Sources of records for Albania - University of Toronto Mississauga

Sources of records for Albania 

in Flora Europaea 

1. Introduction - page 2 

compiled by Peter W. Ball 

CONTENTS 

2. Flora Europaea vol. 1 - page 6 


4. Flora Europaea vols 3 and 4 - page 40 


6. Plant collections from Northern Albania by Bruno Schütt - page 78

Part 1: Introduction 

The following lists of the flora of Albania were compiled in the period 1961-1968. 

During the early stages in the preparation of manuscripts for Flora Europaea volume 1 it 

became apparent that there was no reliable source of information for geographical data 

for countries in the Balkan peninsula except for Bulgaria. The main Flora covering the 

region was Hayek (1924-1931), but Hayek gives distributions based on areas defined by a 

combination of phytogeographical considerations and pre-1914 political boundaries. 

While most of these areas are wholly within the post 1945 political boundaries some, 

particularly Albania, Epirus, Macedonia and Thrace transcend post 1945 political 

boundaries and so do not present reliable evidence for the occurrence of a species within 

the political areas recognized in Flora Europaea (Webb 1963). This created difficulties 

for two countries, Turkey and Albania. D. A. Webb compiled a list for Turkey-in-Europe 

(which was part of Hayek’s Thrace), and this was published by Webb (1966). I took on 

the task of compiling a list for Albania (see Webb, 1963). 

Although Hayek recognized an area he called Albania this area differed significantly 

from the post 1919 boundaries of Albania. Hayek’s Albania extended the north-eastern 

boundary considerably further east than the post-1919 political boundary, including all of 

the mountain range often referred to as “Northern Albanian Alps”. Much of this mountain 

range is in Serbia and Macedonia (particularly the part named !ar Planina) and at the 

time Hayek was published this portion of the range was much more extensively explored 

than the part which remained in Albania. At the southern boundary Hayek drew the 

boundary slightly further north than the post 1919 boundary, including this part of 

Albania in Epirus, which is primarily part of post 1919 Greece. Following the end of the 

first world war several collecting expeditions were sent to Albania resulting in many 

additions to the flora which were only partially included in Hayek. A further source of 

confusion arises from many pre-1914 collections in which the labels state that the 

collection was from Albania when in fact it came from Serbia or Macedonia. Some of 

these collections are from areas which are now along political boundaries (especially high 

mountain peaks such as Korab, Koritnik and Pastrik) so introducing an element of 

uncertainty about which country they came from. During the post-1920 period these high 

mountain border peaks were often out-of-bounds for plant collectors so it was not 

possible to verify these older reports of the occurrence of a species in Albania. 

The list was compiled from published papers, mostly derived from the extensive 

bibliography published by Markgraf (1932) together with a supplement Markgraf (1963). 

Some post-1945 collections and publications could not be included as they were not 

available to me at the time. However some did become available after the publication of 

volumes 1 and 2 of Flora Europaea so the list for volumes 3 and 4 is more complete than 

those for the earlier volumes. The bibliography indicates (*) those publications which 

became available at a later time. For a variety of reasons the list was never completed for 

volume 5 of Flora Europaea. The information that had been accumulated by 1968 is 

included here, but it does not coincide with the records published in Flora Europaea 

volume 5. There are about 50 species included in Flora Europaea volume 5 which are not 

in that list. It is likely that these were derived from Flora of Tirana (Tir) and from other 

sources that became available after 1970. However the list includes about 90 species 

2

eported from Albania but which are not reported for Albania in Flora Europaea. Some of 

these appear to be the only report of the species from Albania. 

In addition to published records I was given access to unpublished material. The most 

important of these was a manuscript by Bruno Schütt which was made available by the 

Curator of Botany of the Uebersee Museum, Bremen (BREM). This manuscript is 

included as part 6 of this document. Another important addition was a complete list of the 

collections made by Alston and Sandwith kindly provided by N. Sandwith. Although 

Alston & Sandwith published a proportion of their records this list contained a number of 

records which appear to be the only reports of a species from Albania. I was also given 

access to complete lists of collections made in Albania during the 1930’s by several 

different people and housed at Royal Botanic Gardens, Kew. A few reports of these 

collections were published by Turrill (1932, 1939) and Chator & Turrill (1934) but most 

are unpublished. 

It is important to understand that this list does not purport to represent a comprehensive 

list of the flora of Albania. Its main purpose was to ensure that species reported from 

Albania in Flora Europaea were based on evidence which could be verified either in print 

or by an author examining herbarium specimens. Unfortunately some important 

collections of Albanian plants were destroyed during the second World War and later, 

particularly those of Markgraf at B. Some material at W was also lost in 1945, and much 

material at Budapest (BP) was lost in 1956. 

The list is arranged in the sequence published by Hayek (1924-1932), the species being 

numbered according to Hayek. Because Flora Europaea followed a slightly different 

family sequence from Hayek a few families are out of sequence when they occur in a 

different volume of Flora Europaea. For example volume 1 of Hayek is covered by 

volumes 1 and 2 of Flora Europaea and volume 2 of Hayek is covered by volumes 3 and 

4 of Flora Europaea. Within a family the genera and species are arranged in the Hayek 

sequence, new genera being placed at the end of the family and new species at the end of 

the genus. Species or infra specific taxa included in the list without a reference are 

reported from a minimum of 3 sources by at least two independent authors. Species 

reported from one or two sources, or reported by only one author, include appropriate 

references. It is important to note that one source may report a species from several 

localities and in a few cases a species reported by several different authors may be 

restricted to a single locality. Post-1919 collections are assumed to be in present-day 

Albania although some are from border mountains. Pre-1919 collections were traced to 

present-day Albania as far as it was possible to do so. Where there was a considerable 

element of doubt about the locality, records were not included. Baldacci (1899), and 

Baldacci (1901) are particularly difficult as Baldacci (1899) collected in areas that are 

now mainly in Greece and Baldacci (1901) crossed into Montenegro although most of 

these collections appear to be from Albania. Many of the D"rfler and other collections 

reported by Hayek (1917) and Hayek (1924) are from boundary areas so some care has to 

be exercised with these records and in a few cases it is not possible to be sure which 

country a collection came from. Such records are indicated as boundary or border 

records, especially when they are the only available record for a species, although many 

of these were verified by the unpublished records of Schütt. 

Below is a key to the abbreviations used in the list for sources of records together with 

references to those publications also mentioned above. Extensive bibliographies for 

3

Albania were published by Markgraf (1932, 1963) and these should be consulted for 

older sources of information. 

List of abbreviations and bibliography. 

A&S Alston, A.G. H. & N.Y. Sandwith, 1940. Results of two Botanical 

Expeditions to South Albania. Jour. Bot. (London) 78. 

Unpublished list by N. Y. Sandwith. 

B94 Baldacci, A., Rivista critica della collezione botanica fatta nel 1892 in 

Albania. Malpighia 8. 

B 94b Baldacci, A., 1894. Contributo alla conscenza della Flora dalmata, 

montenegrina, albanese, epirota e greca. N. Gior. Bot. Ital. 1. 

B96. Baldacci, A., 1896. Rivista della collezione botanica fatta nel 1894 in 

Albania. Bull. Herb. Boiss. 4. 

B99 Baldacci, A., 1899. Rivista dalla collezione botanica fatta nel 1896 in 

Albania. N. Giorn. Bot. Ital. 6. 

B01 Baldacci, A., 1901. Rivista della collezione botanica fatta nel 1897 

nell’Albania settentrionale. Mem. Acad. Sci. Ist. Bologna, ser. 9, 5. 

B17 Baldacci, A., 1917. Itinerari albanasi. Rome. 

BB Baldacci, A. & Beguinot, 1918. Contributo alla flora autumnale et invernale 

del dintorni di Vallona. N. Giorn. Bot. Ital. 25. 

Bm33 Bornmüller, J., 1933. Zur Flora von Montenegro, Albanien und 

Macedonien. Magyar Bot. Lapok 32. 

Chator, D. A., & W. B. Turrill, A Contribution to the Flora of Central 

Albania. Kew Bull. 1934. 

Form. Formanek, E., 1895. Beiträg zur Flora von Albanien, Korfu und Epirus. 

Verh. Natf. Vereins Brünn. 33 

See Vandas, 1909. 

Gr Grimus, V. von, 1871. Beiträge zur Flora Albaniens. Verh. Zool.-Bot. Ges. 

Wien 21. 

Giu Giusseppi. Specimens at Kew. See Chator & Turrill and Turrill. 

H17 Hayek A. von, 1917. Beiträge zur Kenntnis der Flora des albanischmontenegrinischen 

Grenzgebietes. Denschr. Akad. Wiss. Wien, Math.natw. 

Kl. 94 

H24 Hayek, A. von, 1924. Zweiter Beiträg zur Kenntnis der Flora Albanien. 

Denkschr. Akad. Wiss. Wien, Math.-natw. Kl. 99. 

Hayek, A. von, Prodromus Florae Peninsulae Balcanicae. Vol. 1, 1924, Vol. 

2, 1927, Vol. 3, 1932 (completed by F. Markgraf). 

Held. Heldreich, von, 1879. Ein Beitrag zur Flora von Epirus, geliefert von Herrn 

N. K. Chodzes. Verh. Bot. Vereins Prov. Brandenburg 21. Sitzber. 

Hep Hepburn, I. Specimens at K. Some published by Turrill and Chator & Turrill 

J20 Janchen, E., Vorarbeiten zu einer Flora der Umgebung von !kodra in Nord- 

Albanien. Österr. Bot. Zeitschr. 69. 

4

Jav. Csiki, E., A. Jávorka & E. B. Kümmerle, 1926. Additamenta ad Floram 

Albaniae. Pteridophyta by E. Kümmerle, Anthophyta by A. Jávorka. 

Budapest. 

M27 Markgraf, F., 1927. An den Grenzen des Mittelmeergebiets: 

Pflanzengeographie von Mittelalbanien. Feddes Repert. Beih. 45. 

M32 Markgraf, F., 1932. Pflanzengeographie von Albanien. Bibliotheca Botanica 

105. 

M42* Markgraf, F., 1942. Reisebericht aus Albanien. Notizbl. Bot. Gart. Mus. 

Berlin-Dahlem. 15. 

Markgraf, F., 1963. Bericht Albanien 1961. Webbia 18. 

K48 Kitanov, B., 1948. Contribution á l’étude de la Flora d’Albanie de l’Est. Ann. 

Fac. Phil. Univ. Skopje Sect. Sci. Nat. 1. 

Pamp. Pampanini, R., 1915. Contributo alla conoscenza della Flora d’Albania. Bull. 

Soc. Bot. Ital. 84. 

Penn. Pennington. Specimens at Kew. See Chator & Turrill and Turrill. 

R. Rechinger fil., K. H., 1939. Zur Flora von Albanien und Mazedonien. Feddes 

Repert. Sp. Nov. 47. 

Rohl Rohlena, J., 1937. Beitrag zur Flora Albaniens. V#stník Kralovské $eské 

Akad. Spolo%nosti Nauk Trída Matemat. P&irod. 1937. 

Rod Rodriguez, 1923. Plantes de l’Albanie Moyenne. Bull. Mus. Hist. Nat. Paris 

Sch Schütt, B. Unpublished list. See part 6 below. 

Tir* Paparisto, K., Xh. Qosja & M. Demiri, 1962. Flora e Tiranes. Tirana. 

Turrill, W. B., 1932. A Contribution to the Flora of Central Albania. Kew Bull. 

1932. 

Turrill, W. B., 1939. Two new Species and New or Important Records. Kew 

Bull. 1939. 

Vandas, K., 1909. Reliquiae Formanekianae. Brunae (Brno). 

Webb, D. A., 1963. The need for check-lists of the flora of certain European 

territories. Webbia 18. 

