ÎÃËßÄ ÒÀ ÂÈÇÍÀ×ÍÈÊ Ê˲ٲ ÐÎÄÓ CALVOLIA (SAPROGLYPHIDAE)
ÎÃËßÄ ÒÀ ÂÈÇÍÀ×ÍÈÊ Ê˲ٲ ÐÎÄÓ CALVOLIA (SAPROGLYPHIDAE)
ÎÃËßÄ ÒÀ ÂÈÇÍÀ×ÍÈÊ Ê˲ٲ ÐÎÄÓ CALVOLIA (SAPROGLYPHIDAE)
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³ñíèê ÎÍÓ Òîì 10, âèïóñê 3, 2005. Á³îëîã³ÿ<br />
114<br />
ÓÄÊ 595.42:591<br />
Â. Ä. Ñåâàñòüÿíîâ, ä-ð á³îë. íàóê, ïðîô.,<br />
Ä. À. ʳâãàíîâ, êàíä. á³îë. íàóê, äîö.,<br />
Äæàðìà Íàä³ÿ Àë³, ìàã³ñòð çîîë.<br />
Îäåñüêèé íàö³îíàëüíèé óí³âåðñèòåò ³ì. ². ². Ìå÷íèêîâà,<br />
êàôåäðà çîîëî㳿,<br />
âóë. Äâîðÿíñüêà, 2, Îäåñà, 65026, Óêðà¿íà<br />
<strong>ÎÃËßÄ</strong> <strong>ÒÀ</strong> <strong>ÂÈÇÍÀ×ÍÈÊ</strong> <strong>Ê˲ٲÂ</strong> <strong>ÐÎÄÓ</strong> <strong>CALVOLIA</strong><br />
(<strong>SAPROGLYPHIDAE</strong>)<br />
гä Calvolia (Saproglyphidae) ñêëàäàºòüñÿ ç 38 âèä³â êë³ù³â, ³ç ÿêèõ ó øåñòè<br />
âèä³â îïèñàí³ ñàìö³, ñàìèö³ òà ã³ïîïóñè; ó äâîõ âèä³â — ñàìö³ òà ñàìèö³, ó<br />
ï'ÿòè — ò³ëüêè ñàìèö³ ³ ó 25 âèä³â — ò³ëüêè ã³ïîïóñè. Äëÿ êîæíîãî âèäó<br />
íàâåäåíî éîãî ãåîãðàô³÷íå òà åêîëîã³÷íå ì³ñöå çíàõîäæåííÿ. Calvolia<br />
hebeclinii Sicher 1899 âèëó÷àºòüñÿ ç³ ñïèñêó âèä³â ðîäó ÿê nomen dubium, a<br />
Calvolia zaheri Ouds., 1930 — ÿê íåâ³ðíî âèçíà÷åíèé àâòîðîì âèä.<br />
Êëþ÷îâ³ ñëîâà: êë³ù³, ð³ä, âèä, ã³ïîïóñè, âèçíà÷íèê.<br />
гä Calvolia ïîñòóëüîâàíî [1] ó ñêëàä³ ðîäèíè Àñàridae íà îñíîâ³<br />
îñîáëèâîñòåé áóäîâè ã³ïîïóñ³â Calvolia hagensis Ouds, 1911.  ñó÷àñí³é<br />
àêàðîëî㳿 [2, 3] ð³ä Calvolia ðîçãëÿäàºòüñÿ ÿê íàëåæíèé äî ðîäèíè<br />
Saproglyphidae.<br />
Ìåòà ö³º¿ ïóáë³êàö³¿ — ðîçãëÿä ñó÷àñíîãî îá'ºìó ðîäó òà ïðåäñòàâëåííÿ<br />
âèçíà÷íèêà âñ³õ ñòàä³é ðîçâèòêó, îñê³ëüêè ïîïåðåäí³ âèçíà÷íèêè<br />
âæå íåïðèäàòí³ äëÿ êîðèñòóâàííÿ [2–4].<br />
dz ñïèñêó â³äîìèõ ç ë³òåðàòóðè âèä³â ðîäó íàìè áóëè âèêëþ÷åí³<br />
äâà âèäè:<br />
1) C. hebeclinii (Sicher), 1889. Âïåðøå áóâ îïèñàíèé ÿê Glycyphagus<br />
hebeclinii. Ñàìêà áóëà çíàéäåíà â Ñèöè볿 íà òåïëè÷í³é ðîñëèí³<br />
ðîäîì ç Àíòèëüñüêèõ îñòðîâ³â, Ãâ³àíè, Áðàçè볿, Ïåðó òà Íîâî¿<br />
Çåëàí䳿. À. À. Çàõâàòê³í [2] â³äçíà÷àâ äóæå íåäîñêîíàë³ ðèñóíêè<br />
â ïåðøîìó îïèñ³ âèäó. Îñê³ëüêè á³ëüøå ñòà ðîê³â âèä â ë³òåðàòóð³<br />
