blepharozia pulcherrima fo. corticola Müll. Frib.– Jungermannia pulcherrima Weber– Ptilidium ciliare var. pulcherrimum (Weber) Hoffm. ex Pfeiff.– Ptilidium ciliare var. wallrothianum (Nees) Gottsche et al.– Ptilidium pulcherrimum fo. rupicola Schiffn.– Ptilidium pulcherrimum var. subpinnatum Jørg.– Ptilidium pulcherrimum var. wallrothianum Husn.First citation for Romania: Loitlesberger 1898.Moldova, incl. sub Blepharozia pulcherrima fo. corticola (Papp 1943:664; Ştefureac & al. 1973:143).Muntenia (Mohan 1988b:110).Transylvanian Alps (Müller 1954:572).Ecology: corticolous, humicolous, saprolignicolous, acidophytic, xero-mesophytic, sciophytic; circumpolar; boreal; Hch.Characteristic species to Vaccinio-Piceetea.Herbaria and Exsiccate: BUCA, HIPBC, PHHMDistribution in Europe: No, Dk, Se, Fi, Ie, Br, Fr, Nl, Be, De, Po, Ch, At, Cz, Sk, Hu, Es, It, Si, Hr, Sb, Me, Ba, Ro, Bg, Ee, Lv,Lt, By, Md, Ua, Cm, Rn, Rw, Rc, Re, Rs, Ca.Distribution in Romania:Maramureş County, Şerban Mountain, leg. & det. Boros Á. (Boros & Vajda 1969:412); betweenTocila Valley and Băiuţ, common spruce forest, 600 m alt., 21-23.08.1998 (Jakab 1999:55).Rodna Mountains, Lala Valley, 1200 m alt., 17.08.1937, sub fo. rupicola (Ştefureac 1938:241,244,249-251,258, Fig.5a-c).Ţibleş Mountains, Piciorul Ţiglei, Ţibleş Valley, Dosul Obcinei, Mesteacănul Valley, 1984-1985(Mohan 1988a:16).Suceava County, Breaza, 3.06.1910, sub P. ciliaris var. pulcherimum (Lilienfeldówna 1911:735);Bistriţa Aurie Valley, between Iacobeni and Ciocăneşti, the edge of Suhărzelul and Suhărzelul MareBrooks (Pascal 1969:107; Pascal & Mititelu 1971; Pascal 1973; Ştefureac & Pascal 1981:478; Ştefureac1986f:194); Piatra Ţibău, 1967 (Ştefureac & Pascal 1970a:246); Tinovu Mare de la Poiana Stampei(Ştefureac 1977a:49,52,53); Cristişor-Neagra Broştenilor, Borcut Meadow, in common spruce forest(Lungu 1973b:72; Lungu 1981c:90).Giumalău Mountain, Giumalău, subalpine zone, det. Ştefureac T. (Cristurean 2000:172); ♀ -Giumalău Secular Forest, 47°26'12"N / 25°29'20"E, 1200 m alt., 13.09.2006, leg. & det. Ştefănuţ S.[BUCA B3667], s - [BUCA B3701]; Giumalău Secular Forest, 47°26'12"N / 25°29'20"E, 1210 m alt.,1.09.2008, leg. & det. Ştefănuţ S. [BUCA B3956].Rarău Mountain, Pietrele Doamnei, 15.05.1910, sub P. ciliaris var. pulcherimum (Lilienfeldówna1911:735); Rarău, sub P. ciliaris var. pulcherrimum fo. corticola (Papp 1927b:149); Rarău-Todirescu,sub Blepharozia ciliaris var. pulcherimum fo. rupicola (Papp 1931a:219); Codrul Secular Slătioara, 820-1100 m alt. (Ştefureac 1941:1154,1283, Tabs.3,8; Ştefureac 1969a: Tab.16; Ştefureac 1986f:193); CodrulSecular Slătioara, Rarău Peak, incl. sub Blepharozia pulcherrima fo. corticola and rupicola (Papp1969b:188); ♀ - Rarău Mountain, 47°27'15"N / 25°33'45"E, 1450 m alt., 25.07.2007, leg. & det. ŞtefănuţS. [BUCA B3770].Bârgău Mountains, Cucureasa Reserve, Coşna Forest Hut, 930-1230 