Zar<strong>and</strong> Mountains, Milova Valley, in beech forest <strong>and</strong> oak grove, Jernova Valley (Csűrös M.1978:77; Ardelean 2006:109, Fig.26).Codru-Moma Mountains, Monesei Valley, Lungă Valley, Crişul Văratecului Valley, BrihenilorValley, on wet soil (Mohan 1996:50; Ardelean 2006:109, Fig.26).Cluj County, Dobrin Mountain (Péterfi M. 1918:61); Cluj, leg. & det. Péterfi M. (Györffy1924b:38); ♀, ♂ - Cluj-Napoca Botanical Garden, 350 m alt., 18.06.1915, leg. & det. Péterfi M. [FRE705b] (Borza 1935:64; Boros 1958:5,7; Borza & Nyárády E.I. 1940:11); Turda’s Gorges, on wet soil, onstone, 400-450 m alt., 07.1929 (Papp 1946a:13); ♀ - Cluj-Napoca, on walls, 26.06.1946, leg. & det.Morariu I. [PHHM 6207327, 6207335, 6207336]; Cluj-Napoca Botanical Garden (Györffy1947a:173,175,178-180; Boros 1958:5,7); Cluj-Napoca Botanical Garden (Boros 1958:5,7; Plămadă1963:179,189; Plămadă 1969:170; Ştefureac 1982c:197,200, Tab.2); the peatbogs from spring <strong>and</strong> uppercourse <strong>of</strong> Someşul Rece, incl. Muntele Mare <strong>and</strong> Dobrin (Pop E. 1960:397); the peatbog from the left side<strong>of</strong> Izbuc (Pop E. 1960:375); the peatbogs <strong>and</strong> marshes from upper course <strong>of</strong> Someşul Cald <strong>and</strong> Bălăceşti-Călăţele zone (Pop E. 1960:385); Puşcaşului Valley <strong>and</strong> in Mera Forest, on wet soil, 1963-1965 (Plămadă1968:24); Cluj, Someşul Valley at Gat, Petöfi A. France Street (Plămadă 1969:170); Someşul Cald Valley(Boros & Vajda 1974:8); Turzii Gorges (Plămadă & Goia 1994:87); Valea Popii (Goia 2002:115, Tab.1).Bihor County, Vadu Crişului Cave (Boros 1942e:154); Crişul Repede Defile, near waterfall, onsoil (Viehmann, Pleşa & Rusu 1964; Boşcaiu & Gergely 1966:103); Mădăraş, Marţihaz, Tulca, infountains, det. Ştefureac T. (Pop I. 1968:69); Ponor Hill, 450-590 m alt., 11.05.1967, ass. Chrysosplenio-Cardaminetum amarae (Tx. 1937) Mass 1959 (Coldea 1978:127, Tab.1; S<strong>and</strong>a & Popescu A. 1991a:42,Tab.1); Iadului Valley, Stâna de Vale, peatbogs, associations with Eriophorum <strong>and</strong> Sphagnum (Raţiu O.& al. 1982:9); Lacul Vărgat, at 4 km south-east to Săcueni, floating reed islet, 145 m alt., 05-09.1980,det. Dihoru G., ass. Carici-Menyanthetum Soó (1938) 1955, ass. <strong>The</strong>lypteridi-Alnetum Klika 1940, ass.Calamagrosti-Salicetum cinerae Soó & Zólyomi 1955, ass. Salici cinerae-Sphagnetum recurvi (Zólyomi1931) Soó 1954 (Karácsonyi 1982:30,33,35, Tabs.1,2); Someşul Cald Gorges, on soil <strong>and</strong> rocks (Goia &Mătase 2001:16).Pădurea Craiului Mountains, Ieduţului Valley, ass. Carici stellulatae-Sphagnetum (recurvipalustris)(Soó 1934) Soó 1954 (Raţiu O. & Boşcaiu 1967:37).Bihor Mountains, Izbucul Ponorului, in wet places, 2-9.07.1961 (Páll 1963a:8); Sighiştel Valley,ass. Carpino-Fagetum Paucă 1941 (Pop I. & Hodişan I. 1969:36); Ponor’s Spring, 1000 m alt., 08.1999(Jakab 2000a:93; Jakab 2000b:67).Satu Mare County, † - Ecedea marsh, near Berveni, 1859, leg. & det. Pokorny A. (Pokorny1860:286; Pop E. 1957b:174; Pop E. 1960:144).subsp. montivagans Bischl. et BoisselierSyn.:– Marchantia alpestris (Nees) Burgeff– Marchantia polymorpha