íå çãàäóâàâñÿ, çà ïðàâèëàìè çîîëîã³÷íî¿ íîìåíêëàòóðè<br />
ìè ââàæàºìî éîãî "ãîëîþ íàçâîþ" — nomen dubium.<br />
2) C.(?) zacheri Oudemans, 1930. Ñóìí³âè ùîäî â³äíåñåííÿ äàíîãî<br />
âèäó äî ðîäó Calvolia âèñëîâëþâàëè ñïî÷àòêó À. À. Çàõâàòê³í<br />
[2], à ïîò³ì Â. Ä. Ñåâàñòüÿíîâ [5]. Íàÿâí³ñòü ó ã³ïîïóñó î÷åé íà<br />
ã³ñòåðîñîì³, à òàêîæ ðîçâèòîê íà öâ³ëîìó ñèð³ õàðàêòåðí³ äëÿ<br />
ïðåäñòàâíèê³â ³íøî¿ ðîäèíè — Anoetidae. Ó çâ'ÿçêó ç âèùåñêàçàíèì,<br />
ìè âèâîäèìî äàíèé âèä ç ðîäó Calvolia. Çà ïðàâèëàìè<br />
çîîëîã³÷íî¿ íîìåíêëàòóðè, ï³ñëÿ ïðîâåäåííÿ ðåâ³ç³¿, íàçâà âèäó<br />
"zacheri" ìîæå áóòè çáåðåæåíà, àëå â ³íøîìó ðîä³ òà ðîäèí³.<br />
© Â. Ä. Ñåâàñòüÿíîâ, Ä. À. ʳâãàíîâ, Äæàðìà Íàä³ÿ Àë³, 2005
Îãëÿä òà âèçíà÷íèê êë³ù³â ðîäó Calvolia<br />
Âèçíà÷íèê ïî ñàìèöÿì<br />
1(2) Øê³ðà ã³ñòåðîñîìè, îñîáëèâî ïîçàäó, í³æíî øòðèõîâàíà òà ìàº<br />
äâà íàï³âêóëÿñò³ ãîðáî÷êè............. C. tuberculata A. Z., 1941.<br />
Ãðóç³ÿ.  êà÷àíàõ êóêóðóäçè [2].<br />
2(1) Ïîêðèâè ã³ñòåðîñîìè âåíòðàëüíî òà äîðñàëüíî ãëàäåíüê³.<br />
3(4) sci º íàéäîâøèìè ùåòèíêàìè; âåðøèíè sci çàõîäÿòü çà îñíîâè<br />
d1; d3 íå äîñÿãàþòü d4, îñòàíí³ íå ìåíø íà ïîëîâèíó âèõîäÿòü<br />
çà êðà¿ ò³ëà..................................... C. kamenskii A. Z., 1941.<br />
Áàøêèð³ÿ. Â ðîñëèííèõ ðåøòêàõ íà ñ³òö³ êîìáàéíó [2].<br />
4(3) Ðîçì³ðè òà ïðîïîðö³¿ âêàçàíèõ ùåòèíîê ³íø³.<br />
5(12) Âåðøèíà àáî é çíà÷íà ÷àñòèíà ÿéöåâèâ³äíîãî îòâîðó âèñòóïàº<br />
çà åï³ìåðè III.<br />
6(7) Çà åï³ìåðè III âèñòóïຠäî òðåòèíè ÿéöåâèâ³äíîãî îòâîðó, åï³ã³í³é<br />
ìàéæå òîðêàºòüñÿ åï³ìåð II ......... C. miniopteri Dusbabek, 1964.<br />
Ñëîâà÷÷èíà. Íà çâè÷àéíîìó äîâãîêðèë³ [6].<br />
7(6) Çà åï³ìåðè III âèñòóïຠëèøå âåðøèíà ÿéöåâèâ³äíîãî îòâîðó.<br />
̳æ âåðøèíîþ åï³ã³í³þ òà åï³ìåðàìè II çíà÷íèé ïðîì³æîê.<br />
8(9) Âåðøèíà ÿéöåâèâ³äíîãî îòâîðó ãîñòðà ... C. heterocoma Mich., 1903.<br />
Çàõ³äíà ªâðîïà, Äàëåêèé Ñõ³ä.  ãí³çäàõ á³ëêè. Ìè ðîçãëÿäàºìî<br />
öåé âèä çã³äíî îïèñó [4].<br />
9(10) Âåðøèíà ÿéöåâèâ³äíîãî îòâîðó ç âóçüêèì òóïèì íà âåðøèí³<br />
â³äðîñòêîì, àáî áóëàâîïîä³áíî ðîçøèðåíà.<br />
10(11) Âåðøèíà ÿéöåâèâ³äíîãî îòâîðó ð³çêî áóëàâîïîä³áíà<br />
