The Hornwort and Liverwort Atlas of Romania

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Galbena Valley, Arieşul Mare upstream to Arieşeni, Iarba Rea Valley, Gârda Seacă – upstream Casa dePiatră, Gogii Brook – Cobleş, Cobleş Valley, Scărişoara Cave – aven, Mătişeşti, Horea, Divaia Valley, onrotten wood (Goia 2001:76); Arieşul Mare Basin, on rotten wood (Goia & Schumacker 2002:87); ArieşulMic Valley down from the mine, 20.08.1994, ass. Orthodicrano montani-Hypnetum filiformis, ArieşulMic Valley up from the mine, 20.08.1994, ass. Dicrano scoparii-Hypnetum filiformis (Goia &Schumacker 2003:64, Tab.5); between Mătişeşti and Şaua Ursoaia, 730 m alt., ass. Orthodicranomontani-Hypnetum filiformis (Goia & Schumacker 2004:111, Tab.4); Detunata Mountains, 1055-1100 malt., det. Sass-Gyarmati A. (Sass-Gyarmati, Pócs & Orbán 2005:127).Cluj County, Dobrin Mountain (Péterfi M. 1918:61); Sălicea, leg. & det. Péterfi M. (Boros1958:4,8); the peatbogs from spring and upper course of Someşul Rece, incl. Muntele Mare and Dobrin(Pop E. 1960:397).Bihor Mountains, without locality, 300-900 m alt. (Péterfi M. 1908:276; Goia 2001:76); GalbenaValley, 750 m alt., 08.1999 (Jakab 2000b:66); Someşul Cald Gorges, 4.08.1998 (Coldea & Wagner1999:85, Tab.1); Someşul Cald Gorges (Goia & Mătase 2001:17); Cetăţile Ponorului, AB, 46°33'55"N /22°42'15"E, 1050 m alt., 3.09.2007, leg. & det. Ştefănuţ S. [BUCA B3838, B3852], ♀ - [BUCA B3862].Lophocolea bidentata (L.) Dumort. var. bidentataSyn.:– Chiloscyphus coadunatus (Sw.) J.J. Engel & R.M. Schust.– Chiloscyphus cuspidatus (Nees) J.J. Engel & R.M. Schust.– Chiloscyphus latifolius (Nees) J.J. Engel & R.M. Schust.– Jungermannia bidentata L.– Jungermannia coadunata Sw.– Lophocolea alata (Gottsche et al.) Mitt. ex Larter– Lophocolea bidentata var. ciliata Velen.– Lophocolea bidentata var. cuspidata (Nees) Gottsche et al.– Lophocolea coadunata (Sw.) Montin– Lophocolea cuspidata (Nees) Limpr.– Lophocolea cuspidata (Nees) Limpr. var. banatica Ştefur.First citation for Romania: Baumgarten 1846.Moldova, incl. sub L. cuspidata (Papp 1943:660; Ştefureac & al. 1973:144).Muntenia, incl. sub L. cuspidata (Mohan 1988b:108).Transylvania (Pax 1908:129).Ecology: epigaeic, saprolignicolous, acidophytic, meso-hygrophytic, sciophytic; circumpolar; mountain; Hch.Herbaria and Exsiccate: BUCA, PHHM, SIB, BRHEDistribution in Europe: Is, Fo, No, Dk, Se, Fi, Ie, Br, Fr, Nl, Be, De, Po, Ch, At, Cz, Sk, Hu, Pt, Es, Bl, Co, Sa, Sc, It, Si, Hr,Se, Me, Ba, Ro, Bg, Al, Gr, Ee, Lv, Lt, By, Ua, Cm, Rn, Rw, Rc, Re, Az, Ma, Cn.Distribution in Romania:Maramureş Mountains, Cornedei Peak - Prislop, 1550-1590 m alt., 19-20.06.1975, ass.Rhododendro kotschyi-Pinetum mugi (Borza 1959) em. Coldea 1985 subass. sphagnetosum nemoreiPlămadă & Coldea 1985 (Plămadă & Coldea 1985a:120,121, Tab.1); between Tocila Valley and Băiuţ,common spruce forest, 600 m alt., 21-23.08.1998 (Jakab 1999:55).Rodna Mountains, Lala Valley, 1200 m alt., 17.08.1937 (Ştefureac 1938:241,250,258).