Webb, D. A., 1966. The Flora of European Turkey. Proc. Roy. Irish Acad. 

65B. 

Weiss* Weiss, E., 1866. Floristisches aus Istrien, Dalmatien und Albanien. Verh. 

Zool.-Bot. Ges. Wien. 16 

5

Part 2: Flora Europaea vol. 1 

Pteridophyta 

1. Lycopodium selago K48 

6. Selaginella selaginoides K48 

7. S. helvetica M27 

9. Equisetum maximum 

10. E. arvense 

12. E. palustre 

13. E. limosum H24 

14. E. ramosissimum 

15. E. hiemale 

22. Ophioglossum vulgatum M27 M32 

24. Botrychium lunaria 

26. B. matricariaefolium H24 

28. Osmunda regalis M32 Bm33 

29. Polypodium vulgare 

33. Pteridium aquilinum 

34. Adiantum capillus-veneris 

35. Notholaena marantae 

38. Cheilanthes persica 

39. Cryptogramma crispa K48 

42. Phyllitis scolopendrium M27 M32 

46. Asplenium viride 

48. A. trichomanes A. var. trichomanes B. var. csikii Jav 

53. A. adiantum-nigrum I. subsp. adiantum-nigrum II. sibsp. serpentini III. subsp. 

onopteris 

54. A. fissum 

55. A. ruta-muraria 

56. A. lepidum Jav 

57. A. septentrionale 

55 x 56. A. x javorkae Jav 

58. Ceterarch officinarum 

59. Athyrium filix-foemina 

61. Dryopteris linnaeana A&S H17(border) 

62. D. robertiana B01 Sch 

66. D. filix-mas 

67. D. villarsii I. subsp. rigida II. subsp. pallidaI 

68. D. spinulosa K48 

69. Polystichum lonchitis 

69 x 70. P. x illyricum 

70. P. lobatum 

6

71. P. setigerum 

74. Cystopteris fragilis 

75. C. regia 

79. Marsilea quadrifolia Sch 

Gymnospermae 

81. Taxus baccata 

83. Juniperus communis I. subsp. communis II. subsp. nana 

84. J. oxycedrus 

85. J. macrocarpa B96 

86. J. foetidissima 

89. J. phoenicea Sch Gr? 

90. Cupressus sempervirens BB 

91. Abies alba 

A. borisii-regis A&S 

94. Picea excelsa Jav Sch 

97. Pinus halepensis B96 M32 

100. P. mughus 

101. P. heldreichii M32 Bm33 

102 I. P. nigra 

104. P. pinea M32 

105. P. peuce 

106. Ephedra campylopoda 

107. E. distachya 

Betulaceae 

109. Betula pendula H24(boundary) 

112. Alnus incana 

113. A. glutinosa 

114. Carpinus betulus 

115. C. orientalis 

116. Ostrya carpinifolia 

117. Corylus colurna A&S 

118. C. avellana 

Fagaceae 

120. Fagus silvatica 

122. Castanea sativa 

123. Quercus cerris (vars. cerris, austriaca, pseudocerris) 

124. Q. macedonica 

125. Q. aegilops B94 B94b 

126. Q. pseudosuber M27 

127. Q. coccifera 

7

128. Q. ilex 

130. Q. conferta 

131 II. Q. lanuginose 

136. Q. sessiliflora 

137. Q. robur B94 M32 

123 x 124 R 

130 x 136 Jav 

Juglandaceae 

139. Juglans regia 

Salicaceae 

140. Populus alba 

142. P. tremula 

143. P. italica 

144. Salix amygdaloides Sch 

145. S. pentandra Bm33 

147. S. alba 

154. S. arbuscula Jav Sch 

155. S. hastata Jav 

159. S. elaeagnos 

162. S. purpurea I. subsp. purpurea A&S II. subsp. amplexicaulis Jav M27 

163 I. S. retusa 

165. S. reticulata 

Moraceae 

166. Morus alba J20 Sch (?cult) 

167. M. nigra Sch (?cult) 

168. Ficus carica 

169. Humulus lupulus J20 Sch 

Ulmaceae 

171. Ulmus campestris (incl. U. glabra, U. foliacea) 

173. U. scabra B01 

174. U. laevis H24 

176. Celtis australis 

Urticaceae 

179. Urtica dioica 

180. U. membranacea BB H24 

181. U. urens 

8

182. U. pilulifera J20 Sch 

184. Parietaria erecta 

185. P. ramiflora 

Santalaceae 

189. Osiris alba 

191. Thesium alpinum B01 K48 

194. T. linophyllum M27 H24 (boundary) 

195. T. divaricatum 

196. T. ramosum A&S Jav 

198. T. parnassi 

200. T. auriculatum Jav Sch 

203. T. bergeri B99 

Loranthaceae 

206. Loranthus europaeus 

207. Arceuthobium oxycedri 

209. Viscum laxus 

Chenopodiaceae 

353. Polycnemum majus B94 J20 

356 I. Beta maritime 

359. Chenopodium botrys 

362. C. vulvaria Jav 

363. C. hybridum Jav 

365. C. opulifolium 

366 I. C. album 

368. C. urbicum B96 

370. C. glaucum B94b A&S 

371. C. bonus-henricus 

372. C. foliosum Hep 

375. Atriplex hortensis J20 

380. A. hastata M27 A&S 

381. A. rosea B94 

382. A. tartarica B96 

384. A. portulacoides 

388. Camphorosma monspeliaca B17(1894) 

391. Kochia prostrata A&S (sub K. scoparia) 

393. K. scoparia H24 

401. Halocnemum strobilaceum A&S B96 

402. Arthrocnemum glaucum 

403. Salicornia fruticosa (incl. f.radicans) 

404. S. herbacea 

9

406. Suaeda fruticosa B96 

408. S. maritima 

409. S. splendens 

413. Salsola kali A. var. kali B17(1894) A&S B. var. tragus B17(1894) 

414. S. soda B94b A&S 

417. Petrosimonia crassifolia 

Caryophyllaceae 

442. Herniaria hirsute 

443. H. incana 

445. H. parnassica 

446. H. glabra 

448. Paronychia chionaea 

449. P. kapela 

450. P. cephalotes Penn Sch 

457. Polycarpon tetraphyllum J20 Sch 

460. Spergularia salina M27 Sch 

462. S. rubra 

463. S. diandra B96 

467. Scleranthus perennis Sch Jav 

468. S. neglectus 

470. S. uncinatus B01 H17 

472.S. annuus I. subsp. annuus II. subsp. verticillatus B01 J20 

473. Queria hispanica A&S 

479. Minuartia bosniaca Sch 

480. M. setacea Hep 

485. M. velenovskyi H17 

486. M. tenuifolia H17 

488. M. mediterranea 

492. M. verna I. subsp. montana II. subsp. leptophylla A&S Jav IV. subsp. gerardi 

496. M. recurva I. subsp. juressi M27 M32 II. subsp. recurva 

498 I. M. graminifolia subsp. clandestina 

499 I. M. stellata A&S 

503. M. baldaccii subsp. I, II, III. 

504 (1 st ) M. garckeana M27 

513. Arenaria cretica B94 B96 (A. gracilis sensu Baldacci) 

516. A. rotundifolia var. pauciflora B94 B96 var. pancicii B01 Jav 

517. A. biflora 

518. A. serpyllifolia 

519. A. leptoclados 

522. A. conferta 

*A. serpentinii A&S 

525. Moehringia trinervia 

530. M. malyi Jav 

532. M. ciliata Jav Sch 

10

533. M. muscosa B01 Sch 

534. Sagina subulata 

535. S. saginoides B01 

536. S. ciliata B01 

538. S. procumbens Sch 

542. Moenchia graeca Sch 

543. M. mantica 

544. Cerastium dinaricum var. velebiticum H24 Sch 

547. C. grandiflorum 

549. C. banaticum subsp. banaticum K48 Sch subsp. alpinum A&S 

550. C. decalvans 

552. C. arvense I. subsp. arvum B99 II. subsp. ciliatum B01 Sch 

553. C. moesiacum 

554. C. alpinum Jav 

555. C. hekuravense Jav Sch 

556. C. lanatum H24 Sch 

557. C. silvaticum K48 

558 I. C. caespitosum Jav Sch 

559.C. brachypetalum vars. galandulosum, luridum frequent; vars. tauricum, roeseri, 

brachypetalum single records only 

560. C. viscosum 

563. C. glutinosum I. subsp. glutinosum II. subsp. pallens H17 

566. C. semidecandrum H24 

567. C. litigiosum 

570. C. rectum M27 Sch 

578. C. cerastioides 

*C. corcyrense A&S (C. campanulatum sensu A&S) 

579. Myosoton aquaticum 

580. Stellaria nemorum I. subsp. nemorum K48 Sch II. subsp. glochidosperma Jav 

581. S. apetala I. subsp. apetala J20 II. subsp. media III. subsp. neglecta H24 K48 

583. S. graminea 

586. Saponaria bellidifolia 

587. S. glutinosa A&S Hep 

588 B. S. haussknechtii var. major 

589. S. officinalis 

590. S. calabrica 

592 I. Vaccaria pyraminata 

603. Gypsophila sperguliifolia 

609. Tunica illyrica 

611. T. saxifraga 

612. T. rigida A&S 

616. Kohlrauschia velutina A&S Sch 

617. K. prolifera 

618. K. glumacea 

619. Dianthus armeria 

621. D. armeriastrum 

11

622. D. corymbosum Sch 

625 II. D. viscidus var. grisebachii 

631. D. liburnicus J20 Sch 

634. D. suskalovicii Sch 

641. D. carthusianorum 

642. D. sanguineus H24 Sch 

645. D. cruentus 

646. D. turcicus M27 Sch 

647. D. holzmannianus M32 Sch 

648. D. tristis 

649. D. lateritius R B99 

654. D. leucophoenicum 

658. D. stribrnyi Sch 

659. D. zernyi H24 

661. D. stenopetalum A&S 

662. D. pinifolius I. subsp. serbicus II. subsp. lilacinus R 

670. D. haematocalyx I. subsp. haematocalyx B94 Hep III. subsp. sibthorpii A&S B96 

IV. subsp. pindicola A&S *subsp. jirasekii Roh 

682. D. viridescens H24 

683. D. deltoides (vars. C & D) 

690 II. D. dalmatica 

693. D. sylvestris I. subsp. tergestinus var. brevicalyx(bertisceus) II. subsp. nodosus B01 

III. subsp. sylvestris 

695 II. D. strictus subsp. bebius M27 

696. D. integer A. var. integer B. var. minutiflorus Jav 

698. D. nicolai Sch 

* D. ambiguus R. 

715. Velezia rigida B99 K48 

717 I. Drypis spinosa 

718. Cucubalus baccifer Jav 

720 b. Silene caesia var. pindica A&S Sch 

723. S. fabarioides A&S 

726. S. vulgaris subsp. II, III, VI, VII 

734. S. conica 

736. S. graeca 

738. S. vespertina H24 

739. S. gallica 

741. S. nocturna Gr 

742. S. remotiflora A&S 

743. S. trinervia 

746. S. colorata Pamp BB(sericea) 

747. S. apetala H24 

750. S. quadridentata subsp. III many records. Subsp. I, II, VIIb, VIII, IX, XI, markgrafii 

recorded once or twice. 