.................................................... C. zachvatkini Volgin, 1980<br />
Ðîñ³ÿ.  ðîñëèííèõ çàëèøêàõ òà ãí³çäàõ ãðèçóí³â [7].<br />
11(12) Âåðøèíà ÿéöåâèâ³äíîãî îòâîðó ð³çêî çâóæåíà<br />
........................................................... C. priapus Volgin, 1980<br />
Îäåñà.  ãí³çä³ ñîðîêè. Ñåì³ðàáàä.  ãí³çä³ êâàêà [7].<br />
12(5) Âåðøèíà ÿéöåâèâ³äíîãî îòâîðó íà ð³âí³ àáî íå äîñÿãຠåï³ìåð III.<br />
13(14) Åï³ìåðè III çàãíóò³ òà çðîùåí³ ç ê³íöÿìè åï³ã³í³ÿ<br />
......................................................... C. elongata Volgin, 1980<br />
Êàçàõñòàí.  ï³äñò³ëö³ êîð³âíèêà [7].<br />
14(15) Åï³ìåðè III ãîñòð³, ïðÿì³, íå çðîùåí³ ç åï³ã³í³ºì.<br />
15(18) Âåðøèíà ÿéöåâèâ³äíîãî îòâîðó âèñòóïຠïåðåä åï³ìåðàìè ²V.<br />
16(17) Åï³ìåðè ²² ð³çêî çàãíóò³ â íàïðÿì³ äî êðà¿â ò³ëà. Âíóòð³øí³<br />
ïðîïîäîñîìàëüí³ ùåòèíêè íå ìåíø ÷èì âäâîº äîâø³ d1 òà d4<br />
................................................. C. ruehmi Türk E. et ., 1957<br />
ͳìå÷÷èíà. Åðëàíãåí.  õîäàõ êîðî¿ä³â íà àìåðèêàíñüêîìó<br />
äóá³ [4].<br />
17(16) Åï³ìåðè II ïðÿì³, ãîñòð³. Âíóòð³øí³ ïðîïîäîñîìàëüí³ ùåòèíêè<br />
ëåäâå äîâø³ d1 òà d4 ........ C. tarsoinfracta Türk E. et ., 1957<br />
ͳìå÷÷èíà. Åðëàíãåí.  ãíèëîìó áàäèëë³ êàðòîïë³ [4].<br />
18(15) ßéöåâèâ³äíèé îòâ³ð ö³ëêîì ì³æ åï³ìåðàìè III òà ²V.<br />
19(23) Àíàëüí³ ùåòèíêè a3 çíà÷íî äîâø³ àíàëüíîãî îòâîðó. sci, dl òà<br />
d4 ìàéæå ð³âí³.<br />
115
20(22) Âåðøèíà ÿéöåâèâ³äíîãî îòâîðó âõîäèòü â ñåðåäèíó äâîâåðøèííîãî<br />
åï³ã³í³þ ........................ C. dominicola Shiro Ochima, 1979<br />
ßïîí³ÿ. Â ïèëó ïîìåøêàíü [8].<br />
21(22) Âåðøèíà ÿéöåâèâ³äíîãî îòâîðó íå äîñÿãຠåï³ã³í³þ. Éîãî áîêîâ³<br />
â³äðîñòêè çàõîäÿòü çà åï³ìåðè III ..... C. romanovae A. Z., 1941<br />
Ìîñêâà. Â ñêëàä³ çåðíà ïøåíèö³.<br />
23(19) Àíàëüí³ ùåòèíêè a3 çíà÷íî êîðîòø³ àíàëüíîãî îòâîðó, ÿêùî<br />
äîâø³, òî sci, d2 òà d4 äóæå äîâã³.<br />
24(25) sci íå êîðîòø³ sce; d2 òà d4 äîâø³ ïîëîâèíè ã³ñòåðîñîìè. Âåðøèíè<br />
d3 òà d4 âèõîäÿòü çà êðàé ò³ëà .............. C. brenyi Cooreman, 1963<br />
Áåëüã³ÿ. Êðèì ïîáëèçó ßëòè. Íà êîðî¿äàõ [9].<br />