var. alpestris NeesFirst citation for <strong>Romania</strong>: Schur 1852.Ecology: epigaeic, meso-hygrophytic, euryion, photo-sciophytic; subalpine-alpine, circumpolar, Hth.Herbaria <strong>and</strong> Exsiccate: BUCA, SIBDistribution in Europe: Sj, Is, Fo, No, Dk, Se, Fi, Ie, Br, Fr, Ad, De, Po, Ch, At, Cz, Sk, Es, It, Si, Hr, Sb, Me, Ro, Bg, Gr, Ee,Ua, Nz, Fl, Rn, Re, Ma, Cn.Distribution in <strong>Romania</strong>:Postăvaru Mountain, 4.07.1880, leg. & det. Fuss M., sub M. alpestris [SIB HF 1487] (Drăgulescu1983:56), sub var. alpestris (Schiffner 1914:303).Bucegi Massif, Bucegi, leg. & det. Fuss M., sub var. alpestris (Fuss 1878:651); Bucşoiu, sub var.alpestris (Degen 1914:216); Babele Mountain, 45°24'42"N / 25°28'37"E,. 2170 m alt, 7.09.2006, leg. &det. Ştefănuţ S. [BUCA B3500]; Babele Mountain, PH, 45°24'49"N / 25°28'43"E, 2220 m alt., 7.09.2006,leg. & det. Ştefănuţ S. [BUCA B3561]; ♀ - Caraiman Mountain, PH, 45°24'56"N / 25°29'02"E, 2330 malt., 20.07.2007, leg. & det. Ştefănuţ S. [BUCA B3913]; Babele Valley, PH, 45°24'43"N / 25°28'44"E,2150 m alt., 14.07.2008, leg. & det. Ştefănuţ S. [BUCA B3912].Făgăraş Mountains, Arpaş Mountain, 1849-1850, leg. & det. Schur F., sub var. alpestris [SIBHK](Schur 1852:92; Heufler 1853:64,65; Schur 1866:873; Fuss 1878:651; Ştefureac 1949a:287,288;Drăgulescu 2003:7); ♀, ♂ - Arpaş Valley, common spruce forest, sub var. alpestris (Heufler 1853:52-53;175
Fuss 1854:20; Ştefureac 1949a:288; Drăgulescu 2003:7); fer - Bâlea Lake, leg. Zsák Z., sub var. alpestris(Schiffner 1914:303; Ştefureac 1949a:288; Drăgulescu 2003:7).subsp. ruderalis Bischl. et BoisselierSyn.:– Marchantia aquatica (Nees) Burgeff– Marchantia polymorpha fo. aquatica Nees– Marchantia polymorpha var. aquatica NeesFirst citation for <strong>Romania</strong>: Matouschek 1903b.Moldova, sub fo. aquatica Nees (Papp 1943:650).Muntenia, sub fo. aquatica Nees (Mohan 1988b:116).Ecology: epigaeic, meso-xerophytic, euryion, photophytic; circumpolar, Hth.Herbaria <strong>and</strong> Exsiccate: BUCADistribution in Europe: Sj, Is, Fo, No, Dk, Se, Fi, Ie, Br, Fr, Nl, Be, De, Po, Ch, At, Cz, Sk, Hu, Pt, Es, Co, Sa, Sc, It, Si, Hr,Sb, Me, Ba, Ro, Al, Gr, Ct, Tu, Ee, Lv, Lt, By, Md, Ua, Cm, Nz, Fl, Rw, Rc, Re, Ca, Az, Ma, Cn.Distribution in <strong>Romania</strong>:Suceava County, Siminicea, 08.1892, leg. Vlădescu M., sub fo. aquatica (Radian 1915:194);Camionca Valley, below Chicera Hill Peak, 1300 m alt., 1-4.09.1953, sub var. aquatica (Ştefureac1956a:255); Drăgoiasa Fen, 1020-1030 m alt., 1959, leg. Ştefureac T., sub fo. aquatica (Ştefureac1962b:70,71; Ştefureac 1967b:308); Adam <strong>and</strong> Eva Mountains, Pojorâta Commune, in marshy places, inhay-fields, on soil, sub var. aquatica (Mihai & Pascal 1973b:156,161); Cristişor-Neagra Broştenilor,Borcut Meadow, ditch edge, sub var. aquatica (Lungu 1973b:72; Lungu 1981c:94).Rarău Massif, Codrul Secular Slătioara, sub fo. aquatica (Papp 1969b:188).Hăşmaş Mountains, Hăşmaş, sub var. aquatica (Schiffner 1914:303).Harghita County, Rotund Mountain between Borsec <strong>and</strong> Topliţa, sub var. aquatica (Matouschek1903b:160); the peatbog from Jigodin, below Cetăţii Hill (“Várhegy”), det. Ştefureac T., sub var.aquatica (Pop E.1960:155); the peatbog from Ciuc Basin, sub var. aquatica (Pop E.1960:168).Neamţ County, Văratec, 21,25.08.1914, leg. Săvulescu T., sub fo. aquatica (Radian 1915:195);Tâmpeşti, sub fo. aquatica Nees (Papp 1925c:155); Peste Recea Hill, in forest, sub fo. aquatica (Papp1947b:185,186,190).Bacău County, Ceardac on Slănic, 05.1892, leg. Vlădescu M., sub fo. aquatica (Radian1915:194,195; Papp 1957:284); Ruginii Brook (Eftimie 1972:152); Slănic Moldova, small fens on hillsnear Slănic Valley, 07.1907, leg. Brândză M., det. Ştefureac T., sub fo. aquatica (Ştefureac 1942a:3; Papp1957:284).Tulcea County, near Letea Village, on floating reed islet, sub M. polymorpha (Tarnavschi &Nedelcu 1970:172).Siriu Mountain, Lacul dintre Pietre, sub fo. aquatica Nees (Dihoru G. 1975:24).Braşov County, Predeal, 2.06.1883, leg. Brândză D., sub fo. aquatica (Radian 1915:194,195); ♀ -Făgăraş railway station, 440 m alt., 25.08.2005, leg. & det. Ştefănuţ S. [BUCA B3201], ♂ - [BUCAB3202], ♀ - 30.10.2004, leg. & det. Ştefănuţ S. [BUCA B3322], ♀ - 3.04.2006, leg. & det. Ştefănuţ S.[BUCA B3388], ♂ - - 3.04.2006, leg. & det. Ştefănuţ S. [BUCA B3389]; Făgăraş railway station, 440 malt., 1.08.2006, leg. & det. Ştefănuţ S. [BUCA B3543].Prahova County, Sinaia railway station, 3.04.2006, leg. & det. Ştefănuţ S. [BUCA B3390].Piatra Craiului National Park, Great Gorges <strong>of</strong> Dâmboviţa River, AG, 45°23'19"N / 25°11'45"E,800 m alt., 7.07.1998, leg. & det. Ştefănuţ S. [BUCA B2901] (Ştefănuţ & Pop O.G. 2006:78); belowGar<strong>of</strong>iţa Pietrei Craiului Cahlet, AG, 45°30'45"N / 25°10'12"E, 1030 m alt., 26.08.2004, leg. OneteMarilena, det. Ştefănuţ S. [BUCA B3099] (Ştefănuţ & Pop O.G. 2006:78).Leaota Mountain, Vaca Valley, 900 m alt., Mitarca Valley, 1300-1500 m alt., sub fo. aquaticaNees (Papp & Diaconescu 1968:387).Bucharest, ♂ - Cişmigiu Park, 3.04.2004, leg. & det. Ştefănuţ S. [BUCA B2937], ♂ - 6.07.2005,leg. & det. Ştefănuţ S. [BUCA B3178]; Grădina Botanică, det. Ştefănuţ S. (!).Făgăraş Mountains, Pojorta Valley, 800-1000 m alt., 18.09.2001, leg. & det. Ştefănuţ S. [BUCAB2554] (Ştefănuţ 2002:190).Vâlcea County, Bistra Monastery, 16.06.1887, leg. Brândză D., sub fo. aquatica (Radian1915:194,195); Puturoasă Valley at Călimăneşti, 08.1908, leg. Colceag N., sub fo. aquatica (Radian1915:194,195).176
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SORIN ŞTEFĂNUŢThe Hornwort and L
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CONTENTINTRODUCTION ...............
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The beginning of the 20 th century
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asin of the Arieş river. Bryotaxon
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Abbreviations used in the text† -
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The presentation of the methodology
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Fig. 3. The GEOCOD mapping system.
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THE HORNWORT AND LIVERWORT SPECIES
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THE HORNWORT AND LIVERWORT CLASSIFI
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Suborder Lepidoziineae R.M. Schust.