751. S. macrantha 

757. S. waldsteinii forma croatica M32 

12

758. S. multicaulis A. var. multicaulis B. var. serbica 

759. S. saxifraga I. subsp. saxifraga II. subsp. parnassica A&S B. var. hayekiana Sch 

760. S. fruticulosa Penn (many Baldacci records, presumably included under 759 by 

Hayek) 

767. S. graminea H24 K48 

770. S. asterias Sch 

772. S. armeria 

785. S. ungeri A&S 

786. S. sedoides B94 

788. S. coeli-rosa Gr 

789. S. cretica M27 

796. S. sendtneri 

797. S. otites I. subsp. otites II. subsp. parviflora Sch 

799. S. roemeri 

800. S. ventricosa A&S 

803. S. radicosa 

809. S. longiflora I. subsp. longiflora H24 II. subsp. staticifolia A&S 

814. S. viridiflora Jav 

815. S. nutans 

816. S. paradoxa 

818. S. italica I. subsp. italica II. subsp. nemoralis M32 

820. S. cephallenia 

822. S. gigantea var. viridescens A&S 

824. S. schwarzenbergeri A&S 

828. Melandrium nemoralis Sch 

829. M. album I. Sch II. many III. A&S 

831. Lychnis viscaria 

832. L. atropurpurea B01 Penn 

833 (2 nd ). L. coronaria 

834. L. flos-cuculi I. subsp. flos-cuculi J20 Sch II. subsp. subintegra A&S H24 

835. Agrostemma githago J20 Sch 

Lauraceae 

838. Laurus nobilis BB 

Aristolochiaceae 

839. Asarum europaeum Sch 

844. Aristolochia clematitis 

845. A. longa M27 

846. A. pallida 

847. A. macedonica M32 

848. A. rotunda 

13

Berberidaceae 

851. Berberis vulgaris 

853. Epimedium alpinum 

Paeoniaceae 

857. Paeonia corallina H24 Jav 

858. P. officinalis A&S Sch 

Ranunculaceae 

862. Helleborus cyclophyllus BB Roh 

864. H. odorus I. subsp. odorus II. subsp. multifidus 

865. H. viridis M32 

869. Trollius europaeus M32 

872 II. Caltha palustris subsp. laeta 

878 B. Aquilegia amaliae var. dinarica H24 

880.A. vulgaris probably all var. C 

883. Nigella arvensis 

889. N. damascena 

892 III. Aconitum pantocsekianum H17 (border) 

903. Delphinium staphisagria A&S 

904. D. fissum H24 (border) Sch 

907. D. halteratum B96 Bm 

908. D. peregrinum 

913. Consolida ajacis 

914. C. orientalis Hep Sch 

919 II. C. regalis subsp. paniculate 

dubious record for 919 I. 

922. Actaea spicate 

924. Anemone hepatica 

925. A. blanda I. subsp. appeninum II. subsp. blanda Rod Penn 

926. A. nemorosa 

927. A. narcisiflora Sch Jav (border) 

934. A. hortensis 

941. Clematis vitalba 

943. C. flammula 

946. C. viticella 

948. Thalictrum aquilegifolium 

951. T. minus (several vars.) 

952.T. elatum (incl. vars. saxatile, olympicum) 

957. T. lucidum 

958. T. flavum 

957 x 958. B96 J20 

961. Ficaria verna 

14

964. Ranunculus wettsteinii H24 

965. R. crenatus 

R. magellensis H17 Sch 

969. R. platanifolius 

971. R. thora 

R. scutatus Jav 

973. R. brevifolius 

985. R. millefoliatus A. var. millefoliatus B. var. garganicus 

987. R. psilostachys 

995. R. ophioglossifolius I. subsp. ophioglossifolius II. subsp. fontanus H24 

1001. R. serbicus A&S 

1002. R. lanuginosus 

1004. R. acris A&S Jav 

1005. R. degeni Jav 

1006. R. oreophilus I. subsp. concinnatus II. subsp. croaticus B96 III. subsp. oreophilus 

B94 A&S 

1007. R. montanus subsp. I & II 

1008 B. R. demissus var. graecus A&S B94 

1010. R. velutinus A&S H24 

1011. R. hayekii H 17 Sch 

1012. R. polyanthemus I. subsp. polyanthemus II. subsp. breyninus 

1013. R. repens 

1014. R. bulbosus subsp. hirsutus Sch 

1015. R. neapolitanus 

1017. R. sardous 

R. trilobus Penn 

1018. R. marginatus 

1021. R. muricatus 

1024. R. arvensis 

1033. R. paucistamineus (paucistamineus and drouetii reported by H17) 

1042. Adonis annua BB? 

1045. A. flammea 

Nymphaceae 

1046. Nymphaea alba 

1048. Nuphar luteum J20 Sch 

Ceratophyllaceae 

1050. Ceratophyllum demersum 

Papaveraceae 

1053. Chelidonium majus 

1056. Glaucium flavum A&S F 

15

1060. Papaver rhoeas J20 Sch 

1063. P. dubium 

1071. P. apulum 

1076. Hypecoum grandiflorum Rod? (Kaptura de Martanesh) 

1083. Corydalis ochroleuca I. subsp. ochroleuca II. subsp. leiosperma 

1086. C. cava I. subsp. marschalliana II. subsp. blanda Gui 

1090. C. solida II. subsp. solida III. subsp. densiflora 

1095. Fumaria judaica A&S 

1097. F. officinalis 

1097 D. F. densiflora H24 

1100. F. schrammii A&S 

1101. F. vaillantii Heldr. M27 

1104. F. thuretii var. pikermiana A&S 

1106. F. capreolata 

* F. ragusina A&S 

Capparidaceae 

1108. Capparis rupestris A&S 

1109. C. sicula B94b B96 

Cruciferae 

1110. Sisymbrium strictissimum Jav 

1115. S. orientale A&S B94b 

1116. S. altissimum Jav 

1118. Chamaeplium officinale 

1120. C. polyceratium A&S Jav 

1122. Erysimum pusillum 

1126. E. diffusum 

1127. E. graecum B01 J20 

1129. E. silvestre A. var. silvestre B. var. lineariaefolium Roh Sch 

1130. E. helveticum 

1131. E. odoratum A. var. odoratum Sch (A&S = 1143) B. var. carniolicum Sch 

1132. E. pectinatum 

1135. E. korabense Jav 

1137. E. hieracifolium Sch 

1140. E. repandum A&S K48 

1143. Syrenia cuspidate 

1144. Barbarea vulgaris Jav 

1151. B. balcana Jav 

1153. B. bracteosa 

1154. B. longirostris A&S 

1159. Roripa silvestris 

1160. R. prolifera Jav 

1161. R. pyrenaica I. subsp. pyrenaica H24 Sch II. subsp. lippizensis 

16

1164. Nasturtium officinale 

1167. Cardamine acris 

1168. C. pratensis I. subsp. pratensis Penn II. subsp. matthiola M32 

1169. C. glauca 

1170. C. carnosa 

1172. C. graeca 

1175. C. impatiens Sch 

1177. C. flexuosa Jav 

1178. C. hirsute 

1183. C. enneaphyllos 

1187. C. bulbifera 

* C. plumieri M27 

1192. Turritis glabra A&S Sch 

1193. T. pseudoturritis H24 

1196. Arabis turrita 

1198. A. alpina I. subsp. alpina Hep Sch II. subsp. flavescens III. subsp. caucasica 

1200. A. verna 

1202. A. auriculata A&S 

1204. A. hirsuta var. Ihirsuta var. sagittata A&S var. planisiliqua Sch 

1205. A. constricta H24 Sch 

1206. A. corymbiflora Sch 

1207. A. nivalis H24 Jav 

1208. A. muralis 

1211. A. scopoliana Jav 

1213. A. bryoides Jav M32 

* A. tomorensis M32 (= Arabidopsis thaliana fide Markgraf) 

1214. Aubrieta gracilis 

1215. A. croatica B96 

1216. A. intermedia 

1220. Alliaria officinalis Jav Sch 

1221. Myagrum perfoliatum Sch 

1225. Isatis tinctoria A&S Roh 

1229. Bunias erucago 

1232. Hesperis dinarica H24 

1233. H. runcinata Rod (H. silvestris) 

1237. H. matronalis III. subsp. cladotricha 

1239. H. glutinosa A. var. visianii A&S 

1240. Malcolmia angulifolia M27 

1242. M. maritima A. var. maritima M32 B. var. serbica 

( M. flexuosa H24, M. bicolor B94) 

1246 B. M. bicolor var. veluchensis A&S M27 

1248. M. confusa BB 

1251. Matthiola sinuata B01 F. 

1253 C. M. tristis var. thessala H17 Sch 

1260. Lunaria rediviva M27 

1261. L. annua 

17

1262. L. telekiana Jav 

1264. Fibigia clypeata 

1267. Berteroa incana A. var. incana Hep Sch B. var. trichocarpa Jav 

1270. B. obliqua 

1271. B. samolifolia Bm 

1272. B. mutabilis A. var. mutabilis B. var. viridis H24 

Alyssum excluded 

1323. Ptilotrichum rupestre A. var. rupestre B. var. scardicum H24 Jav 

1324. P. baldaccii 

1327. Alyssoides utriculatum K48 Sch 

1330. Clypeola jonthlaspi 

1331. C. microcarpa A&S 

1332. Draba aizoides A. var. aizoides Jav B. var. athoa 

1333 C. D. lasiocarpa var. elongata 

1335. D. scardica 

1336. D. longirostris 

1337. D. lacaitae var. albanica Jav 

1338. D. parnassica B94 Sch 

1341. D. korabensis 

1342. D. muralis 

1345. Erophila praecox BB 

1347. E. majiuscula H24 

* E. boerhaavii var. vindobensis M32 

* E. verna sensu lato 

1353. Diplotaxis tenuifolia 

1354. D. muralis BB 

1369. Sinapis arvensis Sch 

1370. S. alba J20 

1373. Erucastrum nasturtiifolium Penn 

1374. Hirschfeldia incana A&S 

1376. Cakile maritima Penn Sch 

1378. Rapistrum rugosum A. var. rugosum Sch B. var. orientale Pamp 

1384. Calepina irregularis 

1387. Raphanus landra A&S 

1388. R. raphanistrum 

1389. R. sativus BB 

1391. Conringia orientalis A&S 

1396. Lepidium latifolium A&S 

1397. L. graminifolium 

1399. L. virginicum Sch 

1402. L. campestre 

1405. Cardaria draba A&S 

1406. Coronopus procumbens J20 

1409. Biscutella didyma A&S BB 

18

1412. Hutchinsia brevicaulis H24 Jav 

1414. Iberis sempervirens 

1416. I. carnosa B94 

1418. I. umbellata 

1425. Aethionema saxatile A. subsp. saxatile B. subsp. graeca 

1427. Kernera saxatilis 

1428. Thlaspi bellidifolium 

1429. T. microphyllum 

1433. T. rivale A&S 

1435. T. praecox 

1436. T. cuneifolium Jav M32 

1437. T. ochroleucum M32 

1439. T. goesingense M27 M32 

1440. T. avalanum H24 

1441. T. kovacsii A&S 

1444. T. perfoliatum 

1449. Peltaria alliacaea 

1452. Hornungia petraea 

1454. Capsella grandiflora A&S Penn 

1455. C. bursa-pastoris 

1456. C. rubella H24 

1458. Camelina rumelica A&S 

1462. Vogelia apiculata 

Resedaceae 

Reseda phyteuma Jav 

R. luteola Jav 

Crassulaceae 

1855. Sempervivum schlehani Penn Sch (var. blanda) 

(Hayek record an error fide Markgraf) 

