Argeş County, Piatra Nemoeştilor (Loitlesberger 1898:193); Ghimbav Gorges, 810 m alt.,25.08.1994, ass. Asplenio (quadrivalens)-Poetum nemoralis Soó 1944 emend. Gergely 1966 (Alexiu1998:171); Great Gorges <strong>of</strong> Dâmboviţa River, AG, 45°23'18"N / 25°11'44"E, 800 m alt., 10.07.2003, leg.& det. Ştefănuţ S. [BUCA B2794]; ♀ - Great Gorges <strong>of</strong> Dâmboviţa River, AG, 45°23'04"N / 25°11'43"E,750 m alt., 8.06.2005, leg. Pop O., det. Ştefănuţ S. [BUCA B3173].Iezer-Păpuşa Mountains, Păpău Mountain, 1800 m alt. (Loitlesberger 1898:193).Giurgiu County, s - Comana, on soil at the base <strong>of</strong> Quercus cerris bark, 1968-1969 (Ştefureac &Mohan 1970a:353).Făgăraş Mountains, Arpaş Valley, beech forest (Heufler 1853:48-49; Fuss 1854:20; Schur1866:873; Fuss 1878:657; Drăgulescu 2003:14); Bâlea Lake, leg. Zsák Z. (Schiffner 1914:306;Drăgulescu 2003:14); Clăbucet Mountain, AG, 06.1941, sub var. maior (Racoviţă A. 1942:510; RacoviţăA. 1944b:34,37); above Bâlea Labe, below Paltina Peak, on s<strong>and</strong>y soil, 2250 m alt., 13.07.1946, belowCiortea Peak, north-eastern slope, 2320 m alt. (Ştefureac 1949a:289,290; Drăgulescu 2003:14); Voila(Mohan 1971a:9); Pojorta Valley, BV, 800-1000 m alt., 18.09.2001, leg. & det. Ştefănuţ S. [BUCAB2542] (Ştefănuţ 2002:190); Vidraru Lake, AG, 6.07.1998, leg. & det. Ştefănuţ S. [BUCA B3352].Sibiu County, Cisnădioara, Prejba [SIB SCHUR 4604](Schur 1866:873; Fuss 1878:657; Gündisch1977:49-50; Drăgulescu 2003:14); Tălmaciu [SIB HF 1068] (Fuss 1878:657; Gündisch 1977:49-50;Drăgulescu 2003:14); Bradu (Fuss 1878:657; Drăgulescu 2003:14); Sibiu, “in Querceto Cibinensi”, leg.& det. Lerchenfeld J., sub Jungermannia [SIB HL 33711](Fuss 1878:657; Drăgulescu 1983:58;Drăgulescu 2003:14); Gura Râului, 600 m alt. (Gündisch 1977:49-50; Drăgulescu 2003:14).Cindrel Mountains, Cibin Gorges, 900 m alt., Crinţ, 900-1000 m alt., Săroi, 1300 m alt., Păltiniş,1300 m alt., Beşineu, 1600 m alt. (Gündisch 1977:49-50; Drăgulescu 2003:14).Vâlcea County, s – Băiţa Valley, Văroasa Valley, in Carpino-Fagetum, on soil <strong>and</strong> bark (Ştefureac& Peicea 1973:45,46,49,50).Sebeşului Mountains, Sebeş Valley, between Oaşa <strong>and</strong> Ciban, in common spruce forest, 800-1200m alt., incl. sub var. maior (Borza 1959:68,220,283, Tab.38; Drăgulescu 2003:14).Căpăţânei Mountains, Lotru Valley, Bucureasa, Malaia, 9-15.09.1952 (Ştefureac, Popescu &Lungu 1955:527).Parâng Mountains, Latoriţa Valley, calcareous rocks, 900 m alt., 31.08.1954 (Ştefureac, Popescu& Lungu 1956a:513, Tab.1); Latoriţa Valley, Repedea Valley, 08-09.1954, incl. sub var. riparia(Ştefureac, Popescu & Lungu 1959:9,23-25,29,30,34,39, Tabs.3,4,6,7).Şureanu Mountains, Godeanu Valley, Tâmpu <strong>and</strong> Anineş, in beech forest (Balazs 1993:73).Hunedoara County, the Hills from Şoimuş, on Făeragului Valley <strong>and</strong> Piscului Brook (Certejul deSus), Hunedoara on Sânpetru Hill, ass. Carpino-Fagetum Paucă 1941 (S<strong>and</strong>a, Popescu & Peicea1972:315, Tab.7); between Deva <strong>and</strong> Şoimuş Commune, Certeju de Sus (Peicea 1973:440,443,446,Tabs.2,3).Retezat Mountains, Valereasca, upper region (Schiffner 1914:306); Zlătuia Valley, Râul MareValley, in common spruce forest, on wet rocks (Pall 