Ţibleş Mountains, Izvorul Ţibleşului, Ţibleş Valley, Mesteacănul Valley, 1984-1985, sub L.cuspidata (Mohan 1988a:20).Suceava County, Argestrul, 16.05.1910 (Lilienfeldówna 1911:734,735); Seriune Hill, northernslope, Ştirbu Brook Valley, sub L. cuspidata (Raclaru & Barbu 1959: Tab.6). Bistriţa Aurie Valley,between Iacobeni and Ciocăneşti (Pascal 1969:107; Ştefureac & Pascal 1981:479); Ouşorul Mountain, onsoil, 1971 (Mihai 1974b:318); Cristişor-Neagra Broştenilor, Borcut Meadow, in common spruce forest,aldertree thicket and peatbog, incl. sub fo. interrupta, ass. Pogonatetum alloides v. Krusenstjerna 1948(Lungu 1971; Lungu 1973b:72; Lungu 1974:74, Tabs.1,2; Lungu 1976:238, Tab.2; Lungu 1981c:92;Ştefureac & Pascal 1981:479); Barnar Gorges, on soil and humus (Pascal & Seghedin 1977:141).Bârgău Mountains, Cucureasa Reserve, Coşna Forest Hut, 930-1230 m alt., 1967, 1969, ass.Tetraphidetum pellucidae (Barkman 1958) Maurer 1961 (Ştefureac, Mihai & Pascal 1976:36, Tab.1);Străja Valley, Măgura Calului, Piatra Fântânele, Tomnatec Peak, Măgura Ilvei, Iliuţa Bozghi Brook,Tureac Brook, Secu Brook, Cârţibavu Mare Valley, 1983-1984, incl. sub L. cuspidata (Mohan &Hodoroga 1986:212).Rarău Mountain, Rarău, 14.05.1910 (Lilienfeldówna 1911:735); Codrul Secular Slătioara, 860-1400 m alt., incl. sub L. cuspidata (Ştefureac 1941:1154,1279, Tabs.1,3,7; Papp 1969b:188); between147
Dealul Colbului and Pietrele Doamnei, 1520 m alt., 27-28.08.1949, 27.08.1950 (Ştefureac 1951a:60,65);Plaiul Todirescu-Slătioara Reserve, incl. sub L. cuspidata (Ştefureac & Raclaru 1978:196).Giumalău Mountain, Giumalău Secular Forest, 47°26'12"N / 25°29'20"E, 1210 m alt., 1.09.2008,leg. & det. Ştefănuţ S. [BUCA B3935].Stânişoara Mountains, Stânişoarei Culm and Ostra Brook Valley below Puzdrele, on wet soil,1974 (Pascal & Toma M. 1977:42).Iaşi County, Repedea-Bârnova Forest Massif, on rotten woods (Papp 1924a:101; Papp1929:555,558; Papp & Eftimie 1963:108); Repedea and Bârnova forests (Papp 1939a:353).Bistriţa-Năsăud County, Măgheruş Valley, Hederiş Hill, around the springs, Querco-Fagetea, subL. cuspidata (Ştefureac & Cristurean 1973:315).Vaslui County, Stroieşti Forest, on soil, 1966-1969 (Plămadă & Viţalariu 1971:166).Ceahlău Mountain, Ceahlău, incl. sub L. cuspidata (Papp 1927b:149; Papp 1934b:66,68,72; Papp1942b:525; Zanoschi 1970:280; Manoliu & al. 1998:336).Neamţ County, Poieni Forest (Papp 1924a:101; Papp 1947a:159); Peste Recea, Muncelaşu andPoarta Poienilor Hills, in forests (Papp 1947b:182,188,190); Măgura Pietricica, on wet soil (Papp1937d:235,238,240); Neamţ Monastery, on clayey soil and humus, 07-08.1921,1923, leg. Brândză M.,det. Ştefureac T., incl. sub L. cuspidata (Ştefureac 1942a:4).Giurgeu Mountains, Bicaz Gorges, 1957-1960 (Papp & al. 1962:144); Bicaz Gorges, 1978, 1979,ass. Tortello-Ctenidietum (Gams 1927) Stodiek 1937 (Mihai 1981a:18, Tab.5).Harghita County, Vlăhiţa (Olohfalu), 20.08.1901 (Barth 