25(24) sci, d1 òà d4 êîðîòø³ òðåòèíè sce, ò³ëüêè âåðøèíè d4 âèõîäÿòü<br />
çà êðàé ò³ëà ...................................... C. kneissli Krause, 1919<br />
Çàõ³äíà ªâðîïà, íà êîðî¿äàõ òà òðóõëÿâèí³ ñîñíè òà ÿëèíè.<br />
116<br />
Âèçíà÷íèê ïî ñàìöÿì<br />
1(2) Øê³ðà íà ã³ñòåðîñîì³, îñîáëèâî çíèçó òà ïî áîêàì, øòðèõîâàíà, ìàº<br />
ø³ñòü íàï³âêóëÿñòèõ ãîðáî÷ê³â .............. C. tuberculata A. Z., 1941<br />
2(1) Øê³ðà íà ã³ñòåðîñîì³ ãëàäåíüêà.<br />
3(4) Ñòàòåâèé îðãàí (åäåàãóñ) òðóá÷àòèé, áàãàòîðàçîâî çâèâèñòèé, õî÷à á<br />
â 1,6 ðàç³â äîâøèé ò³ëà ........................... C. priapus Volgin, 1980<br />
4(6) Åäåàãóñ ³íøî¿ áóäîâè, êîðîòêèé.<br />
5(11) Åäåàãóñ àáî òîðêàºòüñÿ åï³ìåð IV, àáî çðîùåíèé ç íèìè.<br />
6(7) Àíàëüí³ ùåòèíêè êîðîòø³ àíàëüíî¿ ù³ëèíè<br />
..................................................C. heterocoma Mich., 1903<br />
7(10) Äîâæèíà àíàëüíèõ ùåòèíîê ð³çíà. Åäåàãóñ ç ïåðåõâàòîì ïîñåðåäèí³.<br />
8(9) a2 íàéäîâøà àíàëüíà ùåòèíêà, a3 ïàëè÷êîïîä³áíà, òîâñò³øà a1<br />
......................................... C. dominicola Shiro Ochima, 1979<br />
9(8) a3 íå ïàëè÷êîïîä³áíà, äîâøà a1 òà a2 . C. zachvatkini Volgin, 1980<br />
10(7) Åäåàãóñ ç ïåðåõâàòîì â íàéá³ëüø âóçüê³é ÷àñòèí³<br />
......................................... C. tarsoinfracta Türk E. et ., 1957<br />
11(5) Åäåàãóñ äàëåêî ïîçàäó åï³ìåð IV.<br />
12(13) Êîæíèé ç â³äð³çê³â åï³ìåð III ç ãîñòðèì çóáöåì. d2 íàéêîðîòø³<br />
ùåòèíêè ò³ëà ................... C. miniopteri Dusbabek, 1964<br />
13(14) Áåç çóáö³â íà åï³ìåðàõ III; d2 â ê³ëüêà ðàç äîâøà d1.<br />
14(15) Åï³ìåðè IV ïîñåðåäèí³ ïðÿìîêóòí³. d2 äàëåêî íå äîñÿãàþòü<br />
ïîëîâèíè d1 .................................... C. kneissli Kramer, 1899<br />
15(14) Åï³ìåðè IV, ãîðèçîíòàëüí³. sci ð³âí³ sce .. C. brenyi Cooreman, 1963<br />
Âèçíà÷íèê ïî ã³ïîïóñàì<br />
Â. Ä. Ñåâàñòüÿíîâ, Ä. À. ʳâãàíîâ, Äæàðìà Íàä³ÿ Àë³<br />
1(2) Âåðøèíà ëàïêè IV ç îäí³ºþ ïàðóñíîþ ùåòèíêîþ<br />
........................................................ C. kneissli Kramer, 1899<br />
2(1) Âåðøèíà ëàïêè IV ç äâîìà êîðîòêèìè ùåòèíêàìè.<br />
3(18) Âåíòðóì â³äñóòí³é.