1962:121); Dobrun Brook Valley, Radeşul MareValley, towards Tăul Negru (Páll 1964a:108); Zlătuia Valley, in beech-common spruce mixed forest, onsoil, 1000-1200 m alt., ass. Abieto-Fagetum piceetosum, det. Páll Ş. (Csűrös , Kovács & Moldovan1964:172); Şesele Brook, 1050 m alt., 21.06.1968, ass. Fontinaletum gracilis Plămadă 1974 (Plămadă1974b:Tab.3; Plămadă 1993:55, Tab.5); Retezat National Park, ass. Neckero complanatae-Leucodontetum(Lippmaa 1935) Plămadă 1982, ass. Leucodonto-Madothecietum platyphyllae Plămadă 1982, ass.Anomodontetum attenuati (Barkm. 1939) Peciar 1965, ass. Pterygyn<strong>and</strong>ro-Isothecietum myuri Plămadă(1976) 1978, ass. Neckeretum crispae Ştefur. 1941 (Plămadă 1976a; Plămadă 1976b:20, Tab.2); ZlătuiaValley, 16.09.1967, 6-7.09.1969, ass. Grimmio hartmanii-Isothecietum scabridi Plămadă 1978, TurculuiValley, 12.07.1968, Coama Şesele, 12.07.1968, ass. Pterigyn<strong>and</strong>ro-Isothecietum myuri Plămadă 1978,Turcului Valley, 10.09.1969, ass. Pogonatetum urnigeri Krusenstjerna 1945 retezaticum Plămadă 1978(Plămadă 1978:241-243, Tabs.1-3; Plămadă 1993:54, Tabs.3,4); Zlătuia Valley, ass. Homalothecio-Leucodontetum sciuroides (Størmer 1938) Ježet et Vondr. 1962, 800-1200 m alt., 6-7.09.1969, ass.Neckero complanatae-Leucodontetum, between Zlătuia Valley <strong>and</strong> Turcului Valley, 1100 m alt.,7.09.1969, ass. Leucodonto-Madothecetum platyphyllae, Zlătuia Valley, 800-850 m alt., 6-7.09.1969,Şesele Valley, 950 m alt., 9.09.1969, ass. Anomodontetum attenuati, Zlătuia Valley, 600 m alt.,6.09.1969, between Zlătuia Valley <strong>and</strong> Turcului Valley, 900 m alt., 7.09.1969, Şesele Valley, 950 m alt.,14.07.1970, ass. Pterigyn<strong>and</strong>ro-Isothecietum myuri, Şesele – Pârgului Valley, 950 m alt., 9.09.1969,215
Zlătuia Valley, 1100 m alt., 7.09.1970, ass. Anomodonto attenuati-Neckeretum crispae Plămadă 1982,Zlătuia Valley, 800 m alt., 6.09.1969, ass. Brachythecio-rutabuli-Mnietum cuspidati (Felföldy 1941)Plămadă 1982, Turcului Valley, 950 m alt., 9-10.09.1969, Cârligului Valley, 1000 m alt., 9.09.1969, ass.Dicrano scoparii-Hypnetum cupressiformis (v. Krusenst. 1945) Barkm. 1958 (Plămadă 1982a:303-307,309,315,316, Tabs.1-5,8,9; Plămadă 1993:51,52, Tabs.1,2); Retezat National Park, 1600 m alt., ass.Hieracio rotundatae-Piceetum Pawl. et Br.-Bl. 1939 (Coldea, Lupşa & Plămadă 1984:95, Tab.1);Gemenea Valley, 1600 m alt., 6.09.1967, ass. Scapanietum undulatae Schwick. 1944 (Plămadă 1993:55,Tabs.5,6); Retezat, 800-1200 m alt., ass. Chrysosplenio-Cardaminetum amarae (Tx. 1937) Mass 1959,1650-1750 m alt., ass. Hieracio rotundatae-Piceetum (Coldea 1993:41,46, Tabs.7,10).Ţarcu Mountains, Muntele Mic, 1000-1800 m alt., 03.-08.1943, leg. Boşcaiu N. (Ştefureac1948a:220).Gorj County, Rovinari, „Valea lui Lupe”, 10.1938, det. Papp C. (Racoviţă A. 1941b:253);Rovinari, Lupei Valley, on wet soil, 160 m alt. (Papp 1942a:51).Godeanu Mountains, Lunca Berhinii, 28.06.1967, ass. Chrysosplenio-Cardaminetum (Tx. 1937)Maas 1959 (Boşcaiu 1971a:125, Tab.1).Mehedinţi County, Orşova (Simonkai 1872:67; Hazslinszky 1885:70); in the forest from Suliţatowards Gornenţi, on rotten wood <strong>and</strong> rocks, 26.06.1899 (Radian 1901b:41); Vărănic Hill, way fromBrezniţa Commune towards Gura Văii, 400 m alt., 29.04.1945 (Ştefureac 1947a:138,146); EşelniţaValley, in forest, frequent, on soil (Plămadă 1970b:206); Danube Defile, between Orşova <strong>and</strong> CernaValley, in forest, frequent, on soil, 1966-1968 (Plămadă 1970c:393,395,396); Mraconia Valley,13.05.1969 (Coldea & al. 1970:473, Tab.2); Eşelniţa Valley, 90 m alt., ass. Hypno-Polypodietum Jko etPec. 