The Hornwort and Liverwort Atlas of Romania

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Brook Valley, Valea Mare-Moldova Nouă Reserve, sub P. fabbroniana (Plămadă 1990:75); ♂ - Peşteracu Muscă, 14.08.2006, leg. Cogălniceanu Gina, det. Ştefănuţ S. [BUCA B3712].Semenic Mountain, Semenic, 25.07.1943, leg. Pop E. & Ciobanu I., sub P. fabbroniana (Ştefureac1948a:220); in peatbogs, sub P. fabbroniana (Pop E. 1960:363).Codru-Moma Mountains, Monesei Valley, Lungă Valley, Crişul Văratecului Valley, on wet soil(Mohan 1996:50; Ardelean 2006:106, Fig.23).Arad County, Cladova Brook’s springs, above Arăneag towards Cladovei Ridge and towardsHighiş, near the springs from Geinan above Hălmagiu, sub P. furcigera (Simonkai 1893:360).Zarand Mountains, Arăneag Valley, Highiş Hill, in beech forest, near the springs, sub P.fabbroniana fo. furcifera (Csűrös M. 1978:76);Alba County, Valea Lungă, on wet clayey soil, 24.04.1858, leg. & det. Barth J., sub P. calycina[SIB HF 1508](Fuss 1878:652; Drăgulescu 1983:57); Lal’s Valley, 450-630 m alt., incl. sub P.fabbroniana var. lorea (Papp 1944b:236; Papp 1945a:161); Izbucul Ponorului, sub P. fabbroniana (Boros& Vajda 1974:6); right slope of Ampoiului Valley, hornbeam grove, downstream Găureni, at 16 km toZlatna (Plămadă 1986a:59, Tab.1); Cheia Valley, 700 m alt., 31.07.1994, ass. Pellio-Cratoneuretumfilicini v.Dunk 1972 (Goia 1998:249, Tab.1).Cluj County, Cluj-Napoca, Sălicea Brook, leg. Péterfi M., sub P. fabbroniana (Györffy1924b:39); Southern Ciucea on Surduc Valley (Boros 1942a:34); Cluj-Napoca, Someş Valley (Soó1951:46; Boros 1958:7); Pleşca Valley, Cluj-Napoca, sub P. fabbroniana (Soó 1949:53; Boros 1958:7);Făget, Greben Valley, Feleac Hill, sub P. fabbroniana (Boros 1958:3,7); Cluj-Napoca Botanical Garden,sub P. fabbroniana, incl. fo. furcigera (Plămadă 1963:179; Plămadă 1969:170; Ştefureac 1982c:197,Tab.2); Cluj, on soil, 1961-1966, sub P. fabbroniana, incl. fo. furcigera (Plămadă 1967:470); Mera ForestBasin, on soil and wet stones, 1963-1965, sub P. fabbroniana, incl. fo. furcigera (Plămadă 1968:24);Someş Valley, in wet places, sub P. fabbroniana (Plămadă 1969:170); Turda’s Gorges (Plămadă & Goia1994:87).Bihor County, Crişul Repede Defile (Boros 1942a:31; Boros 1942e:154); Aştileu (Boros1942e:154); Tâlharului Cave (Boros 1942e:154); Crişul Repede Defile, on the edge of Lake from Peşteracu Apă and near the Waterfall, sub P. fabbroniana, ass. Glycerio-Sparganietum (Boşcaiu & Gergely1966:104); Victoria Spa - Oradea, 1948, det. Ştefureac T., sub P. fabbroniana (Olteanu-Cosma 1959:70);Vadu Crişului Cave, leg. Plămadă E. (Viehmann, Pleşa & Rusu 1964; Boşcaiu & Gergely 1966:104);Vadu Crişului Cave, 1968, leg. Neacşu M., det. Ştefureac T., sub P. fabbroniana (Ştefureac 1970h:482;Ştefureac 1985e:110,111, Tab.1).Bihor Mountains, Izbucul Ponorului, on edge of Cetăţilor Brook, 2-9.07.1961 (Páll 1963a:8);Cetatea Rădesei Cave, 1971 (Ştefureac 1985e:110,111, Tab.1); Galbena Valley, 750 m alt., 08.1999(Jakab 2000b:66); Someşul Cald Gorges (Goia & Mătase 2001:17); Cetăţile Ponorului, AB, 46°33'55"N /22°42'15"E, 1050 m alt., 3.09.2007, leg. & det. Ştefănuţ S. [BUCA B3869].Pellia epiphylla (L.) CordaSyn.:– Blasia epiphylla Fr.