The Hornwort and Liverwort Atlas of Romania

New subspecies of Leiocolea bantriensisAmong the bryophytes collected from Lăptici Peatbog, Bucegi Massif, I found some unusualliverworts, which has characters close to Leiocolea bantriensis (Hook.) Jörg. subsp. bantriensis, L.rutheana (Limpr.) Müll. Frib. var. rutheana and L. rutheana var. laxa (Schiffn. ex Burrell) Paton.Based on the presence of hairs on underleaves lamina has been described a new subspecies ofLeiocolea bantriensis:Leiocolea bantriensis (Hook.) Jörg. subsp. wallfischii Ştefănuţ, subsp. nov.Diagnosis. Subspecies subsp. bantriensis similis, differens, autem, pro eo foliis late, amphigastria ad laminae cumcapillus adaxiali crebis.Type. ROMANIA, Southern Carpathians, Bucegi Massif, Lăptici Peatbog, 1470 m alt., coord. 45°22'26"N /25°26'23"E, 28 July 2005, leg. Ştefănuţ S., BUCA B5209 - holotype.Leiocolea bantriensis subsp. wallfischii* is a gregarious liverwort, grows in large tufts, sometimesscattered among mosses, robust, light to deep green, with age become reddisch brown or fuscous. Plants3–4 mm wide × 4–8 cm long. Steam 350–400 μm in diameter, erect. Antical cortical cells 20-28 μmwide. Rhizoids numerous, long, pale brownisch. Leaves quite imbricate, widely spreading, broadly ovatereniform,dorsally convex, wider than long, ca. 2 mm wide × 1.6 mm long, ca. 1.2 × as wide as long,very obliquely inserted, leaves on lax shoots with postical end of insertion line decurrent. Leaves bilobed,sinus rounded, leaf cells averaging 32–38 μm, apex of leaf lobes 1–2-celled or rounded. Underleaves verylarge, to 0.5–1.2 mm long, usually 2–3-fid and lobes ciliate (Fig. 5a). Underleaves lamina with long hairs,from a few uniseriate cells to over 10 uniseriate cells long (Fig. 5b).Dioicous. Plants with young perianths. Antheridia missing. Asexual reprodution lacking.Ecology: aquatic, epigaeic, acidophytic, hydrophytic, photo-sciophytic; boreal-mountain; Hch.This subspecies grows in the water drainage from Lăptici Peatbog ditch, Bucegi Massif, Romania,along with Sanionia uncinata (Hedw.) Loeske and Fissidens taxifolium Hedw. On the edge of ditch growsSphagnum spp., Preissia quadrata and below Scapania paludicola and Aneura pinguis.abFig. 5. Leiocolea bantriensis (Hook.) Jörg. subsp. wallfischii Ştefănuţ, subsp. nov. a) underleaf, b) adaxial face ofunderleaf lamina with hairs.*The name of wallfischii is derived from the romanian bryologist Richard Wallfisch (1914-2008).21