1862. S. heuffelii A. var. heuffelii A&S Hep Sch B. var. glabrum Jav M32 * var. dejae 

M32 * var. albanicum K48 

1863. S. hirtum ?Jav (as S. soboliferum) 

1865. Sedum maximum Jav 

1867. S. magellense 

1868. S. cepaea 

1869. S. stellatum A&S 

1872. S. dasyphyllum 

1873. S. album (subsp. I & III; subsp. II Sch) 

1874. S. serpentini 

1875. S. erythraeum Jav Sch 

1876. S. sartorianum 

1877. S. acre 

19

1879. S. stribrnyi A&S 

1880. S. boloniense 

1881. S. horakii Sch 

1884. S. ochroleucum 

1885. S. rupestre M27 

1888. S. grisebachii Sch 

1889. S. annuum Penn Sch 

1891. S. flexuosum Jav (border) Sch 

1892. S. atratum 

1895. S. glaucum A. var. glaucum C. var. pallidum J20 

1896. S. rubens 

1901. Cotyledon umbilicus-veneris 

1902. C. horizontalis A&S 

1904. C. pendulinus 

Saxifragaceae 

1911. Saxifraga taygetea 

1912. S. rotundifolia 

1913. S. heucherifolia H24 (boundary) Sch 

1914. S. chrysopleniifolia A&S 

1915. S. bulbifera 

1916. S. graeca A&S M27 

1920. S. tridactylites 

1921. S. adscendens II. subsp. adscendens III. subsp. blavii B96 B01 

1926. S. moschata 

1927 I. S. sedoides subsp. prenja Sch 

1929. S. androsacaea Jav Sch 

1930. S. glabella 

1933. S. aizoides Hep Sch 

1934 I. S. media subsp. porophylla (incl. montenegrina, grisebachii) 

1935. S. sempervivum 

1939. S. scardica Sch 

1940. S. marginata (incl. vars. A, B) 

1941. S. spruneri Sch 

1946. S. aizoon (incl. sturmiana, malyi, orientalis) 

1947 B. S. oppositifolia var. meridionalis 

1950. Parnassia palustris 

20

Part 3: flora Europaea vol. 2 

Euphorbiaceae 

269. Chrozophora tinctoria 

271. Mercurialis perennis Jav. Sch 

272. M. ovata 

273. M. annua 

276. Euphorbia palustris J20 Sch 

277. E. lingulata 

278. E. polychroma A&S Penn 

280. E. fragifera H24 J20 

282. E. glabriflora 

283. E. verrucosa 

284. E. spinosa Sch 

287. E. pubescens 

292. E. capitulata 

295. E. stricta 

296. E. platyphyllos (vars. A, B) 

299. E. helioscopia 

301. E. villosa 

305. E. niciciana 

306. E. sequieiana 

309. E. veneta A&S J20 

310. E. heldreichii A&S 

312. E. amygdaloides (incl. D) 

319. E. cyparissias 

321. E. agraria Jav M27(incl. C) 

322. E. terracina A&S 

328. E. paralias 

330. E. herniariaefolia (incl. B) 

332. II. E. pinea BB 

333. E. graeca 

337. E. falcata 

338. E. exigua 

341. E. myrsinites 

344. E. peplis 

346. E. chanaesyce A&S J20 

Buxaceae 

349. Buxus sempervirens 

21

Thelygonaceae 

426. Thelygon cynocrambe 

Cistaceae 

1473. I. Cistus incanus 

1473. II. C. creticus 

1475. C. salvifolius 

1475 X 1476. M27 

1476. C. monspeliensis B01 Sch 

* C. atchleyi Hepb 

1478. Tuberaria guttata 

1482. Helianthemum salicifolium 

1487. H. apenninum Rod 

1488. H. chamaecistus (several vars.) 

1489. H. ovatum I. subsp. ovatum II. subsp. grandiflorum Sch 

1490. H. nitidum 

1492. H. canum (several vars.) 

*H. jonium var. psilosepalum A&S 

1496. Fumana thymifolia 

1498. F. vulgaris 

1500. F. bonapartei 

Violaceae 

1502. Viola odorata 

1506. V. alba subsp. scotophylla M27 

1509. V. hirta M32 

1512. V. pyrenaica A&S 

1519. V. riviniana 

1520. V. silvestris 

1530. V. kosanini 

1532. V. grisebachiana Jav Gui 

1533. V. albanica 

1536 add. V. dukadjinica M27 ?Gui 

1540. V. zoysii 

1544. V. athois Jav Sch 

1545. V. gracilis 

1546. V. heterophylla I. subsp. graeca R III. subsp. epirotica H24 Gui 

1551. V. hymettia M32 Sch 

1554. V. orphanidis 

1556. V. speciosa H17 H24 

1559. V. saxatilis I subsp. aetolica II subsp. macedonica 

1560. V. arvensis B94? J20? (probably 1561) 

1561. V. kitaibeliana 

22

Tamaricaceae 

1571. Tamarix pallasii F M32 

1574. T. hampeana M27 

1575. T. parviflora 

1578. T. africana 

* Myricaria germanica Sch 

Cucurbitaceae 

1582. Bryonia alba A&S 

1583. B. dioica F M27 

1586. Ecballium elaterium 

Guttiferae 

1605. Hypericum perforatum I. subsp. vulgare III. subsp. veronense Jav Sch IV. subsp. 

angustifolium 

1607. I. H. acutum A&S J20 

1611. H. hirsutum A&S 

1614. H. haplophylloides 

1617. H. apollinis 

1622. H. spruneri Form 

1628. H. rumelicum (incl. Var. B) 

1631. H. perfoliatum 

1634. H. grisebachii M32 

1637. H. alpinum 

1640. H. barbatum 

Malvaceae 

1643. Hibiscus trionum A&S Rohl 

H. manihot B96 (cult.) 

Gossypium mexicanum B96 (cult.) 