1963 (Schneider-Binder & al. 1970:315, Tab.3).Domogled Mountain, Domogled (Degen 1901:14); Domogled, leg. & det. Papp C. (Papp1939b:7).Caraş-Severin County, Mehadia (Simonkai 1872:67); Cerna Valley, 450 m alt. (Herzog1919:290); Herculane Spa, on wet soil, 200 m alt., leg. Bujorean G., det. Papp C., sub var. major fo.reptans (Papp 1944a:98); Ruşchiţa [PHHM 123/1366, 342/2868, 199/1919, 5915/3335, 5954/3345](Mohan 1971a:9).Poiana Ruscă Mountains, Rusca Montană (Pančić 1861:94); Luncani, incl. sub var. major(Simonkai 1872:67; Fuss 1878:657); Poiana Ruscă, Govăjdia Forest, on soil, 700 m alt., Luncani, GladnaMontană, 600 m alt., leg. Pteancu P., det. Papp C. (Papp 1944a:98).Zar<strong>and</strong> Mountains, Arăneag, beech forest, Milova Valley, on rocks <strong>and</strong> bark (Csűrös M.1978:76; Ardelean 2006:105, Fig.21).Codru-Moma Mountains, Monesei Valley, Măgura Zăgăului, Lungă Valley, on beech bark(Mohan 1996:49; Ardelean 2006:105, Fig.21).Arad County, Săvârşin (Simonkai 1872:67); Cladova Valley, Fântâna Rece Hill, in montane zone<strong>of</strong> Arăneag, Rândunicii Valley above Chişindiei, on Chicioara, Drocea, Dezna Veche Valley, Nădălbeşti,Moneasa, Răşirata, Dulcele, Zimbru, Aciuvei’s Mountains, Calvaria Hill above Săvârşin, Troiaş Brook,Piatra Albă, Zam Valley, Hălmagiu Mountains zone towards Găina (Simonkai 1893:361,362; Fuss1878:657).Mureş County, Sighişoara, leg. & det. Baumgarten J.C.G. [SIB HF 1516](Baumgarten 1846:220;Schur 1866:873; Fuss 1878:657; Drăgulescu 1983:58); Sighişoara (Höhr 1914:87); west to Şaeş, 500-580m alt., det. Páll Ş, ass. Fagetum silvaticae transsilvanicum (Csűrös & Kovács 1962:220).Alba County, Piatra Ceţii, Trascău Mountains, 3.07.1868 (Barth 1868:141); Galda de Sus, leg. &det. Barth J. (Fuss 1878:657); Valea Lungă, on soil, 3.04.1866, leg. & det. Barth J. [SIB HF 1517,1518](Fuss 1878:657; Drăgulescu 1983:58); Mihalţ, on old trees trunks, leg. & det. Fuss M. (Borza &Lupşa 1964:151); Feneş Gorges, Groşilor Brook on Dumbrăviţa Hill at Zlatna, 680 m alt. (Papp1944b:236; Papp 1945a:161); Scărişoara, on soil, 1200 m alt. (Papp 1942a:51; Goia 2001:67); FeneşGorges (Papp 1945a:163); Bărăşti, Albac Commune, on wet soil, 656 m alt., sub fo. major (Papp1946a:13,15); Colţii Trascăului, Bedeleu Mountains (Páll 1960b:90); Întregalde Gorges, in beech forests,06.1960 (Páll 1964b:25); Măgina, Pădurea Şcolii Hill, above the way, ass. Lathyro (hallersteinii)-Fagetum, 15.08.1967, det. Plămadă E (Gergely 1968:344); right slope <strong>of</strong> Ampoiului Valley downstreamGăureni, hornbeam grove, at 16 km to Zlatna, hornbeam-beech mixed forest on left slope, upstream, at12,5 km to Zlatna, 880 m alt. <strong>and</strong> at 20,5 km to Zlatna, right slope, downstream, oak grove (Plămadă216
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SORIN ŞTEFĂNUŢThe Hornwort and L
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CONTENTINTRODUCTION ...............