1905:13).Harghita Mountains, Rotund Mountain, 1100 m alt., 28.08.2001, leg. Bálint L. & Csízi K., subChiloscyphus coadunatus (Bálint & Orbán 2003:14; Bálint & Orbán 2006:468).Nemira Mountains, Nemira, on siliceous rocks (Barabaş V. 1974:31); Nemira, near upside lakes,on clayey soil, 1970-1977 (Mihai & Barabaş V. 1978:115); Nemira Peak, Şandru Mountain, 1400-1500 malt. (Barabaş V. 1982b:56, Tab.3).Bacău County, Colina Măgura, 514 m alt. (Papp 1957:285; Papp 1958b:388); s - Măgura Hill,north-west to Târgu Ocna, the edge of brooks and cuted forest, on soil, 680 m alt., frequent (Mihai &Barabaş V. 1971:676,682); the forest way from Înţărcătoare towards Zboina Verde, Fundu Caşin, GuraDobrilor, on rotten fir wood and rocks (Eftimie 1972:151); Slănic Brook, on fir tree stump, 2.05-8.11.1971 (Ştefureac & Barabaş V. 1972:77,82).Ciucaş Mountains, Zăgan Mountain, Chiruşca Mountain, Fugarul Brook Valley, Berii BrookValley, Stâncoasă Culm, sub L. cuspidata, Berii Brook Valley, Roşu Brook Valley, Piscul Caprei,Căzăturii Culm, Piscul Gropşoarele, Piatra Laptelui Culm (Mohan 1990b:43).Braşov County, Cetăţii (Lempeş) Hill, in glades, on soil, 05.1961, sub L. cuspidata (Mihai1962:278).Piatra Mare Mountain, Tamina Waterfall, 45°32'58"N / 25°38'33"E, 6.09.1997, leg. & det.Ştefănuţ S. [BUCA B2453].Gârbova Massif, Azuga Valley, Tufa Valley, on humus, 1967-1970 (Lungu & Ştefureac 1970:5).Prahova County, Doftana Valley [PHHM 7743/3873](Mohan 1971a:8; Mohan 1971b:47);Doftana Valley between Trăistieni and Tesila, 850 m alt. (Mohan 1981:99; Mohan 1990a:24); DoftanaValley, in mixed Quercus petraea with Fagus sylvatica forest, 45°10'01,6"N / 25°45'09,2"E, 27.04.2006,leg. & det. Ştefănuţ S. [BUCA B3427] (Vasiliu-Oromulu 2008:87).Piatra Craiului Mountains, the way towards Vlăduşca Pasture, BV, 1100 m alt., 30.10.2001, leg.Pop O., det. Ştefănuţ S. [BUCA B2834] (Dihoru G. & al. 2003:69; Ştefănuţ & Pop O.G. 2006:77);Crăpătura Pietrei Craiului, BV, 45°33'30"N / 25°15'44"E, 1300 m alt., 22.08.2003, leg. & det. Ştefănuţ S.[BUCA B2696, B2714] (Ştefănuţ & Pop O.G. 2006:77); towards Padinile Frumoase, BV, 1700 m alt.,7.08.2004, leg. Pop O., det. Ştefănuţ S. [BUCA B3002] (Ştefănuţ & Pop O.G. 2006:77); Marele Grohotiş,AG, 45°30'36"N / 25°11'45"E, 1650 m alt., 24.07.2002, leg. Pop O., det. Ştefănuţ S. [BUCA B3068],45°30'36"N / 25°11'38"E, 1550 m alt., 8.07.2003, leg. & det. Ştefănuţ S. [BUCA B2646] (Ştefănuţ & PopO.G. 2006:77).Dâmboviţa County, Corbii Mari, 6.05.1998, leg. & det. Ştefănuţ S. [BUCA B3377].Făgăraş Depression, ♀ - Lacul Tătarilor Peatbog, SB, 45°42'56"N / 24°29'07"E, 540 m alt.,8.10.2003, leg. & det. Ştefănuţ S. [BUCA B3085] (Ştefănuţ 2004f:74); Lacul Tătarilor Peatbog, SB,45°42'56"N / 24°29'07"E, 540 m alt., 20.11.2006, leg. & det. Ştefănuţ S. [BUCA B3591].148
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SORIN ŞTEFĂNUŢThe Hornwort and L
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CONTENTINTRODUCTION ...............