1644. Abutilon avicennae J20 

1646. Malva moschata 

1650. M. silvestris 

1651. M. ambigua Rohl 

1653. M. neglecta A&S Rod 

1657. Lavatera arborea H24 

1661. L. thuringiaca 

1662. L. ambigua Sch 

1665. Althaea hirsuta 

1670. A. cannabina 

1673. A. officinalis 

1674. A. pallida 

23

1675. A. rosea 

1678. Malope malacoides M32 

Tiliaceae 

1681. Tilia tomentosa 

1682. I. T. platyphyllos Gr M32 

1682. IV. T. corinthia A&S Hep 

Linaceae 

1684. Linum gallicum 

1685. L. liburnicum 

1686. L. spicatum B. strictum 

1687. L. nodiflorum 

1694. L. capitatum 

1696. L. elegans A&S J20 

1697. L. tauricum C. var. serbicum M27 Sch D. var. albanicum 

1701. L. pubescens 

1703. L. hirsutum I. hirsutum H24(boundary) Sch III. spathulatum B94 Jav 

IV byzantinum Sch 

1706. L. tenuifolium 

1707. L. hologynum 

1709. L. angustifolium 

1710. L. usitatissimum 

1712. II. L. extraaxillare A&S Sch 

1713. L. montanum Jav M32 

1715. I. L. alpinum subsp. alpinum Sch 

1716. IB. L. austriacum var. tommasinii B99 

1717. L. catharticum 

1718. Radiola linoides B01 

Oxalidaceae 

1719. Oxalis acetosella 

1721. O. corniculata A&S Sch 

Geraniaceae 

1722. Geranium phaeum K48 

1723. G. reflexum 

1724. G. aristatum 

1725. G. villosum 

1726. G. molle 

1728. G. divaricatum Sch 

1729. G. pyrenaicum 

24

1730. G. bohemicum H24 (Koritnik) Jav 

1731. G. lanuginosum Rod 

1732. G. pusillum Sch 

1733. G. dissectum 

1734. G. striatum 

1736. G. sanguineum 

1737. I. G. subcaulescens 

1738. G. peloponnesiacum Rod Sch 

1739. G. silvaticum 

1741. G. asphodeloides I. nemorosum M27 M32 II. tauricum A&S M32 

1742. G. rotundifolium 

1743. G. coeruleatum Jav Sch 

1746. G. columbinum 

1747. I. G. tuberosum Gui M27 Rod 

1748. G. macrorhizum I. macrorhizum II. microrhizum Sch 

1749. G. lucidum 

1750. G. purpureum 

1751. G. robertianum 

1757. Erodium laciniatum Rod 

1759. E. malacoides A&S BB 

1760. E. moschatum 

1761. E. cicutarium 

1762. E. ciconium A&S 

1764. E. guicciardii Rod 

Zygophyllaceae 

1773. Tribulus terrestris 

Rutaceae 

1775. Ruta graveolens Rod Sch 

1775. B. R. divaricata H17 J20 

1777. R. chalepensis Penn 

1777. B. R. bracteosa B94b 

1779. Haplophyllum coronatum 

1780. H. patavinum B01 Sch 

1780. B. H. albanicum M27 

1785. H. boissieranum Sch 

1786. Dictamnus albus 

Simaroubaceae 

1787. Ailanthus altissimus [J20 Sch] 

25

Polygalaceae 

1790. Polygala supina var. supina H24 Sch B. var. bosniaca Jav 

1794. B. P. maior var. pindica M27 M32 

1796. P. doerfleri 

1797. I. P. nicaeensis subsp. mediterranea 

1798. P. comosa M27 Penn 

1799. P. vulgaris I. subsp. vulgaris II. subsp. temskyanus 

1800. P. carniolica M27 

1801. II. P. alpestris subsp. croatica 

1804. P. monspeliaca A&S 

Anacardiaceae 

1806. Pistacia terebinthum 

1807. P. lentiscus 

1808. Cotinus coggygria 

1809. Rhus coriaria M27 Sch 

Aceraceae 

1812. Acer tataricum 

1813. A. pseudoplatanus 

1815. A. obtusatum 

1816. A. hyrcanum 

1818. A. monspessulanum 

1821. A. campestre 

1818 X 1821. H17 

1822. A. platanoides H24 (Koritnik) M32 

Hippocastanaceae 

1823. Aesculus hippocastanum 

Aquifoliaceae 

1824. Ilex aquifolium B94 M32 

Celastraceae 

1825. Euonymus verrucosus 

1826. E. europaeus 

Rhamnaceae 

1831. Rhamnus cathartica 

26

1832. R. orbiculata H24 

1833. R. infectoria B. var. pubescens B94 B96 (probably errors for 1832) 

1834. R. tinctoria M27 

1836. R. prunifolia J20 

1840. R. fallax 

1841. R. pumila H24 M32 

1844. R. rupestris 

1845. R. frangula Jav J20 

1846. R. alaternus BB 

1847. Zizyphus jujuba Gr (boundary) 

1849. Paliurus spina-christi 

Vitaceae 

1850. Vitis silvestris 

Rosaceae 

1965. Aruncus silvester Jav 

1966. Filipendula hexapetala 

1967. F. ulmaria A&S (subsp. dentata) 

1968. Rubus saxatilis Jav 

1969. R. idaeus K48 

1972 I. R. candicans subsp. thyrsanthus M27 

1980. R. ulmifolius 

I. subsp. rusticanus II. subsp. dalmaticus M27 Sch IV. Subsp. thessalus Jav 

1980 x 1983 B B96 

1980 III x 1983 Jav 

1983 B. R. tomentosus var. glabratus H24 

1990. R. ochrosetus Sch 

1991. R. serpens M32 

1993. R. hirtus A&S 

1993 add. R. begoniaefolius Jav 

1998. R. nemorosus J20 M27 

1999. R. caesius 

* R. tomentifolius Rohl 

2000. Fragaria vesca 

2001. F. moschata 

2002. F. viridis M27 M32 

2003. Potentilla speciosa 

2005. P. apennina 

2008. P. clusiana Hep Sch 

2010. P. sterilis M32 

2012. P. micrantha 

2014. P. rupestris M32 

2015. P. argentea 

27

2016. P. canescens 

2017. P. recta 

2020. P. hirta 

2021. P. adriatica J20 

2022. P. detommasii var. holosericea A&S B99 

2024. P. pindicola A&S 

2026. P. patula Sch 

2028. P. opaca H17 Sch 

2029. P. australis Sch 

2031. P. crantzii 

2032. P. aurea B01 Sch 

2033. P. ternata 

2038. P. tommasiniana H24(Koritnik) 

2039. P. reptans 

2040. P. erecta Sch 

2041. P. erecta H17 H24(Pastrik) 

2047. Alchenilla arvensis 

2050. A. hoppeana 

2052. A. glaberrima Sch 

2053. A. flabellata H17 

2054. A. acutiloba Jav 

2055 D. A. alpestris var. frigens 

2056. A. pratensis 

2057. A. silvestris A.var. pastoralis H17 B. var. subcrenata M32 D. var. acutangula Sch 

2058. A. pubescens (several vars.) 

* A. vulgaris subsp. coriacea Sch 

2061. Geum coccineum 

2062. G. urbanum 

2063. G. aleppicum Sch 

2064. G. molle 

2065. G. montanum 

2067. G. reptans H24 Jav 

2068. G. bulgaricum 

2069. G. rivale Sch 

* G. heterocarpum A&S 

2070. Dryas octopetala 

2071. Agrimonia eupatoria 

2073. A. agrimonioides 

2074. Sanguisorbia officinalis Jav 

2075. S. albanica Jav 

2076. S. minor 

2078. S. muricata 

2086. Rosa sempervirens 

2088. R. arvensis 

2089. R. gallica 

2091. R. andegavensis A&S H24 

28

2092. R. canina B96 I. subsp. lutetiana Jav II. subsp. spuria H17 IV. subsp. dumalis 

B96 M32 

2093. R. dumetorum A&S Jav 

2095. R. glauca Jav. 

2098. R. heckeliana 

2100. R. tomentella 

2101. R. agrestis 

2102. R. micrantha B94 Sch 

2110. R. sicula B94 B96 

2111. R. glutinosa 

2112. R. tomentosa Jav M27 

2113. R. mollis 

2114. R. pomifera Jav 

2116. R. orientalis Sch 

2117. R. rubrifoliia 

2120. R. pendulina 

2121. R. spinosissima 

* R. elliptica M27 

2123. Cydonia oblonga M27 

2124. Amelanchier ovalis 

2127. Sorbus domestica 

2128. S. aucuparia M27 K48 

2129 I. S. austriaca Jav 

2130 II. S. aria subsp. umbellate 

2131. S. chaemaemespilus Jav Sch 

2132. S. torminalis 

2134. Pyrus amygdaliformis 

2138. P. piraster 

2139. Malus silvestris 

2140. M. pumila M32 

2141. Cotoneaster integerrima 

2143. C. tomentosa Jav 

2145. Pyracantha coccinea 

2148. Crataegus pentagyna H24 

2149. C. orientalis J20 B99 

2150. C. heldreichii A&S 

2154. C. monogyna 

2156. Prunus spinosa 

2157. P. pseudarmeniaca A&S 

2158. P. cerasifera subsp. divaricata M27 

2159. P. insititia Jav B94b 

2165. P. prostrata 

2169. P. mahaleb 

29

Leguminosae 

2173. Cercis siliquastrum M32 Rohl 

2178. Colutea arborescens 

2182. Astragalus hamosus A&S J20 

2192. A. gremlii 

2193. A. glycyphyllus 

2205. A. autrini B94 M32 

2207. A. exscapus A&S Gui 

2213. A. depressus 

2216 II. A. creticus subsp. rumelicus 

2224. A. vesicarius A&S. R 

2230. A. sericophyllus R 

2237. A. onobrychis A. var. onobrychis D. var. chlorocarpus 

2241. A. monspessulanus A. var. atticus A&S B. var. macedonicus M32 Penn B96? BB? 

2243. A. spruneri Sch 

2245. A. fialae H17 Sch 

2249. A. baldaccii A&S B? 

2251. A. angustifolius 

* A. sirinicus A&S 

2252 B. Oxytropis pilosa var. purpureus B01? 

2254 III. O. campestris subsp. dinarica 

2255 b. O. sericea subvar. korabensis H24 Jav 

2256. O. prenja Sch 

2257 I. O. montana subsp. jacquini H24 

2258. O. lapponica Jav (Korab boundary) 

* O. purpurea M27 R 

2259. Biserrula pelecinus A&S 

2260. Glycyrrhiza glabra B94b 

2264. Galega officinalis 

2265. Psoralea bituminosa 

2267. Cicer montbretii A&S 

2272. Vicia narbonensis I. subsp. narbonensis II. subsp. serratifolia 

2273. V. lathyroides Penn 

2274. V. lutea 

2275. V. melanops 

2277. V. peregrina A&S Gr 

2279. V. grandiflora 

2280. V. sepium M27 Sch 

2282. V. sativa II. subsp.cordata III. subsp. angustifolia H17 Penn 

2284. V. hybrida Gr Sch 

2285. V. pannonica II. subsp. striata 

2287. V. bithynica 

2289. V. onobrychioides 

2293. V. silvatica H17 H24 

2300. V. tenuifolia 

30

2301. V. cracca I. subsp. cracca II. subsp. incana 

2302 1 st . V. villosa 

2302 2 nd . V. dasycarpa 

2312. V. hirsuta J20 

2314. V. ochroleuca Sch 

2316. V. tetrasperma 

2318. V. ervilia A&S Rod 

2321. Ervum ervoides 

2322. Lathyrus aphaca 

2323. L. nissolia 

2324. L. clymenum J20 

2326. L. ochrus Gr 

2327. L. cicera Rod 

2330. L. sativus A&S 

2332. L. annuus J20 

2334. L. sphaericus 

2335. L. inconspicuous 

2337. L. setifolius 

2338. L. hirsutus 

2339. L. pratensis 

2341. L. tuberosus A&S 

2342. L. grandiflorus 

2344. L. silvester M32 

2346. L. megalanthus A. var megalanthus B. var. lanceolatus B96 

2351. L. inermis 

2354. L. pannonicus Rod Penn 

2356 B. L. filiformis var. ensifolius 

2357. L. montanus H24 

2359. L. friedrichsthalii 

2360. L. vernus M32 

2361. L. venetus 

2362. L. niger 

2364. Pisum elatius Penn 

2365. P. sativum Penn 

2369. Ononis breviflora A&S B94 

2372. O. ornithopodioides A&S 

2373. O. reclinata var. mollis M27 

2374. O. variegata B96 

2375. O. pusilla B94 

2383. O. antiquorum J20 Sch 

2386. O. spinosa 

2387. O. hircine 

2394. Trigonella monspeliaca 

2398. T. corniculata 

2400. T. hamosa H24 

2405. Medicago marina 

31

2406. M. sativa J20 

2407. M. falcata 

2409. M. lupulina 

2410. M. prostrata 

2412. M. orbicularis 

2415. M. carstiensis 

2416. M. hispida 

2420. M. coronata A&S 

2421. M. rigidula 

2425. M. litoralis M27 

2426. M. turbinata Rod 

2428. M. arabica 

2429. M. minima 

2435. Melilotus italicus A&S 

2437. M. altissimus b. subvar. palustris B96 J20 c. subvar. adriaticus H24 

2438. M. neapolitanus 

2439. M. officinalis 

2440 1 st . M. elegans B94 

2440 2 nd . M. alba 

2441. M. indica 

2442. Trifolium filiforme B01 

2445. T. campestre 

2450. T. patens 

2451. T. velenovskyi 

2452. T. strepens Sch 

2454. T. badium 

2455. T. speciosum A&S Held 

2464. T. nigrescens 

2468. T. pallescens 

2469. T. repens 

2470. T. biasolettii J20 

2471. T. glomeratum J20 

2472. T. suffocatum 

2474 b. T. nervulosum var. albanicum B01 

2477. T. pilczii 

2479. T. vesiculosum 

2480. T. mutabile Sch 

2481. T. physodes 

2482. T. fragiferum 

2483. T. resupinatum 

2484. T. tomentosum 

2486. T. tenuifolium 

2487. T. trichopterum Sch 

2488. T. striatum 

2489. T. scabrum 

2490. T. dalmaticum 