m alt., 1967, 1969, ass.Tetraphidetum pellucidae (Barkman 1958) Maurer 1961, ass. Lophocolea heterophylla-Ptilidiumpulcherrimum Maurer 1961, ass. Orthodicraneto-Plagiothecielletum Barkman 1958 (Ştefureac, Mihai &Pascal 1976:35,36,39, Tabs.1,2); Cârţibavu Mare Valley, Secu Brook, Leşului Valley, Măgura Calului,1983-1984 (Mohan & Hodoroga 1986:211).Căliman Mountains, on fallen rotten common spruce trees, Pinus mugo bark and rocks, 1981-1984 (Mihai 1985:97; Ştefureac 1986f:194).Neamţ County, Neamţ Monastery, on rotten wood, 07-08.1923, leg. Brândză M., det. Ştefureac T.(Ştefureac 1942a:5).Ceahlău Mountain, Patapi, incl. sub Blepharozia pulcherimum var. corticola (Papp 1924b:392;Papp 1934b:68,78; Zanoschi 1970:280; Mihai 1979b; Manoliu & al. 1998:338).Hăşmaş Mountains, Hăşmaş (Schiffner 1914:308).Harghita Mountains, Stâncilor Peak (Schiffner 1914:308); Stâncilor Peak, sub Blepharozia(Degen 1930:976); Greu Mountain, 1100 m alt., 11.07.2000, Asău Valley, 1000-1100 m alt., 22.10.2000,237
Descut, 15.08.2001, Podul de Piatră Mountain, 1551 m alt., 15.08.2001, Rotund Mountain, 1100 m alt.,28.08.2001, Pietros Valley, 900 m alt., 28.08.2001, Boitor Forest, 1000-1100 m alt., 28.08.2001, UlieşulMic, 1150 m alt., 17.04.2001, Luci Meleche Forest, 1050 m alt., 13.08.2001, leg. Bálint L. & Csízi K.(Bálint & Orbán 2003:14).Harghita County, Mohoş (Boros 1943c:114; Bálint & Orbán 2006:470); Mohoş (Pop E.1960:322; Bálint & Orbán 2006:470); Bălan Spa (Bálint & Orbán 2006:474).Nemira Mountains, Nemira Mountain, on soil and common spruce trees trunks, 1350-1550 m alt.(Barabaş V. 1974:26,28); Şandru, Nemira, on trees trunks, 1970-1977 (Mihai & Barabaş V.1978:114,115); Şandru Peak, 1550 m alt. (Barabaş V. 1982b:56, Tab.3); Ţiganca Mountain, 1400 m alt.,ass. Dicrano scopari-Hypnetum cupressiformis Will. 1962, Nemira Mountain, 1350-1600 m alt., ŞandruPeak, 1400 m alt. (Barabaş V. 1982b:57,62,63,65,66, Tabs.5,10-12,14,15).Bacău County, Slănic Brook, on rotten wood, 2.05-8.11.1971 (Ştefureac & Barabaş V.1972:76,77,82); Trotuş Valley at Straja-Asău and Şuliţa-Agăş, ass. Myrtillo-Pinetum Kobendza 1930 em.Passarge 1956 (Mititelu & Barabaş N. 1974:76).Ciucaş Mountains, Zăgan Mountain, Piscul Caprei, Şuviţelor Culm, Roşu Mountain (Mohan1990b:44).Prahova County, Doftana Valley [PHHM 7774/3873](Mohan 1971a:8; Mohan 1971b:47).Postăvaru Mountain, without locality (Schiffner 1914:308).Gârbova Massif, Azuga Valley, Tufa Valley, Rea Valley, on Picea abies bark, 1967-1970 (Lungu& Ştefureac 1970:5).Bucegi Massif, subalpine zone, 1600-2100 m alt. (Puşcaru E. & al. 1950:15).Piatra Craiului Mountains, towards Păşunea Vlăduşca, 30.10.2001, leg. Pop O., det. Ştefănuţ S.