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The beginning of the 20 th century
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asin of the Arieş river. Bryotaxon
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Abbreviations used in the text† -
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The presentation of the methodology
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Fig. 3. The GEOCOD mapping system.
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THE HORNWORT AND LIVERWORT SPECIES
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THE HORNWORT AND LIVERWORT CLASSIFI
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Suborder Lepidoziineae R.M. Schust.
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46. Cephalozia pleniceps (Austin) L
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147. Pallavicinia lyellii (Hook.) C
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13 Corsinia coriandrina (Spreng.) L
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11. Porella obtusata (Taylor) Trevi
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34Phaeoceros carolinianus (Michx.)
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Făgăraş Mountains, Cârţişoara
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Cluj County, Someşul Rece Valley a
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Parâng Mountains, Repedea Valley,
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Argeş County, Rucăr (Loitlesberge
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Cindrel Mountains, Cindrel, 2200 m
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Parâng Mountains, Latoriţa Valley
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Athalamia hyalina (Sommerf.) S. Hat
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Barbilophozia barbata (Schmidel ex
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Bârgău Mountains, Cârţibavu Mar
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First citation for Romania: Schur 1
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- Pleuroschisma triangulare (Schlei
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and Băiuţ, common spruce forest,
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Distribution in Europe: Sj, Is, Fo,
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Poiana Stampei Peatbog, SV, 47°20'
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alt., 8.08.2003, leg. Pop O., det.
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leg. & det. Ştefureac T., ass. Buc
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Rodna Mountains, Puzdra Mountain, 1
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Alba County, Vârciorog Valley, 780
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Bârgău Mountains, Cucureasa Reser
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Suceava County, Tinovul Mare de la
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Suceava County, Rodna Nouă, on Per
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2001:74,75); Arieşul Mare Basin, o
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Ţibleş Mountains, Mesteacănul Va
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Cindrel Mountains, Găuşoara, leg.