Îãëÿä òà âèçíà÷íèê êë³ù³â ðîäó Calvolia<br />
4(13) Ç ÷³òêèìè ïîòîâùåííÿìè ïîêðèâ³â á³ëÿ ïðèñìîêòóâàëüíîãî äèñêó.<br />
5(12) Ïåðåäí³ ïîòîâùåííÿ ïîêðèâ³â äîâø³ àíàëüíî¿ ù³ëèíè.<br />
6(9) Ïîòîâùåííÿ ïîêðèâ³â âåðòèêàëüí³, ìàéæå ïàðàëåëüí³ àíàëüí³é<br />
ù³ëèí³.<br />
7(8) Ïîòîâùåííÿ ïîêðèâ³â øèðîê³. Äèñòàëüíèé ÷ëåíèê ãíàòîñîìè<br />
ñïåðåäó áåç âèð³ç³â. Ñòåðíàëüíèé ùèò íå ãðàíè÷èòü ç åï³ìåðàìè<br />
III. Øòðèõîâêà ïîêðèâ³â ìàéæå áåç àíàñòîìîç³â<br />
........................................................ C. brenyi Cooreman, 1963<br />
8(7) Ïîòîâùåííÿ ïîêðèâ³â òîíê³, íå ñóâîðî ïàðàëåëüí³. Êîìá³íàö³ÿ<br />
âêàçàíèõ îçíàê ³íøà................ C. fraxini Türk E. et ., 1957<br />
ͳìå÷÷èíà. Ìþíõåí.  òðóõëÿâîìó ÿñåí³.<br />
9(6) Ïîòîâùåííÿ íàï³âÿéöåïîä³áí³, ïîçàäó ³ç çíà÷íèìè çóáöÿìè.<br />
10(11) Ïîòîâùåííÿ ð³çêî äóãîïîä³áí³ ç âåðøèíàìè, íàïðàâëåíèìè íàçóñòð³÷<br />
îäíà îäí³é ..................... C. minima Türk E. et ., 1957<br />
ͳìå÷÷èíà. Íà êîðî¿äàõ, â òðóõëÿâîìó äóá³.<br />
11(12) Ïîòîâùåííÿ ñëàáî âèãíóò³, ïîçàäó ç âåëèêèì çóáöåì, ïåðåäí³é<br />
êðàé äèñêó ç äâîìà çóá÷èêàìè ............ C. natalia A. Z., 1941<br />
Çàõ³äíà ªâðîïà. Ìîñêâà. Íà êîðî¿äàõ.<br />
12(5) Ïîòîâùåííÿ êîðîòø³ àíàëüíî¿ ù³ëèíè, ïîïåðå÷í³, ïîñåðåäèí³<br />
ð³çêî ðîçøèðåí³ .................... C. wichmanni Türk E. et ., 1957<br />
ͳìå÷÷èíà. Íà äóáîâîìó âóñà÷³ òà ³íøèõ Cerambicidae.<br />
13(4) Ïîòîâùåííÿ ïîêðèâ³â ïîðÿä ç ïðèñìîêòóâàëüíèì äèñêîì<br />
â³äñóòí³.<br />
14(15) Çðîùåí³ ëàïêà òà ãîì³ëêà IV óòâîðþþòü ò³á³îòàðçóñ. Îäíà ç éîãî<br />
ïàðóñíèõ ùåòèíîê íå äîâøà 3/4 ³ä³îñîìè .. C. striata Vitzthum, 1914<br />
²òàë³ÿ. Íà êîðî¿äàõ.<br />
15(14) Ò³á³îòàðçóñ â³äñóòí³é. Îäíà ç ùåòèíîê ëàïêè äîâøà ò³ëà.<br />
16(17) ²ä³îñîìà åë³ïòè÷íà. Ñïèíí³ ùèòè ç øèðîêèì ñ³ò÷àñòèì ìàëþíêîì<br />
............................................... C. elli ptica A. Z., 1941<br />
Ìîñêâà. Íà êîðî¿äàõ.<br />
17(16) ²ä³îñîìà âåðåòåíîïîä³áíà. Ñïèíí³ ùèòè â äóæå ãóñòîìó øòðèõóâàíí³<br />
............................................. C. fusiformis A. Z., 1941<br />
Ìîñêâà. Íà êîðî¿äàõ.<br />
18(3) Âåíòðóì ÷³òêî âèðàæåíèé.<br />
19(24) 3 ïîòîâùåííÿìè ïîêðèâ³â ó ïðèñèñíîãî äèñêà.<br />
20(21) Ïîòîâùåííÿ ñïåðåäó òà ïîçàäó äèñêà ... C. goetzae Türk Å. et ., 1957.<br />
ͳìå÷÷èíà. Íà Aphodius prodromus.<br />
21(20) Ïîòîâùåííÿ ò³ëüêè ïî áîêàì äèñêà.<br />
22(23) ̳æ ïîòîâùåííÿìè íå âì³ùàºòüñÿ äèñê. Ðåòèíè î÷åé íå çðîùåí³<br />
.................................. C. tarsoinfracta Türk E. et ., 1957<br />
23(22) ̳æ áîêîâèìè ïîòîâùåííÿìè â³ëüíî âì³ùàºòüñÿ äèñê<br />
............................................ C. longi pes Türk E. et ., 1957<br />
ͳìå÷÷èíà. Åðëàíãåí. Íà Chelidura acanthophagia.<br />