(Dihoru G. & al. 2003:70; Ştefănuţ & Pop O.G. 2006:81); Crăpătura Pietrei Craiului, BV, 45°33'49"N /25°15'54"E, 1050 m alt., leg. & det. Ştefănuţ S., 21.08.2003 [BUCA B2781] (Ştefănuţ & Pop O.G.2006:81); ♀ - Crăpătura Pietrei Craiului, BV, on common spruce bark, 45°33'44"N / 25°15'51"E, 1100 malt., 21.08.2003, leg. & det. Ştefănuţ S. [BUCA B2787] (Ştefănuţ & Pop O.G. 2006:81); Crăpătura PietreiCraiului, BV, on rotten wood, 45°33'37"N / 25°15'48"E, 1200 m alt., 22.08.2003, leg. & det. Ştefănuţ S.[BUCA B2786] (Ştefănuţ & Pop O.G. 2006:81); Marele Grohotiş, AG, 45°30'36"N / 25°11'38"E, 1550 malt., 8.07.2003, leg. & det. Ştefănuţ S. [BUCA B2638] (Ştefănuţ & Pop O.G. 2006:81,82).Făgăraş Mountains, Mozgavului sheephold and Negoiu on Izvorului Valley, 1200-1400 m alt.,sub Blepharozia ciliaris var. pulcherrima (Loitlesberger 1898:196; Drăgulescu 2003:9).Argeş County, Oeşti, 500 m alt. (Herzog 1919:295).Cindrel Mountains, Păltiniş, 1200-1400 m alt., Încărcătura Orlatului, 1400 m alt., Onceşti, 1700m alt. (Gündisch 1977:46; Drăgulescu 2003:9); Beşineu, 1600 m alt., det. Ştefureac T. (Gündisch1977:46; Drăgulescu 2003:9); Păltiniş [HIPBC](Drăgulescu 2003:9).Sebeşului Mountains, Sebeş Valley, between Oaşa and Ciban, in common spruce forest, 800-1200m alt., Şurianu Mountain (Borza 1959:69,283, Tab.38; Drăgulescu 2003:9).Parâng Mountains, Cărării Valley, 08-09.1954 (Ştefureac, Popescu & Lungu 1959:8,40,41, Tab.8).Retezat Mountains, Galeşu Lake Bank (Páll 1964a:108); Zlătuia Valley, 1300-1400 m alt., 6-8.09.1969, ass. Nowellio-Ptilidietum pulcherrimi Plămadă 1982, 8,09.1969, ass. Isopterygio seligeri-Dicranetum scoparii Ježek et Vondr. 1962 (Plămadă 1982a:312,313, Tabs.6,7; Plămadă 1993:52, Tab.2);Bucura Lake, 18.08.2004, leg. Onete Marilena, det. Ştefănuţ S. [BUCA B3062].Ţarcu Mountains, Muntele Mic, 1000-1800 m alt., 03.-08.1943, leg. Boşcaiu N. (Ştefureac1948a:220).Alba County, Vidra-Scărişoara, 800 m alt. (Péterfi M. 1908:276); Izbucul Ponorului (Boros &Vajda 1974:6); Morii Valley, on rotten wood, 600 m alt., 7.08.1994, 22.10.1995, ass. Riccardio-Nowellietum curvifoliae Koppe 1956, ass. Dicrano scoparii-Hypnetum cupressiformis Barkman 1958(Goia 1998:260,262, Tabs.9,12); Morii Valley, 600 m alt., 22.10.1995, ass. Riccardio-Scapanietumumbrosae Philippi 1965, Gârda Valley, 1200 m alt., 7.08.1996, ass. Anastrepto orcadensis-Dicranodontietum denudati Ştefur. 1941, Găina, 980 m alt., 21.08.1992, Cobleş Valley, 1100 m alt.,10.08.1996, ass. Brachythecio salebrosi-Drepanocladetum uncinati Marstaller 1989, Morii Valley, 700 malt., 22.10.1995, ass. Dicrano scoparii-Hypnetum filiformis Barkman 1958 (Goia 2000:112,116, 117,119,Tabs.2,5-7); Arieşul Mic Basin (Goia & Schumacker 2000:92); Morii Valley, on common spruce barkand rotten wood, Băii Brook, on beech bark, Gârda Seacă upstream Casa de Piatră, Ursoaia Saddle,Mătişeşti, on rotten wood (Goia 2001:82); Arieşul Mare Basin, on rotten wood (Goia & Schumacker238
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SORIN ŞTEFĂNUŢThe Hornwort and L
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CONTENTINTRODUCTION ...............