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Parâng Mountains, the edge of Câl
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& det. Ştefănuţ S. [BUCA B3684];
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Alba County, at 20,5 km to Zlatna,
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Cephaloziella spinigera (Lindb.) Wa
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First citation for Romania: Péterf
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Iezer-Păpuşa Mountains, Iezer Val
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- Cladopodiella fluitans var. gigan
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Caraş-Severin County, Cerna Valley
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(Mititelu, Barabaş N. & al. 1968:1
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1885:30; Ştefureac 1949a:287,288;
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104Conocephalum salebrosum Szweyk.,
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Retezat Mountains, Retezat, sub Jun
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Cindrel Mountains, Cânaia, 1750 m
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Iaşi County, Sineşti Forest (Papp
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[BUCA B2793], on rotten beech trunk
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Orthodicrano montani-Hypnetum filif
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Distribution in Romania:Maramureş
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- Gymnocolea inflata var. laxa (Nee
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Ciucaş Mountains, Caprei Brook Val
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Giumalău Mountain, Giumalău Secul
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- Haplozia riparia var. rivularis B
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- Jungermannia karl-muelleri auct.,
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Jungermannia gracillima Sm.Syn.:- A
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Mountain, PH, 45°24'40"N / 25°28'
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Retezat Mountains, Rovine Brook, on
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Romania, alt. 600-2280 m (Váňa 19
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- Solenostoma subellipticum (Lindb.
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Distribution in Romania:Bârgău Mo
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13.07.1946, sub Lophozia alpestris
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Distribution in Romania:Maramureş
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Rodna Mountains, Lala Valley, 1200
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Ştefănuţ S. [BUCA B2574, B2580,
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Dealul Colbului and Pietrele Doamne
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150var. rivularis (Raddi) Warnst.Sy
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Penteleu Massif, fer - Roşu Lake,
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Satu Mare County, Noroieni, Ghiarma
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Lophozia ascendens (Warnst.) R.M. S
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- Jungermannia intermedia Lindenb.-
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Căliman Mountains, on humus and ro
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Hăşmaş Mountains, Hăşmaş, sub
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Scapania brevicaulis TaylorSyn.:- S
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Suceava County, Cristişor-Neagra B
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Piatra Mare Mountain, 1700 m alt.,
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- Scapania amurensis Warnst.- Scapa
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Mureş County, Sighişoara - Fuchsl
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Căliman Mountains, the peatbogs fr
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- Scapania heterophylla M. Howe- Sc
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Slăvei and Muchia Ascuţită, 2000
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Distribution in Europe: Ie, Br, Fr,
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Nemira Mountains, Nemira, on humus,
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- Sphenolobus medelpadicus (Arnell)
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Parâng Mountains, Latoriţa Valley
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Fig. 9. Distribution of Phaeoceros
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Fig. 13. Distribution of Anastrophy
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Fig. 17. Distribution of Anthelia j
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Fig. 21. Distribution of Asterella
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Fig. 25. Distribution of Barbilopho
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Fig. 29. Distribution of Barbilopho
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Fig. 33. Distribution of Bazzania t
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Fig. 37. Distribution of Calypogeia
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Fig. 41. Distribution of Calypogeia
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Fig. 45. Distribution of Cephalozia
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Fig. 49. Distribution of Cephalozia
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Fig. 53. Distribution of Cephalozie
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Fig. 57. Distribution of Cephalozie
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Fig. 61. Distribution of Cephalozie
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Fig. 65. Distribution of Cololejeun
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Fig. 69. Distribution of Corsinia c
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Fig. 73. Distribution of Fossombron
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Fig. 77. Distribution of Frullania
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Fig. 81. Distribution of Gymnocolea
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Fig. 85. Distribution of Harpanthus
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Fig. 89. Distribution of Jungermann
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Fig. 93. Distribution of Jungermann
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Fig. 97. Distribution of Jungermann
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Fig. 101. Distribution of Jungerman
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Fig. 105. Distribution of Leiocolea
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Fig. 109. Distribution of Lejeunea
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Fig. 113. Distribution of Lophocole
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Fig. 117. Distribution of Lophozia
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Fig. 121. Distribution of Lophozia