– Gymnomitrion epiphyllon Huebener– Jungermannia epiphylla L.– Marsilia epiphylla (L.) Lindb.– Pellia epiphylla var. undulata (Nees) Gottsche et al.First citation for Romania: Baumgarten 1846.Romania (Müller 1954:513).Moldova (Papp 1943:654; Plămadă 1968:23,24; Ştefureac & al. 1973:143).Muntenia (Mohan 1988b:112).Transylvania (Pax 1908:128).Ecology: epigaeic, acidophytic, hydro-hygrophytic, sciophytic; circumpolar; mountain; Hth.Herbaria and Exsiccatae: BUCA, PHHM, SIBDistribution in Europe: Fo, No, Dk, Se, Fi, Ie, Br, Fr, Nl, Be, De, Po, Ch, At, Cz, Sk, Hu, Pt, Es, Co, Sa, Sc, It, Si, Hr, Sb, Me,Ba, Mk, Ro, Bg, Gr, Ct, Ee, Lv, Lt, By, Ua, Rn, Rw, Rc, Re, Az, Ma.Distribution in Romania:Maramureş County, Poienile de sub Munte (Warnstorf 1895b:143).Rodna Mountains, (Pax 1908:128; Plămadă 1968:23,24); Iezerul Pietrosului, MM, 47°35'51"N /24°38'48"E, 1840 m alt., 28.08.1999, leg. & det. Ştefănuţ S. [BUCA B2358].207
Ţibleş Mountains, Piciorul Ţiglei, Piciorul Mesteacănului, Ţibleş Valley, Suplai Commune, 1984-1985 (Mohan 1988a:16).Botoşani County, Alba Forest – Ursoaiei Valley, on the edge of the brooks, on wet soil (Mihai1968c:391).Suceava County, Argestrul, 16.05.1910 (Lilienfeldówna 1911:731,734); Bistriţa Aurie Valley,between Iacobeni and Ciocăneşti, north-wester slope of Brezuţa Peak, the edge of Suhărzelul andSuhărzelul Mare Brooks (Pascal 1969:107); Cârlibaba Brook Valley, Iedu, Deaca, Liţu, Ţibău, Ciocăneşti(Pascal & Mititelu 1971; Ştefureac & Pascal 1981:477); Adam and Eva Mountains, Pojorâta Commune,in marshy places (Mihai & Pascal 1973b:156,161); Ouşorul Mountain, on soil, 1971 (Mihai1974b:318,320); the bog from Neagra Broştenilor Meadow, ass. Cratoneuretum filicino-commutati (Kuhn1937) Oberd. 1977 (Lungu 1981b: 417, Tab.2); Cristişor-Neagra Broştenilor, Borcut Meadow, ineutrophic fen, incl. sub fo. undulata Nees (Lungu 1973b:72; Lungu 1981c:90).Rarău Mountain, Codrul Secular Slătioara, 800-1340 m alt., 17.07.1937, incl. sub var. and fo.undulata (Ştefureac 1941:1154,1242,1278, Tabs.4,5,7,8,9; Ştefureac 1969a: tab 16; Papp 1969b: 188);Plaiul Todirescu-Slătioara Reserve (Ştefureac & Raclaru 1978:196).Bârgău Mountains, Cucureasa Reserve, Coşna Forest Hut, 930-1230 m alt., 1967, 1969(Ştefureac, Mihai & Pascal 1976:35,36); Cârţibavu Mare Valley, Sângeorz-Băi, Poiana Ilvei, Secu Brook,Ilva Mică Commune, Ilva Mare Commune, Tureac Brook, Leşului Valley, Măgura Calului, PiatraFântânele, 1983-1984 (Mohan & Hodoroga 1986:211).Bistriţa-Năsăud County, Piatra Corbului, Monari, Simioneşti, 1980-1981 (Mohan & Hodoroga1983:51; Mohan 1990a:24).Iaşi County, Bârnova Forest (Papp 1939a:340); Repedea-Bârnova Forest Massif, on soil umed(Papp & Eftimie 1963:108; Plămadă 1968:23,24).Neamţ County, Cuejdiu Valley, incl. fo. undulata (Papp 