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The beginning of the 20 th century
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asin of the Arieş river. Bryotaxon
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Abbreviations used in the text† -
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The presentation of the methodology
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Fig. 3. The GEOCOD mapping system.
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THE HORNWORT AND LIVERWORT SPECIES
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THE HORNWORT AND LIVERWORT CLASSIFI
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Suborder Lepidoziineae R.M. Schust.
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46. Cephalozia pleniceps (Austin) L
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147. Pallavicinia lyellii (Hook.) C
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13 Corsinia coriandrina (Spreng.) L
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11. Porella obtusata (Taylor) Trevi
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34Phaeoceros carolinianus (Michx.)
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Făgăraş Mountains, Cârţişoara
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Cluj County, Someşul Rece Valley a
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Parâng Mountains, Repedea Valley,
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Argeş County, Rucăr (Loitlesberge
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Cindrel Mountains, Cindrel, 2200 m
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Parâng Mountains, Latoriţa Valley
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Athalamia hyalina (Sommerf.) S. Hat
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Barbilophozia barbata (Schmidel ex
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Bârgău Mountains, Cârţibavu Mar
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First citation for Romania: Schur 1
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- Pleuroschisma triangulare (Schlei
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and Băiuţ, common spruce forest,
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Distribution in Europe: Sj, Is, Fo,
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Poiana Stampei Peatbog, SV, 47°20'
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alt., 8.08.2003, leg. Pop O., det.
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leg. & det. Ştefureac T., ass. Buc
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Rodna Mountains, Puzdra Mountain, 1
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Alba County, Vârciorog Valley, 780
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Bârgău Mountains, Cucureasa Reser
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Suceava County, Tinovul Mare de la
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Suceava County, Rodna Nouă, on Per
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2001:74,75); Arieşul Mare Basin, o
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Ţibleş Mountains, Mesteacănul Va
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Cindrel Mountains, Găuşoara, leg.
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Parâng Mountains, the edge of Câl
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& det. Ştefănuţ S. [BUCA B3684];
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Alba County, at 20,5 km to Zlatna,
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Cephaloziella spinigera (Lindb.) Wa
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First citation for Romania: Péterf
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Iezer-Păpuşa Mountains, Iezer Val
Page 97 and 98: - Cladopodiella fluitans var. gigan
Page 99 and 100: Caraş-Severin County, Cerna Valley
Page 101 and 102: (Mititelu, Barabaş N. & al. 1968:1
Page 103 and 104: 1885:30; Ştefureac 1949a:287,288;
Page 105 and 106: 104Conocephalum salebrosum Szweyk.,
Page 107 and 108: Retezat Mountains, Retezat, sub Jun
Page 109 and 110: Cindrel Mountains, Cânaia, 1750 m
Page 111 and 112: Iaşi County, Sineşti Forest (Papp
Page 113 and 114: [BUCA B2793], on rotten beech trunk
Page 115 and 116: Orthodicrano montani-Hypnetum filif
Page 117 and 118: Distribution in Romania:Maramureş
Page 119 and 120: - Gymnocolea inflata var. laxa (Nee
Page 121 and 122: Ciucaş Mountains, Caprei Brook Val
Page 123 and 124: Giumalău Mountain, Giumalău Secul
Page 125 and 126: - Haplozia riparia var. rivularis B
Page 127 and 128: - Jungermannia karl-muelleri auct.,
Page 129 and 130: Jungermannia gracillima Sm.Syn.:- A
Page 131 and 132: Mountain, PH, 45°24'40"N / 25°28'
Page 133 and 134: Retezat Mountains, Rovine Brook, on
Page 135 and 136: Romania, alt. 600-2280 m (Váňa 19
Page 137 and 138: - Solenostoma subellipticum (Lindb.