32

2492. T. pallidum 

2495. T. cinctum 

2503. T. arvense 

2507. T. stellatum A. var. stellatum C. var. xanthinum A&S 

2508 A. T. incarnatum var. molineri 

2511. T. angustifolium 

2512. T. purpureum 

2514. T. noricum 

2515. T. praetutianum 

2517. T. patulum 

2519. T. pratense 

2521. T.wettsteinii 

2522. T. ochroleucum 

2523. T. pannonicum Sch 

2524. T. pignatii 

2525. T. medium 

2526. T. hirtum B01 Jav 

2529. T. lappaceum 

2530. T. alpestre 

2532. T. subterraneum 

2535. Dorycnium hirsutum 

2538. D. germanicum 

2539. D. herbaceum 

2540. Lotus ornithopodiodes A&S H24 

2549. L. corniculatus I. subsp. corniculatus III. subsp. tenuis. 

2550 L. preslii A&S 

2553. L. angustissimus B01 Rohl 

2556. L. edulis 

2562. Securigera securidaca 

2563. Hymenocarpus circinnatus 

2564. Anthyllis hermanninae B94b 

2565. A. barba-jovis Rod 

2567. A. aurea 

2568 I. A. montana subsp. jacquinii 

2569. A. vulneraria subsp. I, II, III, IV, V 

2570. Physanthyllis tetraphylla A&S 

2577. Lupinus graecus A&S Held 

2579. Argyrolobium linnaeanum B94b 

2583. Cytisus nigricans 

2584. C. triflorus Rod 

2586. C. diffusus A. var. diffusus Penn Gui B. var. pseudoprocumbens M27 K48 

2587. C. decumbens A. var. decumbens M32 C. var. rectipilosus H24 

2593. C. hirsutus subsp. hirsutus I. subsp. leucotrichus H24 

2599. C. tommasinii 

2601 C. C. austriacum var. pindicolum 

2603. C. heuffelli H24 

33

2605. C. jankae H24 Sch 

2610. Podocytisus caramanicus B99 

2612. Laburnum alpinum M27 M32 

2613. Calycotome infesta B96b 

2614. C. villosa A&S 

2616. Petteria ramentacea B01 H17 

2617. Genista radiata 

2619. G. hassertiana 

2628. G. silvestris I. subsp. silvestris H24 Jav II. subsp. dalmatica H24 

2630. G. pilosa B01 

2635 B. G. sericea var. rigida B94 

2637. G. tinctoria 

2641. G. depressa III. subsp. depressa Sch IV. subsp. csikii 

2642. G. januensis M32 

2648. G. sagittalis 

2647. Spartium junceum 

2648. Ornithopus compressus 

2650 I. Coronilla emerus subsp. emeroides 

2652. C. valentina BB 

2653. C. glauca B94 

2658. C. varia 

2659. C. elegans H24 

2660. C. cretica 

2662. C. scorpioides 

2664. Hippocrepis comosa I. subsp. comosa II. subsp. glauca 

2668. H. unisiliquosa H24 Penn 

2670. Scorpiurus muricatus M27 

2671. S. subvillosus A&S J20 

2686 Onobrychis alba I. subsp. alba subsp. laconica A&S H17 

2687. O. arenaria I. subsp. lasiostachya R III. subsp. arenaria M27 Sch 

2688. O. viciaefolia M32 Jav 

2689. O. montana a. var. montana c. var. scardica 

2691. O. ocellata B01 (check) 

2692. O. oxyodonta 

2694. O. aequidentata subsp. foveolata A&S Penn 

2695. O. caput-galli 

Thymelaeaceae 

2697. Lygia passerina 

2702. Daphne mezereum 

2705. D. blagayana 

2707 II. D. oleoides 

2712. D. cneorum H24 Jav 

2713. D. gnidium B94b 

34

Lythraceae 

2716 (1 st ). Lythrum salicaria 

2716 (2678). L. virgatum Sch 

2716 (2 nd ). L. flexuosum B94 J20 

2718. L. hyssopifolia 

2720. L. tribracteatum B94 Roh 

2722. Peplis portula Jav 

Myrtaceae 

2725. Myrtus communis 

Punicaceae 

2726. Punica granatum 

Onagraceae 

2727. Ludwigia palustris A&S J20 Sch 

2728. Epilobium hirsutum 

2729. E. parviflorum 

2730. E. montanum 

2731. E. collinum Sch 

2732. E. lanceolatum A&S Sch 

2733. E. roseum 

2734. E. gemmascens M32 

2735. E. adnatum B01 

2737. E. obscurum Sch 

2738. E. lamyi J20 

2739. E. palustre Sch 

2740. E. alpestre Sch 

2742. E. anagallidifolium H24 Jav 

2743. Chamaenerium angustifolium A&S B01 

2744. C. palustre 

2747. Circaea lutetiana Sch 

2750. Trapa natans Gr 

Halorrhagidaceae 

2751. Myriophyllum verticillatum J20 Sch 

2752. M. spicatum J20 

Hippuridaceae 

2753. Hippuris vulgaris J20 Sch 

35

Callitrichaceae 

2755. Callitriche verna J20 Sch 

2759. C. stagnalis 

Cornaceae 

2762. Cornus mas 

2763. C. sanguinea 

2764. C. australis Sch 

Araliaceae 

2765. Hedera helix 

Umbelliferae 

2766. Hydrocotyle vulgare A&S 

2767. Sanicula europaea 

2770 I. Astrantia maior subsp. elatior 

2772. Eryngium maritimum 

2775. E. palmatum 

2777. E. creticum 

2779. E. amethystinum 

2780. E. campestre 

2786. Bupleurum falcatum Jav Hep 

2789. B. sibthorpianum 

2790. B. junceum 

2791 I. B. commutatum subsp. eucommutatum H24 

2796. B. tenuissimum var. tenuissimum A&S var. columnae A&S 

2799. B. karglii 

2801. B. semidiaphanum A&S. Form 

2804. B. veronense (B. aristatum) 

2805. B. flavicans 

2807. B. longifolium M32 

2808. B. lancifolium Gr J20 

2809. B. rotundifolium A&S H24 

2810. Trinia glauca 

2814. T. dalechampii A&S 

2817. Apium graveolens Form 

2819. A. nodiflorum A&S B94 

2825. Ammi majus A&S Sch 

2826. A. visnaga 

2827. Ptychotis ammoides B94b 

2829. Carum carvii Sch 

2831. C. lumpeanum H24 Jav 

36

2832. C. multiflorum A&S 

2834. C. graecum 

2836. C. rupestre 

2838. Bunium montanum 

2839. B. alpinum 

2846 Freyera cynapioides A. var. cynapioides C. var. balcanica M32 Sch 

2847. F. parnassica A&S B99 

2854. Pimpinella alpestris M32 Sch 

2855. P. saxifraga J20 

2856. P. polyclada A&S B99 

2857. P. tragium 

2859. P. peregrina 

2862. Pancicia serbica K48 Sch 

2865. Sium latifolium Hep 

2867. Berula erecta 

2868. Crithmum maritimum 

2874. Seseli rigidum Sch 

2883. S. tommasinii 

2895. S. peucedanoides 

2898. Oenanthe aquatica J20 Sch 

2899. O. tenuifolia A&S 

2902. O. silaifolia A. var. silaifolia C. var. media Jav 

2907. O. pimpinelloides 

2908. O. angulosa Sch 

2909. O. incrassans 

2911. O. fistulosa 

2914. Portenschlagia ramosissima J20 

2916. Athamanta macedonica 

2917. A. densa B96 

2918. A. haynaldi 

* A. albanica A&S 

2921. Foeniculum vulgare 

2922. F. piperitum B96. Form 

2924. Silaum flavescens Hep 

2925. Meum athamanticum 

2926. Cnidium silaifolium A. var. silaifolium B. var. orientale 

2930. Ligusticum seguierii Sch 

2931. L. mutellina Jav Sch 

2932. L. albanicum Jav Sch 

2934. Angelica silvestris 

2935. A. elata A&S 

2941. Levisticum officinale A&S 

2944. Johrenia pichleri B96 

2949 B. Ferula candelabrum A&S 

2951. Ferulago silvatica A. var. silvatica A&S M32 C. var. monticola H24 

2954. F. nodosa 

37

2955. F. galbanifera Jav 

2958. Opopanax chironium Sch 

2959. O. hispidum Pamp 

2962. Peucedanum officinale Sch 

2963. P. longifolium A&S 

2968. P. oligophyllum 

2969 B. P. schottii var. petraeum A&S 

2970. P. vittijugum A&S 

2973. P. neumayeri A. var. neumayeri J20 B. var. serpentini Jav Sch 

2976. P. aegopodioides A&S M32 

2977. P. latifolium Sch 

2978. P. alsaticum J20 K48 

2980. P. cervaria A. var. cervaria M32 C. var. latifolium J20 

2983. P. austriacum 

2985. Pastinaca sativa I. subsp. sativa A&S J20 II. subsp. urens H24 Jav 

2989. Heracleum orsinii H24 Jav 

2990. H. pallinianum var. oetum 

2991. H. sphondylium Sch 

2992. H. sibiricum A&S Sch 

2996. Malabaila aurea 

2997. M. involucrata A&S 

2999. Tordylium maximum 

3000. T. officinale A&S J20 

3001. T. apulum 

3003. Laser trilobum Jav 

3004. Elaeosilinum asclepium B96 Pamp 

3005 II. Laserpitium siler subsp. garganicum A&S 

3005 II x 3006. A&S 

3006. L. zernyi H24 Jav 

3009. L. marginatum M32 

3016. Daucus pumilus M27 

3019. D. broteri Sch 

3020. D. guttatum M32 Sch 

3021. D. carota 

3023. Orlaya daucorlaya 

3024. O. grandiflora 

3025. O. platycarpa M27 

3026. Turgenia latifolia A&S Sch 

3027. Caucalis daucoides A&S 

3030. Torilis nodosa 

3031. T. leptophylla A&S 

3034. T. arvensis I. subsp. arvensis II. subsp. purpurea 

3035. T. anthriscus A&S Sch 

3038. Physocaulis nodosa A&S Sch 

3040. Chaerophyllum aromaticum A&S B01 

3043. C. aureum 

38

3044. C. temulum A&S 

3047. C. villarsii Sch 

3048. C. cicutaria Sch 

3049. C. coloratum 

3052. Anthriscus silvester A&S Penn 

3053. A. nemorosus A&S 

3054. A. fumarioides Sch 

3056 B. A. cerefolium var. trichosperma A&S Jav 

3058. A. vulgaris H24 

3060. Scxandix pecten-veneris 

3063. S. grandiflora A&S 

3064. Myrrhis odorata Sch 

3068. Danaa cornubiensis 

3069. Scaligera cretica A&S B94 

3070. Conium maculatum A&S B01 

3071. C. divaricatum A&S 

3072. Smyrnium olusatrum A&S 

3075. S. perfoliatum 

3081. Prangos ferulacea 

3085. Echinophora spinosa B96 F 

3088. Bifora radians A&S H24 

3089. B. testiculata A&S 

39

Part 4: Flora Europaea vols. 3 & 4 

Plumbaginaceae 

3090. Plumbago europaea 

3091. Acantholimon echinus 

3093. Goniolinum dalmaticum A&S 

3097. Statice sinuata A&S 

3099. S. angustifolia 

3108. S. oleifolia 

3111. S. anfracta A&S 

3118. Armeria canescens 

3121. A. rumelica 

3124. A. alpina Sch 

Pyrolaceae 

3127. Ramischia secunda 

3128. Pyrola minor 

3133. Monotropa hypopitys 

Ericaceae 

3139. Arbutus unedo 

3140. A. andrachne B99 

3141. Arctostaphylos uva-ursi 

3142. A. alpina B17 (field record) 

3144. Erica arborea 

3145. E. carnea 

3146. E. verticillata 

3149. Vaccinium myrtillus 

3150. V. uliginosum Jav (border) Sch 

3152. V. vitis-idaea Sch 

Empetraceae 

3153. Empetrum nigrum Jav (field record) 

Primulaceae 

3154 I. Primula vulgaris subsp. vulgaris 

3155 II. P. elatior subsp. intricate 

3156. P. veris I. subsp. veris Penn M32 II. subsp. columnae 

3159. P. longiflora H24 Jav 

3165. Androsace villosa 

3166. A. hedreantha H24 Jav (border) 

40

3172. Soldanella alpina 

3173. S. pindicola M27 M32 

3174. S. dimoniei 

3175. S. hungarica Sch 

3185. Cyclamen neapolitanum 

3187. Lysimachia nummularia 

3190. L. punctata 

3191. L. vulgaris 

3192. L. atropurpurea 

3193. L. dubia 

3194. Asterolinum linum-stellatum BB 

3196. Anagallis arvensis 

3197. A. femina 

3196 x 3197 A. x doerfleri Jav H24 

3199. Centunculus minimus J20 

3200. Samolus valerandi 

3201. Coris monspelianus var. annua B94 

Ebenaceae 

3202. Diospyros lotus Tir 

* D. kaki Tir 

Styracaceae 

3203. Styrax officinalis Tir 

Convolvulaceae 

3209. Convolvulus cneorum Tir 

3212. C. lineatus B94 

3213. C. cantabricus 

3214. C. radicosus A&S Gui (C. nitidus auct.) 

3220. C. arvensis 

3222. C. hirsutus 

3224. C. elegantissimus 

3225. Calystegia sepium 

3226. C. silvestris 

3227. C. soldanella M27 Pamp 

3228. Cressa cretica B96 

3231. Cuscuta breviflora Tir 

3233. C. europaea 

3235. C. epithymum 

* C. campestris A&S 

* C. globularis M32 

Boraginaceae 

41

3242. Heliotropium supinum B96 

3243. H. dolosum A&S 

3244. H. europaeum A. var. europaeum B. var. lineatum M32 

3250. Cynoglossum officinale Sch Tir 

3251.C. columnae 

3253. C. creticum 

3254. C. montanum 

3255. C. germanicum A&S Sch 

3257. Solenanthus scardicus 

3258. S. albanicus R (n.b. the type locality is on the Al/Gr border and cannot be precisely 

localized) 