1932:462,464); Doamna, Gârcina,Cuejdiu (Papp 1934a:193; Lungu 1963a:241); Neamţ Monastery, on clayey soil, 07-08.1921,1923, leg.Brândză M., det. Ştefureac T. (Ştefureac 1942a:4; Plămadă 1968:23,24); Tarcău-Grinduş Mountain (Papp1942b:524); Cernegura Mountain, Sterghinosu Brook Valley, 400-550 m alt. (Lungu 1963a:242).Ceahlău Mountain, Durău Valley, in common spruce forest, on wet soil (Mihai 1984a:89).Harghita County, Vărşag, on the marshy edge of Târnava River, 1958-1962 (Páll 1963b:17);around Luci Reserve, in common spruce forest (Mititelu & Elekes E.S. 1984:124).Harghita Mountains, Minei Valley – riverhead, 1200 m alt., 11.07.2000, on soil, leg. Bálint L. &Csízi K. (Orbán & Bálint 2002:60; Bálint & Orbán 2003:14); Groapa Mare, 1300 m alt., 16.08.2001, leg.Bálint L. & Csízi K. (Bálint & Orbán 2003:14).Giurgeu Mountains, Bicaz Gorges, on rocks, 1957-1960, sub fo. undulata Nees (Papp & al.1962:144,147,149).Nemira Mountains, Şandru, on wet soil, 1970-1977 (Mihai & Barabaş V. 1978:115); Cheşcheş,Şandru (Barabaş V. 1982b:70,72, Tabs.20,23).Bacău County, Hoţului Brook, at west to Hârja, incl. fo. undulata (Papp 1957:284); Gura Dobrilor(Eftimie 1972:150).Ciucaş Mountains, Roşu Brook Valley, Zăgan Mountain, Albele Culm, Piscul Gropşoarele, RoşuMountain (Mohan 1990b:44).Prahova County, Doftana Valley [PHHM 7763/3873](Mohan 1971a:8; Mohan 1971b:45; Mohan1990a:24); Teleajen Valley, Negraşului Valley, Ordojoaia Valley, sub P. endiviifolia (Mohan 1990a:24).Bucegi Massif, Ialomiţa Valley, sub P. endiviifolia (Mohan 1990a:24).Piatra Craiului Mountains, Marele Grohotiş, AG, 45°30'36"N / 25°11'45"E, 1650 m alt.,8.07.1998, leg. & det. Ştefănuţ S. [BUCA B2605] (Ştefănuţ & Pop O.G. 2006:79); Marele Grohotiş, AG,45°30'34"N / 25°11'46"E, 1650 m alt., 8.07.2003, leg. & det. Ştefănuţ S. [BUCA B2633] (Ştefănuţ & PopO.G. 2006:79); Pârâul Rece, AG, 45°30'38"N / 25°11'34"E, 1500 m alt., 8.07.2003, leg. & det. ŞtefănuţS. [BUCA B2631] (Ştefănuţ & Pop O.G. 2006:79).Leaota Mountain, Mitarca Valley, 1300-1600 m alt. (Papp & Diaconescu 1968:386,387).Iezer-Păpuşa Mountains, Iezer Valley, Bătrâna Valley, Dâmboviţa Valley, sub P. endiviifolia(Mohan 1990a:24).Argeş County, ♂ - Boteni, 03.1893, leg. Vlădescu M. (Radian 1915:197).Făgăraş Depression, ♀♂ - Mlaca Mică from Arpaşu de Sus, 45°42'47"N / 24°29'25"E, 555 m alt.,30.10.2004, leg. & det. Ştefănuţ S. [BUCA B3312].208
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SORIN ŞTEFĂNUŢThe Hornwort and L
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CONTENTINTRODUCTION ...............
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The beginning of the 20 th century
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asin of the Arieş river. Bryotaxon
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Abbreviations used in the text† -
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The presentation of the methodology
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Fig. 3. The GEOCOD mapping system.