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The beginning of the 20 th century
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asin of the Arieş river. Bryotaxon
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Abbreviations used in the text† -
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The presentation of the methodology
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Fig. 3. The GEOCOD mapping system.
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THE HORNWORT AND LIVERWORT SPECIES
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THE HORNWORT AND LIVERWORT CLASSIFI
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Suborder Lepidoziineae R.M. Schust.
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46. Cephalozia pleniceps (Austin) L
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147. Pallavicinia lyellii (Hook.) C
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13 Corsinia coriandrina (Spreng.) L
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11. Porella obtusata (Taylor) Trevi
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34Phaeoceros carolinianus (Michx.)
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Făgăraş Mountains, Cârţişoara
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Cluj County, Someşul Rece Valley a
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Parâng Mountains, Repedea Valley,
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Argeş County, Rucăr (Loitlesberge
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Cindrel Mountains, Cindrel, 2200 m
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Parâng Mountains, Latoriţa Valley
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Athalamia hyalina (Sommerf.) S. Hat
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Barbilophozia barbata (Schmidel ex
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Bârgău Mountains, Cârţibavu Mar
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First citation for Romania: Schur 1
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- Pleuroschisma triangulare (Schlei
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and Băiuţ, common spruce forest,
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Distribution in Europe: Sj, Is, Fo,
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Poiana Stampei Peatbog, SV, 47°20'
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alt., 8.08.2003, leg. Pop O., det.
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leg. & det. Ştefureac T., ass. Buc
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Rodna Mountains, Puzdra Mountain, 1
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Alba County, Vârciorog Valley, 780
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Bârgău Mountains, Cucureasa Reser
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Suceava County, Tinovul Mare de la
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Suceava County, Rodna Nouă, on Per
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2001:74,75); Arieşul Mare Basin, o
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Ţibleş Mountains, Mesteacănul Va
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Cindrel Mountains, Găuşoara, leg.
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Parâng Mountains, the edge of Câl
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& det. Ştefănuţ S. [BUCA B3684];
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Alba County, at 20,5 km to Zlatna,
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Cephaloziella spinigera (Lindb.) Wa
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First citation for Romania: Péterf
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Iezer-Păpuşa Mountains, Iezer Val
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- Cladopodiella fluitans var. gigan
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Caraş-Severin County, Cerna Valley
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(Mititelu, Barabaş N. & al. 1968:1
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1885:30; Ştefureac 1949a:287,288;
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104Conocephalum salebrosum Szweyk.,
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Retezat Mountains, Retezat, sub Jun
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Cindrel Mountains, Cânaia, 1750 m
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Iaşi County, Sineşti Forest (Papp
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[BUCA B2793], on rotten beech trunk
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Orthodicrano montani-Hypnetum filif
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Distribution in Romania:Maramureş
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- Gymnocolea inflata var. laxa (Nee
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Ciucaş Mountains, Caprei Brook Val
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Giumalău Mountain, Giumalău Secul
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- Haplozia riparia var. rivularis B
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- Jungermannia karl-muelleri auct.,
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Jungermannia gracillima Sm.Syn.:- A
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Mountain, PH, 45°24'40"N / 25°28'
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Retezat Mountains, Rovine Brook, on
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Romania, alt. 600-2280 m (Váňa 19
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- Solenostoma subellipticum (Lindb.