3259. S. appeninus H24 Bm33 

3264. Lappula echinata 

3265. L. barbata A&S 

3267. Asperugo procumbens A&S H24 

3268. Symphytum officinale Tir 

3272. Symphytum tuberosum 

3273. S. bulbosum Sch Tir 

3275. S. ottomanum 

3282. Anchusa hybrida 

3284. A. procera Sch 

3286. A. italica 

3288. A. stylosa Tir 

3291 A. arvensis Tir. 

3293. A. variegata 

3295. A. barrelierii 

3300. Nonnea pallens A&S 

3302 B. Alkanna pulmonaria var. pindica A&S 

3307. A. scardica H24 

3308. A. graeca 

3310. A. boeotica Sch 

3315. A. tinctoria 

3316. Pulmonaria angustifolia M27 

3317. P. rubra H24 

3318. P. mollissima 

3320. P. officinalis 

3321. Myosotis scorpioides 

3322. M. lingulata A&S 

3325. M. sicula J20 

3329. M. sylvatica I. subsp. sylvatica II. subsp. alpestris 

3330. M. versicolor A&S 

3331. M. arvensis 

3332. M. collina 

3333. M. micrantha 

3336. M. refracta A&S 

42

3337. M. sparsiflora H24 

3338. Moltkea petraea 

3339. M. doerfleri 

3340. Halacsya sendtneri 

3343. Lithospermum purpureo-coeruleum 

3344. L. officinale 

3345. L. arvense Sch Tir 

3346. L. incrassatum 

3349. L. apulum Tir 

3351. Onosma stellulatum B94 B96 BB b. subvar. serpentini Jav M42 

3355. O. tauricum M32 Sch 

3356. O. mattirolii 

3359. O. aucherianum I. subsp. aucherianum A&S M27 II. subsp. javorkae 

3360. O. tubiflorum I. subsp. tubiflorum R II. subsp. viride 

3361. O. arenarium subsp. arenarium A&S Hep I. subsp. pyramidatum R 

III. subsp. albanicum H24 Tir 

3364 I. O. visianii subsp. visianii 

* O. canescens M27 

3370. Cerinthe minor 

3372. C. major 

3373. C. glabra Tir 

3375. C. retorta A&S Penn 

3376. Echium italicum 

3377. E. rubrum 

3378. E. vulgare 

3379. E. diffusum Tir 

3380. E. plantaginium 

3381. E. confusum Pamp 

3383. E. parviflorum Tir 

Solanaceae 

3385. Lycium europaeum B94 

3389. Atropa belladonna 

3391. Hyoscyamus nigrum H17 

3393. Hyoscyamus albus 

3396. Physalis alkekengi 

* P. pubescens Tir 

* Capsicum annuum 

3397. Solanum sodomaeum Tir 

3399. Solanum dulcamara 

3400. S. pseudocapsicum [B 01] 

3402. S. alatum B96 

3403. S. luteum Jav 

3404. S. nigrum 

3405. S. lycopersicum Tir 

43

* S. melongena Tir 

* S. miniatum Tir 

* S. jasminoides Sch 

* S. tuberosum Tir 

3408. Datura stramonium 

* Nicotiana tabacum Tir [B96 cult.] 

Scrophulariaceae 

3415. Verbascum densiflorum a. subvar. densiflorum b. subvar. thapsiforme M32 

3419. V. phlomoides 

3420. V. longifolium 

3423. V. pannosum Jav K48 

3428. V. nicolai Sch 

3430. V. guicciardii M27 

3436. V. baldaccii Sch 

3444. V. undulatum B. var. integrifolium B99 

3455 I. V. speciosum subsp. speciosum A&S M32 

3457. V. pulverulentum 

3457 x 3471 

3462. V. banaticum 

3468. V. lychnitis Sch 

3471. V. sinuatum 

3473. V. mallophorum A&S K48 

3476. V. bosnense 

3479. V. bornmulleri 

3481. V. nigrum A. var. nigrum B. var. abietinum M32 

3482. V. glabratum 

3486. V. blattaria 

3494. V. rohlenae J20 Jav 

3497. V. phoeniceum 

3513. Antirrhinum majus 

3516. A. orontium A&S M27 

3517. Linaria reflexa BB 

3520. L. arvensis Gr 

3521. L. parviflora B99 

3522. L. alpina 

3524. L. pellisseriana 

3525. L. concolor Sch 

3526. L. chalepensis A&S Gr 

3528. L. peloponnesiaca 

3531. L. italica Sch 

3531. L. genistifolia Jav 

3533. L. dalmatica 

3534. L. vulgaris 

3535. L. cymbalaria I. subsp. cymbalaria J20 II. subsp. longipes A&S 

44

3536. L. microcalyx Penn Sch 

3538. Kickxia spuria Tir 

3539. K. elatine I. subsp. elatine II. subsp. sieberi 

3540. K. commutata 

3541. K. cirrhosa B94 

3544. Chaenorrhinum minus I. subsp. viscidum II. subsp. litorale Sch Tir 

Pawlownia tomentosa Sch (status?) 