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Fig. 125. Distribution of Lunularia
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Fig. 129. Distribution of Marchanti
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Fig. 133. Distribution of Marsupell
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Fig. 137. Distribution of Marsupell
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Fig. 141. Distribution of Moerckia
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Fig. 145. Distribution of Nardia ge
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Fig. 149. Distribution of Odontosch
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Fig. 153. Distribution of Pedinophy
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Fig. 157. Distribution of Peltolepi
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Fig. 161. Distribution of Porella a
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Fig. 165. Distribution of Preissia
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Fig. 169. Distribution of Radula li
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Fig. 173. Distribution of Riccardia
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Fig. 177. Distribution of Riccia ca
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Fig. 181. Distribution of Riccia cr
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Fig. 185. Distribution of Riccia go
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Fig. 189. Distribution of Ricciocar
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Fig. 193. Distribution of Scapania
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Fig. 197. Distribution of Scapania
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Fig. 201. Distribution of Scapania
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Fig. 205. Distribution of Scapania
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Fig. 209. Distribution of Scapania
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Fig. 213. Distribution of Scapania
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Fig. 217. Distribution of Tritomari
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Evaluating the distribution dataFol
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Fig. 224. Hornworts and Liverworts
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Signs used:Bolded names - species i
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Marchantiaceae50. VU - Bucegia roma
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Fig. 228. The distribution of zones
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Anthoceros communis Steph. = Phaeoc
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Blasia funckii Corda = Blasia pusil
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Cephalozia crassiflora Spruce = Cep
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Cephaloziella spinigera (Lindb.) Wa
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Diplomitrium lyellii (Hook.) Corda
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Gymnocolea inflata subsp. acutiloba
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Jungermannia acuta var. gypsacea Go
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Jungermannia dentata Raddi = Cephal
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Jungermannia limprichtii Lindb. = L
- Page 425 and 426:
Jungermannia scalaris var. denudata
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Leiocolea heterocolpos (Thed. ex C.
- Page 429 and 430:
Lophozia incisa var. inermis Müll.
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Marchantia polymorpha var. dilatata
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Mastigobryum deflexum (Mart.) Ångs
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Odontoschisma sphagni var. denudatu
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Porella platyphylla (L.) Pfeiff.Por
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Riccia maior Roth = Riccia glauca L
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Scapania heterophylla M. Howe = Sca
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Sphagnocetis communis var. macrior
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Mittheilungen des siebenbürgischen
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91. Borza A., 1922a. Schedae ad "Fl
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142. Boşcaiu N., Täuber F., Colde
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199. Coldea G., 1975. Étude phytos
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256. Csűrös Ş., Moldovan I., Cs
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317. Dihoru G., 1987b. Contribuţie
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367. Domin K., 1923. Grundzüge der
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426. Gergely I., 1969. Pajişti mez
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487. Györffy I., 1906c. Polytrichu
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540. Györffy I., 1943b. Erdélyi P
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598. Ionescu Venera, Cristurean I.,
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658. Lungu Lucia, 1967a. Contribuţ
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712. Mihai G., 1971e. Aspecte ale b
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767. Mihăilescu Simona, Ştefănu
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824. Mohan G., 1988c. Die Gattung T
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879. Niţu I.T, 1973. Solurile cu e
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941. Papp C., 1931b. O schiţă a v
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999. Papp C., Eftimie Elena, 1963.
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1059. Péterfi L.Ş., 1963. Flagela
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1119. Plămadă, Viţalariu, 1971.
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1178. Pop I., Hodişan I., 1958. Co
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1231. Prunescu-Arion Elena, Toniuc
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1285. Raţiu O., 1970. Consideraţi
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1342. Rüegsegger F., 1986. Frullan
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1396. Schiffner V., 1913b. Über ei
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1455. Söderström L., Urmi E., Vá
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1515. Ştefănuţ S., 2004d. The Ho
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1567. Ştefureac T., 1960b. Cercet
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1624. Ştefureac T., 1976c. Rolul b
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1675. Ştefureac T., Cristurean I.,
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1727. Tarnavschi I., Nedelcu G., 19
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1785. Wallfisch R., 2008. Contribut
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var. fragilis (A. Roth) Müll. Frib
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Riccardia incurvata Lindb. 28, 248,