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Distribution in Romania:Bârgău Mo
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13.07.1946, sub Lophozia alpestris
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Distribution in Romania:Maramureş
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Rodna Mountains, Lala Valley, 1200
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Ştefănuţ S. [BUCA B2574, B2580,
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Dealul Colbului and Pietrele Doamne
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150var. rivularis (Raddi) Warnst.Sy
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Penteleu Massif, fer - Roşu Lake,
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Satu Mare County, Noroieni, Ghiarma
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Lophozia ascendens (Warnst.) R.M. S
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- Jungermannia intermedia Lindenb.-
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Căliman Mountains, on humus and ro
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Hăşmaş Mountains, Hăşmaş, sub
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Herbaria and Exsiccate: BUCADistrib
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peatbog on the border of Câlcescu
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? - Iezer-Păpuşa Mountains, Iezer
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Făgăraş Mountains, s - above Avr
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Nemira Mountains, on edge of Nemira
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2003:7); Măgura Cisnădiei [SIB HL
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Fuss 1854:20; Ştefureac 1949a:288;
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- Nardia brevissima (Dumort.) Lindb
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1919:295); Ciortea Mountain, on wet
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- Sarcoscyphus sphacelatus var. med
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Siriu Mountain, Poiana din Fundul M
Page 187 and 188: Iaşi County, below Repedea, 7.10.1
Page 189 and 190: Valley, Arieşul Mic - upstream min
Page 191 and 192: Alba County, Valea Lungă, 10.02.18
Page 193 and 194: Rarău Massif, between Dealul Colbu
Page 195 and 196: Harghita County, Harghita Spa, on r
Page 197 and 198: Căliman Mountains, in alpine zone,
Page 199 and 200: Stânişoara Mountains, Suha Mare B
Page 201 and 202: Ecology: humicolous, acidophytic, h
Page 203 and 204: Bacău County, Slănic Brook (Papp
Page 205 and 206: Rarău Mountain, Codrul Secular Sl
Page 207 and 208: 1915:197,198); Cândeşti, 07.1891,
Page 209 and 210: Ţibleş Mountains, Piciorul Ţigle
Page 211 and 212: Ceahlău Mountain, Durău Valley, i
Page 213 and 214: major (Ştefureac 1962b:69); Benea-
Page 215 and 216: Ciucaş Mountains, Berii Brook Vall
Page 217 and 218: Zlătuia Valley, 1100 m alt., 7.09.
Page 219 and 220: - Plagiochila dillenii (Taylor) Tay
Page 221 and 222: Wallfisch R. (Dihoru G. & al. 2003:
Page 223 and 224: Ştefănuţ M. 2003:79); Iezerul Pi
Page 225 and 226: utabuli-Mnietum cuspidati (Felföld
Page 227 and 228: Iaşi County, Mârzeşti Forest, Bu
Page 229 and 230: 3.04.2004, leg. Atanasiu Paulina, d
Page 231 and 232: Arad County, Săvârşin, sub Madot
Page 233 and 234: Preissia quadrata (Scop.) NeesSyn.:
Page 235 and 236: Diana, BV, 1100 m alt., 22.08.2001,
Page 237: Rodna Mountains (Pax 1908:130); Pie
Page 241 and 242: Bârgău Mountains, Cucureasa Reser
Page 243 and 244: B3521, B3522] (Ştefănuţ 2006b:34
Page 245 and 246: 16,20.08.1994, ass. Pterigynandretu
Page 247 and 248: - Marchantia crinita Michx.- Marcha
Page 249 and 250: Giumalău Mountain, Giumalău Secul
Page 251 and 252: Riccardia multifida (L.) GraySyn.:-
Page 253 and 254: Penteleu Massif, the left side of C
Page 255 and 256: Herbaria and Exsiccate: SIBDistribu
Page 257 and 258: 558,560; Papp 1939a:358; Papp & Eft
Page 259 and 260: Riccia glauca L. var. glaucaSyn.:-
Page 261 and 262: Distribution in Europe: Is, Fo, No,
Page 263 and 264: Bihor County, Tărian, sub Riccia (
Page 265 and 266: 1960:1113); upper side of Mălăie
Page 267 and 268: Scapania brevicaulis TaylorSyn.:- S
Page 269 and 270: Suceava County, Cristişor-Neagra B
Page 271 and 272: Piatra Mare Mountain, 1700 m alt.,
Page 273 and 274: - Scapania amurensis Warnst.- Scapa
Page 275 and 276: Mureş County, Sighişoara - Fuchsl
Page 277 and 278: Căliman Mountains, the peatbogs fr
Page 279 and 280: - Scapania heterophylla M. Howe- Sc
Page 281 and 282: Slăvei and Muchia Ascuţită, 2000
Page 283 and 284: Distribution in Europe: Ie, Br, Fr,
Page 285 and 286: Nemira Mountains, Nemira, on humus,
Page 287 and 288: - Sphenolobus medelpadicus (Arnell)
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Parâng Mountains, Latoriţa Valley
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Fig. 9. Distribution of Phaeoceros
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Fig. 13. Distribution of Anastrophy
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Fig. 17. Distribution of Anthelia j
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Fig. 21. Distribution of Asterella
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Fig. 25. Distribution of Barbilopho
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Fig. 29. Distribution of Barbilopho
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Fig. 33. Distribution of Bazzania t
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Fig. 37. Distribution of Calypogeia
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Fig. 41. Distribution of Calypogeia
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Fig. 45. Distribution of Cephalozia