Page 139 and 140: Distribution in Romania:Bârgău Mo
Page 141 and 142: 13.07.1946, sub Lophozia alpestris
Page 143 and 144: Distribution in Romania:Maramureş
Page 145 and 146: Rodna Mountains, Lala Valley, 1200
Page 147: Ştefănuţ S. [BUCA B2574, B2580,
Page 151 and 152: 150var. rivularis (Raddi) Warnst.Sy
Page 153 and 154: Penteleu Massif, fer - Roşu Lake,
Page 155 and 156: Satu Mare County, Noroieni, Ghiarma
Page 157 and 158: Lophozia ascendens (Warnst.) R.M. S
Page 159 and 160: - Jungermannia intermedia Lindenb.-
Page 161 and 162: Căliman Mountains, on humus and ro
Page 163 and 164: Hăşmaş Mountains, Hăşmaş, sub
Page 165 and 166: Herbaria and Exsiccate: BUCADistrib
Page 167 and 168: peatbog on the border of Câlcescu
Page 169 and 170: ? - Iezer-Păpuşa Mountains, Iezer
Page 171 and 172: Făgăraş Mountains, s - above Avr
Page 173 and 174: Nemira Mountains, on edge of Nemira
Page 175 and 176: 2003:7); Măgura Cisnădiei [SIB HL
Page 177 and 178: Fuss 1854:20; Ştefureac 1949a:288;
Page 179 and 180: - Nardia brevissima (Dumort.) Lindb
Page 181 and 182: 1919:295); Ciortea Mountain, on wet
Page 183 and 184: - Sarcoscyphus sphacelatus var. med
Page 185 and 186: Siriu Mountain, Poiana din Fundul M
Page 187 and 188: Iaşi County, below Repedea, 7.10.1
Page 189 and 190: Valley, Arieşul Mic - upstream min
Page 191 and 192: Alba County, Valea Lungă, 10.02.18
Page 193 and 194: Rarău Massif, between Dealul Colbu
Page 195 and 196: Harghita County, Harghita Spa, on r
Page 197 and 198: Căliman Mountains, in alpine zone,
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Stânişoara Mountains, Suha Mare B
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Ecology: humicolous, acidophytic, h
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Bacău County, Slănic Brook (Papp
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Rarău Mountain, Codrul Secular Sl
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1915:197,198); Cândeşti, 07.1891,
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Ţibleş Mountains, Piciorul Ţigle
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Ceahlău Mountain, Durău Valley, i
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major (Ştefureac 1962b:69); Benea-
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Ciucaş Mountains, Berii Brook Vall
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Zlătuia Valley, 1100 m alt., 7.09.
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- Plagiochila dillenii (Taylor) Tay
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Wallfisch R. (Dihoru G. & al. 2003:
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Ştefănuţ M. 2003:79); Iezerul Pi
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utabuli-Mnietum cuspidati (Felföld
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Iaşi County, Mârzeşti Forest, Bu
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3.04.2004, leg. Atanasiu Paulina, d
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Arad County, Săvârşin, sub Madot
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Preissia quadrata (Scop.) NeesSyn.:
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Diana, BV, 1100 m alt., 22.08.2001,
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Rodna Mountains (Pax 1908:130); Pie
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Descut, 15.08.2001, Podul de Piatr
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Bârgău Mountains, Cucureasa Reser
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B3521, B3522] (Ştefănuţ 2006b:34
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16,20.08.1994, ass. Pterigynandretu
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- Marchantia crinita Michx.- Marcha
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Giumalău Mountain, Giumalău Secul
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Riccardia multifida (L.) GraySyn.:-
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Penteleu Massif, the left side of C
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Herbaria and Exsiccate: SIBDistribu
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558,560; Papp 1939a:358; Papp & Eft
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Riccia glauca L. var. glaucaSyn.:-
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Distribution in Europe: Is, Fo, No,
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Bihor County, Tărian, sub Riccia (
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1960:1113); upper side of Mălăie
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Scapania brevicaulis TaylorSyn.:- S
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Suceava County, Cristişor-Neagra B
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Piatra Mare Mountain, 1700 m alt.,
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- Scapania amurensis Warnst.- Scapa
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Mureş County, Sighişoara - Fuchsl
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Căliman Mountains, the peatbogs fr
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- Scapania heterophylla M. Howe- Sc
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Slăvei and Muchia Ascuţită, 2000
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Distribution in Europe: Ie, Br, Fr,
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Nemira Mountains, Nemira, on humus,
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- Sphenolobus medelpadicus (Arnell)
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Parâng Mountains, Latoriţa Valley
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Fig. 9. Distribution of Phaeoceros