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46. Cephalozia pleniceps (Austin) L
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147. Pallavicinia lyellii (Hook.) C
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13 Corsinia coriandrina (Spreng.) L
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11. Porella obtusata (Taylor) Trevi
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34Phaeoceros carolinianus (Michx.)
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Făgăraş Mountains, Cârţişoara
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Cluj County, Someşul Rece Valley a
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Parâng Mountains, Repedea Valley,
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Argeş County, Rucăr (Loitlesberge
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Cindrel Mountains, Cindrel, 2200 m
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Parâng Mountains, Latoriţa Valley
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Athalamia hyalina (Sommerf.) S. Hat
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Barbilophozia barbata (Schmidel ex
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Bârgău Mountains, Cârţibavu Mar
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First citation for Romania: Schur 1
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- Pleuroschisma triangulare (Schlei
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and Băiuţ, common spruce forest,
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Distribution in Europe: Sj, Is, Fo,
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Poiana Stampei Peatbog, SV, 47°20'
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alt., 8.08.2003, leg. Pop O., det.
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leg. & det. Ştefureac T., ass. Buc
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Rodna Mountains, Puzdra Mountain, 1
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Alba County, Vârciorog Valley, 780
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Bârgău Mountains, Cucureasa Reser
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Suceava County, Tinovul Mare de la
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Suceava County, Rodna Nouă, on Per
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2001:74,75); Arieşul Mare Basin, o
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Ţibleş Mountains, Mesteacănul Va
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Cindrel Mountains, Găuşoara, leg.
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Parâng Mountains, the edge of Câl
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& det. Ştefănuţ S. [BUCA B3684];
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Alba County, at 20,5 km to Zlatna,
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Cephaloziella spinigera (Lindb.) Wa
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First citation for Romania: Péterf
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Iezer-Păpuşa Mountains, Iezer Val
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- Cladopodiella fluitans var. gigan
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Caraş-Severin County, Cerna Valley
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(Mititelu, Barabaş N. & al. 1968:1
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1885:30; Ştefureac 1949a:287,288;
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104Conocephalum salebrosum Szweyk.,
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Retezat Mountains, Retezat, sub Jun
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Cindrel Mountains, Cânaia, 1750 m
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Iaşi County, Sineşti Forest (Papp
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[BUCA B2793], on rotten beech trunk
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Orthodicrano montani-Hypnetum filif
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Distribution in Romania:Maramureş
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- Gymnocolea inflata var. laxa (Nee
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Ciucaş Mountains, Caprei Brook Val
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Giumalău Mountain, Giumalău Secul
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Iaşi County, Mârzeşti Forest, Bu
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3.04.2004, leg. Atanasiu Paulina, d
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Arad County, Săvârşin, sub Madot
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Preissia quadrata (Scop.) NeesSyn.:
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Diana, BV, 1100 m alt., 22.08.2001,
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Rodna Mountains (Pax 1908:130); Pie
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Descut, 15.08.2001, Podul de Piatr
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Bârgău Mountains, Cucureasa Reser
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B3521, B3522] (Ştefănuţ 2006b:34
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16,20.08.1994, ass. Pterigynandretu
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- Marchantia crinita Michx.- Marcha
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Giumalău Mountain, Giumalău Secul
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Riccardia multifida (L.) GraySyn.:-
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Penteleu Massif, the left side of C
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Herbaria and Exsiccate: SIBDistribu
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558,560; Papp 1939a:358; Papp & Eft
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Riccia glauca L. var. glaucaSyn.:-
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Distribution in Europe: Is, Fo, No,
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Bihor County, Tărian, sub Riccia (
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1960:1113); upper side of Mălăie
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Scapania brevicaulis TaylorSyn.:- S
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Suceava County, Cristişor-Neagra B
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Piatra Mare Mountain, 1700 m alt.,
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- Scapania amurensis Warnst.- Scapa
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Mureş County, Sighişoara - Fuchsl
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Căliman Mountains, the peatbogs fr
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- Scapania heterophylla M. Howe- Sc
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Slăvei and Muchia Ascuţită, 2000
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Distribution in Europe: Ie, Br, Fr,
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Nemira Mountains, Nemira, on humus,
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- Sphenolobus medelpadicus (Arnell)
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Parâng Mountains, Latoriţa Valley
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Fig. 9. Distribution of Phaeoceros
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Fig. 13. Distribution of Anastrophy
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Fig. 17. Distribution of Anthelia j
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Fig. 21. Distribution of Asterella
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Fig. 25. Distribution of Barbilopho
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Fig. 29. Distribution of Barbilopho