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THE HORNWORT AND LIVERWORT SPECIES
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THE HORNWORT AND LIVERWORT CLASSIFI
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Suborder Lepidoziineae R.M. Schust.
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46. Cephalozia pleniceps (Austin) L
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147. Pallavicinia lyellii (Hook.) C
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13 Corsinia coriandrina (Spreng.) L
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11. Porella obtusata (Taylor) Trevi
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34Phaeoceros carolinianus (Michx.)
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Făgăraş Mountains, Cârţişoara
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Cluj County, Someşul Rece Valley a
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Parâng Mountains, Repedea Valley,
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Argeş County, Rucăr (Loitlesberge
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Cindrel Mountains, Cindrel, 2200 m
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Parâng Mountains, Latoriţa Valley
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Athalamia hyalina (Sommerf.) S. Hat
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Barbilophozia barbata (Schmidel ex
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Bârgău Mountains, Cârţibavu Mar
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First citation for Romania: Schur 1
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- Pleuroschisma triangulare (Schlei
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and Băiuţ, common spruce forest,
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Distribution in Europe: Sj, Is, Fo,
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Poiana Stampei Peatbog, SV, 47°20'
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alt., 8.08.2003, leg. Pop O., det.
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leg. & det. Ştefureac T., ass. Buc
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Rodna Mountains, Puzdra Mountain, 1
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Alba County, Vârciorog Valley, 780
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Bârgău Mountains, Cucureasa Reser
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Suceava County, Tinovul Mare de la
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Suceava County, Rodna Nouă, on Per
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2001:74,75); Arieşul Mare Basin, o
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Ţibleş Mountains, Mesteacănul Va
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Cindrel Mountains, Găuşoara, leg.
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Parâng Mountains, the edge of Câl
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& det. Ştefănuţ S. [BUCA B3684];
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Alba County, at 20,5 km to Zlatna,
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Cephaloziella spinigera (Lindb.) Wa
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First citation for Romania: Péterf
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Iezer-Păpuşa Mountains, Iezer Val
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- Cladopodiella fluitans var. gigan
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Caraş-Severin County, Cerna Valley
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(Mititelu, Barabaş N. & al. 1968:1
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1885:30; Ştefureac 1949a:287,288;
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104Conocephalum salebrosum Szweyk.,
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Retezat Mountains, Retezat, sub Jun
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Cindrel Mountains, Cânaia, 1750 m
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Iaşi County, Sineşti Forest (Papp
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[BUCA B2793], on rotten beech trunk
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Orthodicrano montani-Hypnetum filif
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Distribution in Romania:Maramureş
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- Gymnocolea inflata var. laxa (Nee
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Ciucaş Mountains, Caprei Brook Val
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Giumalău Mountain, Giumalău Secul
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- Haplozia riparia var. rivularis B
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- Jungermannia karl-muelleri auct.,
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Jungermannia gracillima Sm.Syn.:- A
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Mountain, PH, 45°24'40"N / 25°28'
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Retezat Mountains, Rovine Brook, on
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Romania, alt. 600-2280 m (Váňa 19
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- Solenostoma subellipticum (Lindb.