24(19) Áåç ïîòîâùåíü ïîêðèâ³â ïîðÿä ç äèñêîì.<br />
117
25(26) Ãðàíèöÿ ñòåðíàëüíîãî ùèòà ïîïåðå÷íà. Ðåòèíè î÷åé íå çàìêíóò³<br />
.......................................... C. astoma Türk E. et ., 1957<br />
ͳìå÷÷èíà. Íà ìóðàõàõ ormica rufa òà Lasius fuliginosus.<br />
26(25) ßêùî ñòåðíàëüíèé ùèò ³ñíóº, òî éîãî ãðàíèöÿ íàï³âÿéöåïîä³áíà,<br />
åë³ïòè÷íà, àëå íå ïîïåðå÷íà.<br />
27(28) Âåíòðàëüíèé ùèò íå óòâîðþº îêðåìèõ êîêñàëüíèõ ïîë³â<br />
..................................................... C. calliphorae Vitzhum, 1922<br />
ͳìå÷÷èíà. Íà ì'ÿñí³é ìóñ³ Calli phora vomitoria.<br />
28(29) Êîêñàëüí³ ïîëÿ çðîùåí³ óòâîðþþ÷è ÷³òê³ ïðàâ³ òà ë³â³ ïîëÿ.<br />
Ïîêðèâè áåç àáî â ð³çíîìàí³òíîìó ìàëþíêó.<br />
29(30) Ïîêðèâè ãëàäåíüê³ .............................. C. marie Samsinak, 1989<br />
Àôðèêà. Çà¿ð. Íà ìóñ³ Gymnometopina magna [10].<br />
30(31) Ïîêðèâè â ð³çíîìàí³òíîìó ìàëþíêó — øòðèõóâàíí³, ÿìêàõ ³<br />
ò. ï.<br />
31(32) Øê³ðà â ãóñòèõ êðàïêàõ ....... C. waldorfae ain et Jonson, 1974<br />
Àíãë³ÿ. Íà Sminthurus fuscus [11].<br />
32(33) Øê³ðà â ñ³òêîïîä³áíîìó øòðèõóâàíí³.<br />
33(34) Ðîñòðàëüíèé âèñòóï ïðîïîäîñîìè øèðîêèé ïîïåðå÷íèé<br />
....................................................... C. laesionis Mahunka, 1973<br />
Àôðèêà. Ãàíà. Íà ãíîéîâèêàõ [12].<br />
34(35) Ðîñòðàëüíèé âèñòóï íå ïîïåðå÷íèé. Ò³ëî ÿéöåïîä³áíå, àáî àìôîðîïîä³áíå.<br />
35(36) Ò³ëî ð³çêî àìôîðîïîä³áíå ç ÿçèêîïîä³áíîþ îï³ñòîñîìîþ<br />
........................................................ C. thraca Vitzhum, 1922<br />
Áîëãàð³ÿ. Òàòàðñòàí. Íà ìóõàõ â ãíîºñõîâèùàõ.<br />
36(35) Ò³ëî îâàëüíå, ÿéöåïîä³áíå.<br />
37(38) Ïðîïîäîñîìà ó î÷åé ç ð³çêî â³äîêðåìëåíîþ â³ä ³íøèõ ïîêðèâ³â<br />
ïëîùàäêîþ, âêðèòîþ ïîâçäîâæí³ìè ñêëàäêàìè. Ñóñ³äí³ ïîêðèâè<br />
â çíà÷íèõ ïîïåðå÷íèõ êîì³ðêàõ .......... C. reticulata A. Z., 1941<br />
Øâåéöàð³ÿ. Íà áäæîë³ Osmia dalmatica.<br />
38(39) Ìàëþíîê âñ³º¿ ïîâåðõí³ ïðîïîäîñîìè îäíàêîâèé.<br />
39(42) Ïîêðèâè â ãóñòèõ ïîðàõ àáî â çíà÷íèõ (îñîáëèâî íà ã³ñòåðîñîì³)<br />
ÿìêàõ.<br />
40 (41) Ïîêðèâè â ãóñòèõ ïîðàõ. Ïðîïîäîñîìà â ïîïåðå÷íèõ, ã³ñòåðîñîìà<br />
â ïîâçäîâæí³õ êîì³ðêàõ ......... C. hagensis Oudemans, 1911<br />
Òèïîâèé âèä ðîäó. Íàäàºìî éîãî ä³àãíîç ïî [3]. Ãîëëàíä³ÿ. Â<br />
ãíèþ÷îìó ëèñò³.<br />
41(40) Ïîêðèâè â çíà÷íèõ ãëèáîêèõ ÿìêàõ ... C. biscupliurata Mahunka, 1974<br />
Àôðèêà. Ãàíà. Íà êîìàõàõ êîêîñîâèõ ïàëüì [13].<br />
42(39) ѳòêà ïîêðèâ³â áåç ïîð òà ÿìîê.<br />
42(43) Î÷³ çíàõîäÿòüñÿ â ºäèí³é ðåòèíàëüí³é ïëÿì³<br />
....................................................... C. bulgarica Storkan, 1935<br />
Áîëãàð³ÿ. Ñìîëåíñüêà îáë. Íà êîìàõàõ.<br />
44(43) ̳æ ÿâíèìè î÷èìà çíà÷íèé ïðîì³æîê.<br />
45(46) ³äñòàíü ì³æ î÷èìà äîð³âíþº ïîïåðå÷íèêó ¿õ ë³íç<br />