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Fig. 13. Distribution of Anastrophy
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Fig. 17. Distribution of Anthelia j
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Fig. 21. Distribution of Asterella
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Fig. 25. Distribution of Barbilopho
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Fig. 29. Distribution of Barbilopho
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Fig. 33. Distribution of Bazzania t
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Fig. 37. Distribution of Calypogeia
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Fig. 41. Distribution of Calypogeia
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Fig. 45. Distribution of Cephalozia
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Fig. 49. Distribution of Cephalozia
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Fig. 53. Distribution of Cephalozie
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Page 317 and 318:
Page 319 and 320:
Fig. 65. Distribution of Cololejeun
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Fig. 69. Distribution of Corsinia c
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Fig. 73. Distribution of Fossombron
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Fig. 77. Distribution of Frullania
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Fig. 81. Distribution of Gymnocolea
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Fig. 85. Distribution of Harpanthus
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Fig. 89. Distribution of Jungermann
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Page 335 and 336:
Page 337 and 338:
Fig. 101. Distribution of Jungerman
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Fig. 105. Distribution of Leiocolea
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Fig. 109. Distribution of Lejeunea
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Fig. 113. Distribution of Lophocole
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Fig. 117. Distribution of Lophozia
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Fig. 121. Distribution of Lophozia
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Fig. 125. Distribution of Lunularia
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Fig. 129. Distribution of Marchanti
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Fig. 133. Distribution of Marsupell
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Fig. 137. Distribution of Marsupell
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Fig. 141. Distribution of Moerckia
Page 359 and 360:
Fig. 145. Distribution of Nardia ge
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Fig. 149. Distribution of Odontosch
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Fig. 153. Distribution of Pedinophy
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Fig. 157. Distribution of Peltolepi
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Fig. 161. Distribution of Porella a
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Fig. 165. Distribution of Preissia
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Fig. 169. Distribution of Radula li
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Fig. 173. Distribution of Riccardia
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Fig. 177. Distribution of Riccia ca
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Fig. 181. Distribution of Riccia cr
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Fig. 185. Distribution of Riccia go
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Fig. 189. Distribution of Ricciocar
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Fig. 193. Distribution of Scapania
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Page 387 and 388:
Page 389 and 390:
Page 391 and 392:
Page 393 and 394:
Page 395 and 396:
Fig. 217. Distribution of Tritomari
Page 397 and 398:
Evaluating the distribution dataFol
Page 399 and 400:
Fig. 224. Hornworts and Liverworts
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Signs used:Bolded names - species i
Page 403 and 404:
Marchantiaceae50. VU - Bucegia roma
Page 405 and 406:
Fig. 228. The distribution of zones
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Anthoceros communis Steph. = Phaeoc
Page 409 and 410:
Blasia funckii Corda = Blasia pusil
Page 411 and 412:
Cephalozia crassiflora Spruce = Cep
Page 413 and 414:
Cephaloziella spinigera (Lindb.) Wa
Page 415 and 416:
Diplomitrium lyellii (Hook.) Corda
Page 417 and 418:
Gymnocolea inflata subsp. acutiloba
Page 419 and 420:
Jungermannia acuta var. gypsacea Go
Page 421 and 422:
Jungermannia dentata Raddi = Cephal
Page 423 and 424:
Jungermannia limprichtii Lindb. = L
Page 425 and 426:
Jungermannia scalaris var. denudata
Page 427 and 428:
Leiocolea heterocolpos (Thed. ex C.
Page 429 and 430:
Lophozia incisa var. inermis Müll.
Page 431 and 432:
Marchantia polymorpha var. dilatata
Page 433 and 434:
Mastigobryum deflexum (Mart.) Ångs
Page 435 and 436:
Odontoschisma sphagni var. denudatu
Page 437 and 438:
Porella platyphylla (L.) Pfeiff.Por
Page 439 and 440:
Riccia maior Roth = Riccia glauca L
Page 441 and 442:
Scapania heterophylla M. Howe = Sca
Page 443 and 444:
Sphagnocetis communis var. macrior
Page 445 and 446:
Mittheilungen des siebenbürgischen
Page 447 and 448:
91. Borza A., 1922a. Schedae ad "Fl
Page 449 and 450:
142. Boşcaiu N., Täuber F., Colde
Page 451 and 452:
199. Coldea G., 1975. Étude phytos
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The Hornwort and Liverwort Atlas of Romania

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