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Fig. 33. Distribution of Bazzania t
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Fig. 37. Distribution of Calypogeia
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Fig. 41. Distribution of Calypogeia
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Fig. 45. Distribution of Cephalozia
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Fig. 49. Distribution of Cephalozia
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Fig. 53. Distribution of Cephalozie
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Fig. 57. Distribution of Cephalozie
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Fig. 61. Distribution of Cephalozie
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Fig. 65. Distribution of Cololejeun
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Fig. 69. Distribution of Corsinia c
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Fig. 73. Distribution of Fossombron
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Fig. 77. Distribution of Frullania
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Fig. 81. Distribution of Gymnocolea
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Fig. 85. Distribution of Harpanthus
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Fig. 89. Distribution of Jungermann
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Fig. 93. Distribution of Jungermann
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Fig. 97. Distribution of Jungermann
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Fig. 101. Distribution of Jungerman
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Fig. 105. Distribution of Leiocolea
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Fig. 109. Distribution of Lejeunea
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Fig. 113. Distribution of Lophocole
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Fig. 117. Distribution of Lophozia
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Fig. 121. Distribution of Lophozia
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Fig. 125. Distribution of Lunularia
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Fig. 129. Distribution of Marchanti
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Fig. 133. Distribution of Marsupell
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Fig. 137. Distribution of Marsupell
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Fig. 141. Distribution of Moerckia
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Fig. 145. Distribution of Nardia ge
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Fig. 149. Distribution of Odontosch
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Fig. 153. Distribution of Pedinophy
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Fig. 157. Distribution of Peltolepi
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Fig. 161. Distribution of Porella a
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Fig. 165. Distribution of Preissia
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Fig. 169. Distribution of Radula li
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Fig. 173. Distribution of Riccardia
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Fig. 177. Distribution of Riccia ca
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Fig. 181. Distribution of Riccia cr
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Fig. 185. Distribution of Riccia go
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Fig. 189. Distribution of Ricciocar
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Fig. 193. Distribution of Scapania
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Fig. 197. Distribution of Scapania
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Fig. 201. Distribution of Scapania
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Fig. 205. Distribution of Scapania
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Fig. 209. Distribution of Scapania
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Fig. 213. Distribution of Scapania
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Fig. 217. Distribution of Tritomari
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Evaluating the distribution dataFol
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Fig. 224. Hornworts and Liverworts
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Signs used:Bolded names - species i
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Marchantiaceae50. VU - Bucegia roma
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Fig. 228. The distribution of zones
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Anthoceros communis Steph. = Phaeoc
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Blasia funckii Corda = Blasia pusil
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Cephalozia crassiflora Spruce = Cep
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Cephaloziella spinigera (Lindb.) Wa
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Diplomitrium lyellii (Hook.) Corda
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Gymnocolea inflata subsp. acutiloba
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Jungermannia acuta var. gypsacea Go
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Jungermannia dentata Raddi = Cephal
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Jungermannia limprichtii Lindb. = L
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Jungermannia scalaris var. denudata
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Leiocolea heterocolpos (Thed. ex C.
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Lophozia incisa var. inermis Müll.
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Marchantia polymorpha var. dilatata
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Mastigobryum deflexum (Mart.) Ångs
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Odontoschisma sphagni var. denudatu
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Porella platyphylla (L.) Pfeiff.Por
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Riccia maior Roth = Riccia glauca L
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Scapania heterophylla M. Howe = Sca
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Sphagnocetis communis var. macrior
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Mittheilungen des siebenbürgischen
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999. Papp C., Eftimie Elena, 1963.
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1059. Péterfi L.Ş., 1963. Flagela
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1119. Plămadă, Viţalariu, 1971.
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1178. Pop I., Hodişan I., 1958. Co
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1231. Prunescu-Arion Elena, Toniuc
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1342. Rüegsegger F., 1986. Frullan
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1396. Schiffner V., 1913b. Über ei
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1567. Ştefureac T., 1960b. Cercet
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1624. Ştefureac T., 1976c. Rolul b
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1675. Ştefureac T., Cristurean I.,
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1727. Tarnavschi I., Nedelcu G., 19
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1785. Wallfisch R., 2008. Contribut
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var. fragilis (A. Roth) Müll. Frib
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Riccardia incurvata Lindb. 28, 248,