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Fig. 49. Distribution of Cephalozia
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Fig. 53. Distribution of Cephalozie
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Fig. 65. Distribution of Cololejeun
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Fig. 69. Distribution of Corsinia c
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Fig. 73. Distribution of Fossombron
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Fig. 77. Distribution of Frullania
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Fig. 81. Distribution of Gymnocolea
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Fig. 85. Distribution of Harpanthus
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Fig. 89. Distribution of Jungermann
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Fig. 101. Distribution of Jungerman
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Fig. 105. Distribution of Leiocolea
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Fig. 109. Distribution of Lejeunea
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Fig. 113. Distribution of Lophocole
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Fig. 117. Distribution of Lophozia
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Fig. 121. Distribution of Lophozia
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Fig. 125. Distribution of Lunularia
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Fig. 129. Distribution of Marchanti
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Fig. 133. Distribution of Marsupell
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Fig. 137. Distribution of Marsupell
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Fig. 141. Distribution of Moerckia
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Fig. 145. Distribution of Nardia ge
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Fig. 149. Distribution of Odontosch
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Fig. 153. Distribution of Pedinophy
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Fig. 157. Distribution of Peltolepi
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Fig. 161. Distribution of Porella a
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Fig. 165. Distribution of Preissia
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Fig. 169. Distribution of Radula li
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Fig. 173. Distribution of Riccardia
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Fig. 177. Distribution of Riccia ca
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Fig. 181. Distribution of Riccia cr
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Fig. 185. Distribution of Riccia go
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Fig. 189. Distribution of Ricciocar
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Fig. 193. Distribution of Scapania
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Fig. 217. Distribution of Tritomari
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Evaluating the distribution dataFol
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Fig. 224. Hornworts and Liverworts
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Signs used:Bolded names - species i
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Marchantiaceae50. VU - Bucegia roma
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Fig. 228. The distribution of zones
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Anthoceros communis Steph. = Phaeoc
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Blasia funckii Corda = Blasia pusil
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Cephalozia crassiflora Spruce = Cep
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Cephaloziella spinigera (Lindb.) Wa
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Diplomitrium lyellii (Hook.) Corda
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Gymnocolea inflata subsp. acutiloba
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Jungermannia acuta var. gypsacea Go
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Jungermannia dentata Raddi = Cephal
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Jungermannia limprichtii Lindb. = L
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Jungermannia scalaris var. denudata
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Leiocolea heterocolpos (Thed. ex C.
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Lophozia incisa var. inermis Müll.
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Marchantia polymorpha var. dilatata
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Mastigobryum deflexum (Mart.) Ångs
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Odontoschisma sphagni var. denudatu
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Porella platyphylla (L.) Pfeiff.Por
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Riccia maior Roth = Riccia glauca L
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Scapania heterophylla M. Howe = Sca
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Sphagnocetis communis var. macrior
Page 445 and 446:
Mittheilungen des siebenbürgischen
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91. Borza A., 1922a. Schedae ad "Fl
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142. Boşcaiu N., Täuber F., Colde
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199. Coldea G., 1975. Étude phytos
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256. Csűrös Ş., Moldovan I., Cs
Page 455 and 456:
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The Hornwort and Liverwort Atlas of Romania

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