.................................................... C. heterocoma (Mich.) 1903<br />
118<br />
Â. Ä. Ñåâàñòüÿíîâ, Ä. À. ʳâãàíîâ, Äæàðìà Íàä³ÿ Àë³
Îãëÿä òà âèçíà÷íèê êë³ù³â ðîäó Calvolia<br />
46(45) ³äñòàíü ì³æ î÷èìà ïåðåâèùóº ïîïåðå÷íèê ¿õ ë³íç<br />
............................................... C. elisabethae Zachvatkin, 1941<br />
Áóõàðà. óïîïóñè íà ñë³ïóøîíö³ Ellobius talpinus<br />
Òàêèì ÷èíîì, ó ïðåäñòàâëåíîìó âèçíà÷íèêó ïðåäñòàâëåíî 38 â³äîìèõ<br />
íàì âèä³â ðîäó, ç ÿêèõ: ó øåñòè âèä³â îïèñàí³ ñàìö³, ñàìèö³ òà<br />
ã³ïîïóñè; ó äâîõ âèä³â — ñàìö³ òà ñàìèö³, ó ï'ÿòè — ò³ëüêè ñàìèö³,<br />
³ ó 25 âèä³â — ò³ëüêè ã³ïîïóñè.<br />
ßê ñïðàâåäëèâî â³äì³÷àâ À. À. Çàõâàòê³í ùå ó 1941 ðîö³ [2], ã³ïîïóñè<br />
º ÷èñòî ôîðåòîìîðôíîþ ñòà䳺þ ó æèòò³ êë³ù³â, òîáòî íàñë³äêîì<br />
ïðèñòîñóâàííÿ äî çàêð³ïëåííÿ òà ïåðåñóâàííÿ íà õàçÿ¿í³.  çâ'ÿçêó ç<br />
öèì íå äóæå êîðåêòíî áóäóâàòè ô³ëîãåíåòè÷íó ñèñòåìó êë³ù³â (â<br />
òîìó ÷èñë³ ³ ðîäó Calvolia), âèêîðèñòîâóþ÷è ìîðôîëîã³þ ã³ïîïóñ³â.<br />
Òîìó ìè áà÷èìî ìîæëèâ³ñòü óòî÷íåííÿ ñèñòåìàòè÷íîãî ïîëîæåííÿ<br />
ïðåäñòàâíèê³â äîñë³äæåíîãî íàìè ðîäó íå ò³ëüêè çà ðàõóíîê îïèñó<br />
íîâèõ êë³ù³â, ùî çíàéäåí³ ó ïðèðîä³, àëå é çà äîïîìîãîþ ëàáîðàòîðíèõ<br />
äîñë³äæåíü (âèâåäåíí³ ç ã³ïîïóñ³â ñàìö³â òà ñàìèöü). Öå íåìèíó÷å<br />
ïðèçâåäå äî âèÿâëåííÿ âèä³â, ÿê³ áóëè îïèñàí³ äâ³÷³ — ïî ã³ïîïóñàì<br />
òà ïî ³ìàãî, à òàêîæ äî ïåðåì³ùåííÿ äåÿêèõ âèä³â äî ³íøèõ<br />
ðîä³â (êîëè ã³ïîïóñè ñõîæ³, à äîðîñë³ ñòà䳿 çíà÷íî â³äð³çíÿþòüñÿ).<br />
˳òåðàòóðà<br />
1. Oudemans À. Ñ. Acarologische Aanteekeningen. 38 // Entomol Ber. — 1911. — V. 3, ¹ 2. —<br />
P. 183–191.<br />
2. Çàõâàòêèí À. À. Òèðîãëèôîèäíûå êëåùè. Tyroglyphoidea (Ôayía ÑÑÑÐ. Ïàóêîîáðàçíûå.).<br />
— Ì.-Ë.: ÀÍ ÑÑÑÐ, 1941. — 494 ñ.<br />
3. ain A. Notes sur les hypopes des Saproglyphidae (Acarina, Sarcoptiformes) II. Redefinition<br />
des genres // Acarologia. — 1972. — V. XIV, f. 2. — P. 225–259.<br />
4. Turk E. et Turk . Systematik und Okologie der Tyroglyphiden Mitteleuropas // I. J. Stammer:<br />
— Leipzig: Acad. Veelagsges, 1957. — T. 1, Abt. 1. — P. 1–231.<br />
5. Ñåâàñòüÿíîâ Â. Ä. Ââåäåíèå â èçó÷åíèå êëåùåé àíîåòèä // Çîîëîãè÷åñêèé æóðíàë. —<br />
1962. — Ò. 62, âûï. 3. — C. 1338–1345.<br />
6. Dusbabek . Some new Species of Tyroglyphoid mites (Acarina, Tyroglyphoidea), parasitic bats<br />
// Acta socient. zool. Bohemoslovenical. — 1964. — V. 28, ¹ 3. — P. 220–233.<br />
7. Âîëãèí Â. È. Íîâûå âèäè êëåùåé ñåì. Saproglyphidae (Acariformes, Acaroidea) // Ïàðàçèòîë.<br />
ñá. — 1980. — Ò. 29. — Ñ. 159–168.<br />
8. Ochima Shiro. New Acaroid mites in hause dust // Annotationes zool. Japonensis. — 1979.<br />
— V. 52, ¹ 4. — P. 240–245.<br />