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Distribution in Romania:Bârgău Mo
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13.07.1946, sub Lophozia alpestris
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Distribution in Romania:Maramureş
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Rodna Mountains, Lala Valley, 1200
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Ştefănuţ S. [BUCA B2574, B2580,
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Dealul Colbului and Pietrele Doamne
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150var. rivularis (Raddi) Warnst.Sy
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Penteleu Massif, fer - Roşu Lake,
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Satu Mare County, Noroieni, Ghiarma
Page 157 and 158: Lophozia ascendens (Warnst.) R.M. S
Page 159 and 160: - Jungermannia intermedia Lindenb.-
Page 161 and 162: Căliman Mountains, on humus and ro
Page 163 and 164: Hăşmaş Mountains, Hăşmaş, sub
Page 165 and 166: Herbaria and Exsiccate: BUCADistrib
Page 167 and 168: peatbog on the border of Câlcescu
Page 169 and 170: ? - Iezer-Păpuşa Mountains, Iezer
Page 171 and 172: Făgăraş Mountains, s - above Avr
Page 173 and 174: Nemira Mountains, on edge of Nemira
Page 175 and 176: 2003:7); Măgura Cisnădiei [SIB HL
Page 177 and 178: Fuss 1854:20; Ştefureac 1949a:288;
Page 179 and 180: - Nardia brevissima (Dumort.) Lindb
Page 181 and 182: 1919:295); Ciortea Mountain, on wet
Page 183 and 184: - Sarcoscyphus sphacelatus var. med
Page 185 and 186: Siriu Mountain, Poiana din Fundul M
Page 187 and 188: Iaşi County, below Repedea, 7.10.1
Page 189 and 190: Valley, Arieşul Mic - upstream min
Page 191 and 192: Alba County, Valea Lungă, 10.02.18
Page 193 and 194: Rarău Massif, between Dealul Colbu
Page 195 and 196: Harghita County, Harghita Spa, on r
Page 197 and 198: Căliman Mountains, in alpine zone,
Page 199 and 200: Stânişoara Mountains, Suha Mare B
Page 201 and 202: Ecology: humicolous, acidophytic, h
Page 203 and 204: Bacău County, Slănic Brook (Papp
Page 205 and 206: Rarău Mountain, Codrul Secular Sl
Page 207: 1915:197,198); Cândeşti, 07.1891,
Page 211 and 212: Ceahlău Mountain, Durău Valley, i
Page 213 and 214: major (Ştefureac 1962b:69); Benea-
Page 215 and 216: Ciucaş Mountains, Berii Brook Vall
Page 217 and 218: Zlătuia Valley, 1100 m alt., 7.09.
Page 219 and 220: - Plagiochila dillenii (Taylor) Tay
Page 221 and 222: Wallfisch R. (Dihoru G. & al. 2003:
Page 223 and 224: Ştefănuţ M. 2003:79); Iezerul Pi
Page 225 and 226: utabuli-Mnietum cuspidati (Felföld
Page 227 and 228: Iaşi County, Mârzeşti Forest, Bu
Page 229 and 230: 3.04.2004, leg. Atanasiu Paulina, d
Page 231 and 232: Arad County, Săvârşin, sub Madot
Page 233 and 234: Preissia quadrata (Scop.) NeesSyn.:
Page 235 and 236: Diana, BV, 1100 m alt., 22.08.2001,
Page 237 and 238: Rodna Mountains (Pax 1908:130); Pie
Page 239 and 240: Descut, 15.08.2001, Podul de Piatr
Page 241 and 242: Bârgău Mountains, Cucureasa Reser
Page 243 and 244: B3521, B3522] (Ştefănuţ 2006b:34
Page 245 and 246: 16,20.08.1994, ass. Pterigynandretu
Page 247 and 248: - Marchantia crinita Michx.- Marcha
Page 249 and 250: Giumalău Mountain, Giumalău Secul
Page 251 and 252: Riccardia multifida (L.) GraySyn.:-
Page 253 and 254: Penteleu Massif, the left side of C
Page 255 and 256: Herbaria and Exsiccate: SIBDistribu
Page 257 and 258: 558,560; Papp 1939a:358; Papp & Eft
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Riccia glauca L. var. glaucaSyn.:-
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Distribution in Europe: Is, Fo, No,
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Bihor County, Tărian, sub Riccia (
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1960:1113); upper side of Mălăie
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Scapania brevicaulis TaylorSyn.:- S
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Suceava County, Cristişor-Neagra B
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Piatra Mare Mountain, 1700 m alt.,
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- Scapania amurensis Warnst.- Scapa
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Mureş County, Sighişoara - Fuchsl
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Căliman Mountains, the peatbogs fr
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- Scapania heterophylla M. Howe- Sc
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Slăvei and Muchia Ascuţită, 2000
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Distribution in Europe: Ie, Br, Fr,
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Nemira Mountains, Nemira, on humus,
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- Sphenolobus medelpadicus (Arnell)
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Parâng Mountains, Latoriţa Valley
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Fig. 9. Distribution of Phaeoceros
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Fig. 13. Distribution of Anastrophy
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Fig. 17. Distribution of Anthelia j
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Fig. 21. Distribution of Asterella
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Fig. 25. Distribution of Barbilopho
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Fig. 29. Distribution of Barbilopho
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Fig. 33. Distribution of Bazzania t
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Fig. 37. Distribution of Calypogeia
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Fig. 41. Distribution of Calypogeia
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Fig. 45. Distribution of Cephalozia
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Fig. 49. Distribution of Cephalozia
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Fig. 53. Distribution of Cephalozie