9. Cooreman J. Notes et observations sur quelques acariens infeodes aux coleopteres scolytides<br />
de la faune Belge // Bul. inst. roy. Scienc. natur. de Belgique. — 1963. — V. 39, ¹ 30. —<br />
P. 1–33.<br />
10. Samsinak K. Mites of flies of the family Slhaeroceridae // Acarologia. — 1989. — V. 30,<br />
¹ 2. — P. 85–105.<br />
11. ain A., Jonson D. Three new species of hi popi phoretik on Springtais (Collembola) in<br />
England (Acari: Acaridae) // J. nat. Hist. — 1974. — V. 8. — P. 411–420.<br />
12. Mahunka S. Auf insecten lebende milbeh (Acari: Acarida, Tarsonemida) aus Africa. I. //<br />
Acta zool. Acad. Scient. Hungaricae. — 1973. — V. 19, ¹ 1–2. — P. 75–123.<br />
13. Mahunka S. Auf insecten lebende milben (Acari: Acarida, Tarsonemida) aus Africa. III //<br />
Acta zool. Acad. Scient. Hungaricae. — 1974. — V. 20, ¹ 1–2. — P. 137–156.<br />
119
120<br />
Â. Ä. Ñåâàñòüÿíîâ, Ä. À. Êèâãàíîâ, Íàäåæäà Àëè Äæàðìà<br />
Îäåññêèé íàöèîíàëüíûé óíèâåðñèòåò èìåíè È. È. Ìå÷íèêîâà,<br />
êàôåäðà çîîëîãèè,<br />
óë. Äâîðÿíñêàÿ, 2, Îäåññà, 65026, Óêðàèíà<br />
ÎÁÇÎÐ È ÎÏÐÅÄÅËÈÒÅËÜ ÊËÅÙÅÉ ÐÎÄÀ <strong>CALVOLIA</strong><br />
(<strong>SAPROGLYPHIDAE</strong>)<br />
Ðåçþìå<br />
Ðîä Calvolia (Saproglyphidae) ñîñòîèò èç 38 âèäîâ êëåùåé, èç êîòîðûõ ó øåñòè<br />
âèäîâ èçâåñòíû ñàìêè, ñàìöû è ãèïîïóñû; ó äâóõ ñàìöû è ñàìêè, ó ïÿòè òîëüêî<br />
ñàìêè è ó 25 âèäîâ îïèñàíû òîëüêî ãèïîïóñû. Äëÿ êàæäîãî âèäà ïðèâîäèòñÿ åãî<br />
ãåîãðàôè÷åñêîå è ýêîëîãè÷åñêîå ìåñòî îáíàðóæåíèÿ. Calvolia hebeclinii Sicher 1889<br />
èñêëþ÷àåòñÿ èç ñïèñêà âèäîâ ðîäà êàê nomen dubium, à Calvolia zacheri Oudemans<br />
1930 êàê îøèáî÷íî îïðåäåëåííûé àâòîðîì âèä.<br />
Êëþ÷åâûå ñëîâà: êëåùè, ðîä, âèä, ãèïîïóñû, îïðåäåëèòåëü.<br />
V. D. Sevastianov, D. A. Kivganov, Nadezhda Ali Jarma<br />
Odessa National University by I. I. Mechnikov, Zoological Department,<br />
Dvoryanska St., 2, Odesa, 65026, Ukraine<br />
THE REVIEW AND KEY OR MITES O <strong>CALVOLIA</strong><br />
(<strong>SAPROGLYPHIDAE</strong>) GENUS<br />
Summary<br />
The Calvolia (Saproglyphidae) genus consists of 38 mites species, from which the<br />
females, males and hypopuses are known for six species; for two — males and females,<br />
for five — only females, and for 25 species only hypopuses were described. or every<br />
species, its geographical and ecological place of revealing is given. The Calvolia hebeclinii<br />
Sicher 1889 is excluded from the species list of genus as nomen dubium, the Calvolia<br />
zacheri Oudemans, 1930 as an author's erroneously determined species<br />
Keywords: mites, genus, species, hypopuses, key.<br />
Â. Ä. Ñåâàñòüÿíîâ, Ä. À. ʳâãàíîâ, Äæàðìà Íàä³ÿ Àë³
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