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Page 317 and 318:
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Fig. 65. Distribution of Cololejeun
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Fig. 69. Distribution of Corsinia c
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Fig. 73. Distribution of Fossombron
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Fig. 77. Distribution of Frullania
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Fig. 81. Distribution of Gymnocolea
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Fig. 85. Distribution of Harpanthus
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Fig. 89. Distribution of Jungermann
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Page 335 and 336:
Page 337 and 338:
Fig. 101. Distribution of Jungerman
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Fig. 105. Distribution of Leiocolea
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Fig. 109. Distribution of Lejeunea
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Fig. 113. Distribution of Lophocole
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Fig. 117. Distribution of Lophozia
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Fig. 121. Distribution of Lophozia
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Fig. 125. Distribution of Lunularia
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Fig. 129. Distribution of Marchanti
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Fig. 133. Distribution of Marsupell
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Fig. 137. Distribution of Marsupell
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Fig. 141. Distribution of Moerckia
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Fig. 145. Distribution of Nardia ge
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Fig. 149. Distribution of Odontosch
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Fig. 153. Distribution of Pedinophy
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Fig. 157. Distribution of Peltolepi
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Fig. 161. Distribution of Porella a
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Fig. 165. Distribution of Preissia
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Fig. 169. Distribution of Radula li
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Fig. 173. Distribution of Riccardia
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Fig. 177. Distribution of Riccia ca
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Fig. 181. Distribution of Riccia cr
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Fig. 185. Distribution of Riccia go
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Fig. 189. Distribution of Ricciocar
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Fig. 193. Distribution of Scapania
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Fig. 217. Distribution of Tritomari
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Evaluating the distribution dataFol
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Fig. 224. Hornworts and Liverworts
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Signs used:Bolded names - species i
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Marchantiaceae50. VU - Bucegia roma
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Fig. 228. The distribution of zones
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Anthoceros communis Steph. = Phaeoc
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Blasia funckii Corda = Blasia pusil
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Cephalozia crassiflora Spruce = Cep
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Cephaloziella spinigera (Lindb.) Wa
Page 415 and 416:
Diplomitrium lyellii (Hook.) Corda
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Gymnocolea inflata subsp. acutiloba
Page 419 and 420:
Jungermannia acuta var. gypsacea Go
Page 421 and 422:
Jungermannia dentata Raddi = Cephal
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Jungermannia limprichtii Lindb. = L
Page 425 and 426:
Jungermannia scalaris var. denudata
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Leiocolea heterocolpos (Thed. ex C.
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Lophozia incisa var. inermis Müll.
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Marchantia polymorpha var. dilatata
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Mastigobryum deflexum (Mart.) Ångs
Page 435 and 436:
Odontoschisma sphagni var. denudatu
Page 437 and 438:
Porella platyphylla (L.) Pfeiff.Por
Page 439 and 440:
Riccia maior Roth = Riccia glauca L
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Scapania heterophylla M. Howe = Sca
Page 443 and 444:
Sphagnocetis communis var. macrior
Page 445 and 446:
Mittheilungen des siebenbürgischen
Page 447 and 448:
91. Borza A., 1922a. Schedae ad "Fl
Page 449 and 450:
142. Boşcaiu N., Täuber F., Colde
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199. Coldea G., 1975. Étude phytos
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256. Csűrös Ş., Moldovan I., Cs
Page 455 and 456:
317. Dihoru G., 1987b. Contribuţie
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367. Domin K., 1923. Grundzüge der
Page 459 and 460:
426. Gergely I., 1969. Pajişti mez
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487. Györffy I., 1906c. Polytrichu
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540. Györffy I., 1943b. Erdélyi P
Page 465 and 466:
598. Ionescu Venera, Cristurean I.,
Page 467 and 468:
658. Lungu Lucia, 1967a. Contribuţ
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712. Mihai G., 1971e. Aspecte ale b
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The Hornwort and Liverwort Atlas of Romania

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