3546. Scrophularia peregrina 

3547. S. aestivalis Sch 

3548. S. bosniaca 

3549. S. scopolii 

3551. S. nodosa 

3552. S. alata A&S 

3556. S. heterophylla I. subsp. heterophylla II. subsp. laciniata 

3559. S. canina 

3564. Gratiola officinalis 

3573. Veronica spicata I. subsp. spicata J20 Sch II. subsp. orchidacea J20 Sch 

3575. V. andrasovszkyi 

3576. V. satureioides H24 

3581. V. alpina 

3582. V. balcanica Jav Sch 

3583. V. serpyllifolia 

3585. V. peloponnesiaca A&S 

3587. V. acinifolia Tir 

3588. V. arvensis 

3590. V. praecox A&S 

3591. V. triphyllos A&S 

3595. V. tournefortii 

3596. V. polita BB 

3598. V. agrestis H24 

3599. V. cymbalaria A&S 

3600 II. V. hederifolia subsp. triloba J20 

3602. V. prostrata 

3603. V. teucrium I. subsp. pseudochamaedrys Sch II.. subsp. orsiniana R 

3604. V. austriaca (all subsp.) 

3607. V. rhodopaea *var. albanica K48 Tir 

3608. V. chamaedrys 

3609. V. urticifolia Jav Sch 

3611. V. officinalis 

3613. V. montana K48 Sch 

3614. V. aphylla B96 Sch 

3616. V. scutellata Tir 

3617. V. scardica 

3618. V. beccabunga 

3620. V. anagalloides A&S Sch 

3621. V. anagallis 

45

3622. V. aquatica A&S H24 

3623. Wulfenia carinthiaca Sch 

3624. W. baldaccii 

3625. Digitalis ferruginea 

3626. D. laevigata 

3628. D. lanata 

3629. D. leucophaea Sch 

3630. D. ambigua 

3631. D. lutea Tir 

3632. D. viridiflora 

3634. Bellardia trixago 

3635. Parentuchellia viscose 

3636. P. latifolia 

3638. Odontites glutinosa 

3641 II. O. rubra subsp. serotina 

3642 II. Euphrasia rostkoviana subsp. montana H24 Sch 

3644. E. stricta Tir 

3645. E. pectinata I. subsp. pectinata II. subsp. tatarica 

3646. E. liburnica M27 Tir 

3647. E. minima 

3649. E. drosocalyx K48 

3650. E. brevipila K48 

3651 E. hirtella H24 Sch 

3652. E. salisburgensis 

3657. Alectorolophus rumelicus I. subsp. rumelicus II. subsp. sagorskii Sch 

3658. A. pubescens Sch 

3659. A. wagneri Sch 

3661. A. major A&S 

3663 I. A. mediterraneus subsp. mediterraneus 

3664. II. A. melampyroides subsp. hayekii A&S Sch 

3668. A. angustifolius I. subsp. subalpinus Sch II. subsp. angustifolius Jav Sch 

III. subsp. lanceolatus Sch 

3670. A. sintenisii A&S 

3673. A. minor 

* A. behrendsenii M27 

3677. Pedicularis comosa Tir 

3679. P. brachyodonta (all susp.) 

3681 B. P. leucodon var. bosniaca 

3682. P. friderici-augusti Sch 

3683.P. graeca 

3684. P. heterodonta R 

3685. P. petiolaris 

3689. P. hoersmanniana H24 

3691. P. orthantha Sch 

3694. P. verticillata 

* P. fallax M27 

46

3696 III. Melampyrum cristatum subsp. cristatum H24 

3697. M. arvense A&S 

3699. M. fimbriatum 

3700 II. M. barbatum subsp. carstiensis Sch 

3705. M. doefleri H24 

3707. M. heracleoticum 

3712. Latheaea squamata Sch 

Lentibulariaceae 

3715. Pinguicula vulgaris A&S 

3716. P. leptoceras 

3717. P. hirtiflora 

3718. P. louisii 

3719. Utricularia vulgaris M27 

3720. U. neglecta Sch 

3723. U. minor M27 

Orobanchaceae 

3726. Orobanche ramosa Tir 

3727. O. nana 

3729. O. mutelii A&S 

3736. O. novackiana M27 M32 

3739. O. gracilis M32 Sch 

3741. O. major Tir 

3746. O. amethystea M32 

3752. O. versicolor H24 

3754. O. minor 

3756. O. picridis-hieracioides Sch 

3758. O. lutea A&S Hep 

3759. O. teucri H24 

3760. O. caryophyllacea 

3762. O. pancicii M27 

3763. O. alba 

3764 B. O. reticulata var. pallidiflora H24 M32 

Gesneriaceae 

3766. Ramonda serbica 

Acanthaceae 

3772. Acanthus longifolius 

3774. A. spinosissimus 

47

Verbenaceae 

3776. Verbena officinalis 

3779 B. Lippia nodiflora var. repens B94 A&S 

3781. Vitex agnus-castus 

Labiatae 

3782. Ajuga reptans 

3783. A. genevensis 

3784. A. pyramidalis K48 Sch 

3785. A. orientalis 

3786. A. piskoi B99 A&S 

3790. A. chamaepitys 

3791. A. chia A&S Tir 

3795. Teucrium ardoini 

3805. T. scordium B94 (check) 

3807. T. flavum 

3810. T. chamaedrys 

3812. T. montanum 

3817. T. polium 

3819. Rosmarinus officinalis Tir 

3820. Prasium majus A&S M32 

3821 B. Scutellaria orientalis var. pinnatifida Sch 

3822. S. alpina 

3823. S. columnae 

3824. S. peregrina BB A&S 

3825. S. altissima Sch 

3831. S. galericulata J20 Sch 

3838. Marrubium cyllenaeum A&S 

3840. M. incanum J20 Sch 

3841. M. peregrinum 

3843. M. vulgare 

3841 x 3843 Sch 

3845. Sideritis montana 

3847. S. romana F Sch 

3848. S. purpurea 

3850. S. scardica M27 

3852. S. roemeri 

3856. Nepeta cataria 

3858. N. parnassica B94. 

3859. N. spruneri 

3866. N. pannonica 

3869. Glechoma hederacea Tir 

3871. G. hirsute H17 

48

3874. Prunella grandiflora Tir 

3875. P. vulgaris 

3874 x 3875 

3876. P. laciniata 

3877. Melittis melisophyllum 

3881. Phlomis fruticosa 

3887 II. Galeopsis ladanum subsp. latifolia A&S 

3888. G. tetrahit Sch 

3889. G. pubescens Sch 

3891 I. G. speciosa subsp. speciosa Jav 

3892. Lamium galeobdolon I. subsp. montanum II. subsp. vulgare H17 

3896. L. maculatum (all vars.) 

3897. L. garganicum I. subsp. garganicum Jav II. subsp. glabratum 

III B. subsp. striatum var. minus B94 B96 

3898. L. pictum 

3899. L. bifidum Penn 

3900. L. purpureum 

3901. L. amplexicaule BB Tir 

* L. flexuosum Tir 

3905. Leonurus cardiaca 

3907. Ballota nigra I. subsp. foetida II. subsp. ruderalis J20 

3909. B. acuta I. subsp. acuta R Sch II. subsp. macedonica A&S Tir 

3912. Stachys jacquini 

3913. S. scardica 

3917. S. officinalis I. subsp. officinalis Sch Tir II. subsp. skipetarum Jav Sch 

3920. S. cretica 

3923. S. sericophylla Sch 

3925. S. germanica I. subsp. heldreichii Sch II. subsp. germanica 

3926. S. reinerti 

3927. S. obliqua Sch 

3928 II. S. alpina subsp. dinarica H17 Sch 

3930. S. decumbens A&S 

3931. S. silvatica 

3936. S. albanica M27 Tir 

3939. S. grandiflora J20 Sch 

3942. S. atherocalyx 

3945. S. beckeana H17 Sch 

3946. S. anisochila 

3947. S. recta (subspp. III, IV, V, VI) 

3961. S. annua 

3963. S. spinulosa A&S Sch 

3965. S. serbica A&S 

3966. S. arvensis H24 

3967. Salvia verticillata 

3968. S. peloponnesiaca 

3970. S. grandiflora Sch 

49

3971. S. officinalis 

3974. S. triloba A&S 

3975. S. ringens 

3978. S. verbenacea I. subsp. verbenacea II. subsp. clandestina 

3981. S. glutinosa 

3982. S. viridis Penn 

3983. S. horminum 

3984. S. sclarea 

3986. S. argentea A&S 

3988. S. candidissima 

3992. S. virgata Gui 

3994. S. bertolonii Penn Sch 

3995. S. tenorii A&S 

3997. S. nemorosa 

3998. S. amplexicaulis 

3999. Ziziphora rigida Tir 

4000. Z. capitate 

4001. Melissa officinalis 

4004. Satureja hortensis 

4006. S. montana 

4010. S. subspicata 

4014. Micromeria juliana 

4016. M. cremnophila A&S 

4017. M. cristata Hep Jav 

4020. M. parviflora 

4029. M. thymifolia Jav 

4031. Calamintha grandiflora 

4032. C. officinalis 

4035. C. nepeta 

4039. C. suaveolens M32 

4040. C. alpina (subspp. I, II, III) 

4041. C. acinos 

4042. C. exigua A&S Sch 

4044. C. vulgaris 

4051. Hyssopus officinalis H24 M32 

4052. Origanum vulgare I. subsp. vulgare II. subsp. viride M32 

4057. Coridothymus capitatus 

4058. Thymus teucrioides A&S 

4062. T. striatus 

4068. T. tosevii 

4074. T. longidens H24 

4083. T. moesiacus 

4084. T. kosaninii Hayek (Prodromus) 

4087. T. jankae 

4089. T. ocheus Markgraf 

4096. T. heterotrichus A&S 

50

4101. T. hirsutus 

4104. T. leucotrichus H24 

4105 II. T. pulegioides subsp. montanus Sch 

4109. T. longicaulis 

4110. T. rohlenae Sch Tir 

4111. T. lykae 

4114. T. illyricus M32 

4115. T. albanus 

4116. T. zygiformis Jav 

4117. T. balcanus 

* T. stojanovii Markgraf 

4119. Lycopus europaeus 

4120. L. exaltatus Jav 

4121. Mentha pulegium 

4127. M. aquatica 

4127 x 4134 

4129. M. pubescens Sch 

4130. M. piperita Tir 

4132. M. longifolia 

4134. M. incana Jav 

4137. M. rotundifolia M32 

Globulariaceae 

4139. Globularia alypus Tir 

4142. G. cordifolia I. subsp. bellidifolia II. subsp. cordifolia 

* G. nana M32 

Plantaginaceae 

4145. Plantago psyllium 

4147. P. indica 

4151. P. altissima Tir 

4152. P. lanceolata 

4153. P. lagopus BB 

4154. P. major 

4155. P. reniformis 

4156. P. gentianoides 

4157. P. media 

4158. P. argentea 

4159. P. montana 

4163. P. bellardii 

4165. P. coronopus 

4167. P. carinata 

4168. P. maritima B94 B96 

4169. P. crassifolia A&S M27 

51

* P. pauciflora Rohl 

Gentianaceae 

4170. Centaurium spicatum 

4172. C. umbellatum I. subsp. umbellatum III. subsp. rumelicum 

4174. C. uliginosum Jav. 

4175. C. tenuiflorum 

4176. C. pulchellum 

4177. C. maritimum 

4178. I. Gentiana lutea subsp. symphyandra 

4179. G. punctata Sch 

4181. G. cruciata 

4182. G. asclepiadea 

4183. G. pneumonanthe b. nepesae Jav M32 

4185. G. kochiana H17 Sch 

4188. G. verna II. subsp. verna III. subsp. tergestina 

4189. G. utriculata Sch 

4192. G. ciliata Jav Hep 

4194. G. crispata 

4195. G. germanica IV. subsp. bulgarica Jav Sch 

4200. Blackstonia perfoliata 

4201. B. serotina J20 

4202. Microcala filiformis Tir 

Menyanthaceae 

4204. Nymphoides peltata J20 Sch 

Apocynaceae 

4205. Vinca major B. var. pubescens (V. balcanica) 

4211. Nerium oleander 

4213. Periploca graeca 

4214. Gomphocarpus fruticosa A&S 

4216. Cynanchum acutum 

4218. C. huteri 

4219. C. speciosum M27 Tir 

4222. C. vincetoxicum B. var. nivale 

4226. C. fuscatum 

4227. Marsdenia erecta 

Oleaceae 

4228. Fraxinus ornus 

4229. F. excelsior 

52

4230. F. oxycarpa 

* F. angustifolia Tir 

4233. Forsythia europaea 

4234. Syringa vulgaris Tir 

4235. Phillyrea latifolia 

4236. P. media 

4237. P. angustifolia B94 

4238. Olea europaea 

4239. Ligustrum vulgare 

4240. Jasminum officinale Tir 

4241. J. fruticosa B96 B99 

Rubiaceae 

4244. Putoria calabrica 

4245. Sherardia arvensis 

4246. Crucianella latifolia 

4249. C. angustifolia A. var. angustifolia B99 Tir B. var. oxyloba H24 

4251. Asperula arvensis J20 

4252. A. taurina 

4254. A. doerfleri 

4260. A. odorata 

4261. A. laevigata Tir 

4264. A. scutellaria 

4265. A. chlorantha 

4280 I. A. aristata subsp. longiflora 

4283. A. cynanchica A&S Sch 

4293. Galium matteji B94 

4295. G. aristatum Tir 

4296. G. mollugo I. subsp. mollugo II. subsp. erectum A&S B17 

4297. G. lucidum I. subsp. gerardi II. subsp. corrudaefolium 

4298. G. cinereum Tir 

4302. G. pseudoaristatum 

4307. G. pumilum 

4308. G. anisophyllum 

4312. G. verum 

4313. G. firmum 

4319. G. purpureum 

4320. G. apiculatum A&S B96 

4324. G. degenii B96 

4333. G. aparine 

4337. G. parisiense Tir 

4338. G. divaricatum M32 

4340. G. zacynthium A&S B99 

4344. G. palustre I. subsp. palustre Sch Tir II. subsp. elongatum 

4345. G. constrictum A&S Sch 

53

4347. G. tricorne A&S 

4349. G. murale BB 

4350. G. verticillatum A&S 

4351. G. rotundifolium 

4355. G. cruciata 

4357. G. pedemontanum 

4358. Valantia aprica 

4359. V. muralis 

4361. Rubia tinctoria Tir M42? 

4363. Rubia peregrina J20 Tir 

Adoxaceae 

4365. Adoxa moschatellina A&S 

Caprifoliaceae 

4366. Sambucus ebulus 

4367. S. nigra 

4369. Viburnum opulus Tir 

4370. V. lantana 

4371. V. tinus 

4373. Lonicera xylosteum 

4376. L. alpigena I. subsp. alpigena II. subsp. formanekiana 

4380. L. periclymenum Tir 

4381. L. etrusca 

4382. L. caprifolium 

4383. L. implexa Pamp Tir 

Valerianaceae 

4384. Valerianella echinata H24 

4387. V. coronata 

4391. V. eriocarpa Tir 

4392. V. dentata 

4394. V. truncata A&S (V. eriocarpa sensu A&S) 

4396. V. rimosa 

4399. V. locusta 

4401. V. carinata M27 

4402. Fedia cornucopiae BB 

4403. Valeriana officinalis 

4405. V. dioscoridis 

4406. V. tuberosa 

4409. V. bertiscea Jav 

4412. V. montana 

4413. V. crinii Sch 

54

4416. V. pancicii 

4418. Kentranthus velenovskyi Sch 

Dipsacaceae 

4422. Morina persica 

4425. Dipsacus silvester 

4426. D. laciniata 

4428. Cephalaria transsilvanicus 

4431. C. leucantha 

4434. C. ambrosioides 

4437. C. pastricensis 

4438. Succisa pratensis Jav 

4439. Succisella inflexa 

4440. S. petteri J20 

4443. Knautia integrifolia 

4446. K. macedonica 

4447. K. arvensis Sch Tir 

4450. K. purpurea H24 J20 

4454. K. midzorensis H24 

4455. K. silvatica Sch 

4456. K. drymeia 

4461. Pterocephalus perennis 

4463. P. plumosus 

4464. Tremastelma palaestinum Tir 

4468. Scabiosa graminifolia 

4471. S. crenata 

4473 II. S. argentea subsp. ucranica 

4481. S. rotata M32 

4482. S. atropurpurea 

4484. S. leucophylla Sch 

4486. S. lucida Sch 

4487. S. columbaria II. subsp. portae III. subsp. dubia IV. subsp. gramuntia Sch 

4489. S. ochroleuca 

4491. S. webbiana 

4492. S. silaifolia Jav Sch 

4493. S. triniaefolia 

4496. S. silenifolia Tir 

4497. S. tenuis 

Campanulaceae 

4511. Campanula lingulata 

4513. C. sibirica B. var. divergens 

4516. C. alpina H24 Jav 

4523. C. macrostachya Sch 

55

4524. C. cervicaria M32 Sch 

4527. C. moesiaca Sch 

4528. C. glomerata I. subsp. glomerata II. subsp. hispida Sch Tir 

4529. C. foliosa 

4530. C. tymphaea 

4533. C. garganica var. albanica M32 

4539. C. hawkinsiana 

4541. C. cochlearifolia 

4543 I. C. scheuchzeri subsp. witasekiana Jav Tir 

4544. C. linifolia II. subsp. justiniana Sch III. subsp. albanica Jav 

4545. C. rotundifolia IV. subsp. rotundifolia Hep K48 V. subsp. pinifolia H24 Sch 

VI. subsp. velebitica VIII. subsp. racemosa Sch 

4547. C. trachelium I. subsp. trachelium II. subsp. athoa 

4549. C. rapunculoides Sch 

4550. C. bononiensis 

4551. C. pyramidalis J20 Sch 

4552. C. versicolor 

4556. C. erinus 

4558. C. persicifolia 

4559. C. spathulata I. subsp. spruneriana Bm33 II. subsp. spathulata 

4560. C. patula I. subsp. patula Sch Tir II. subsp. abietina 

4561. C. rapunculus 

4562. C. sparsa I. subsp. frivaldszkyi II. subsp. sphaerothrix 

4561. C. pyrygia M32 

4564. C. ramosissima 

* C. debarensis R 

4571. Legouzia speculum-veneris 

4573, L. hybrida A&S Tir 

4579. Asyneuma trichocalycinum 

4580. A. comosiforme H24 

4582. A. canescens 

4584. A. anthericoides Hep 

4585. A. lobelioides M27 

4586. A. limonifolium 

4588. A. tenuifolium Sch 

4593 I. Phyteuma orbiculare subsp. flexuosum 

4594. P. pseudorbiculare 

4597. Edrianthus parnassicus 

4600. E. tenuifolius Gui Sch 

4601. E. graminifolius 

4602. E. wettsteinii Gui 

4605. E. serpyllifolius 

4606. Jasione montana H24 

4608. J. heldreichii A&S Jav 

4609. J. orbiculare 

56

Compositae 

4613. Eupatorium cannabinum 

4614. Solidago virga-aurea I. subsp. virga-aurea Sch Tir II. subsp. alpestris 

4617. Bellis perennis 

4619. B. sylvestris 

4621. B. annua BB 

4624. Aster bellidiastrum Jav Sch 

4625. A. alpinus I. subsp. dolomiticus H24 Sch II. subsp. alpinus M32 

4626. A. amellus BB 

4627. A. tripolium BB H24 

4630. A. punctatus J20 

4631. A. albanicus 

4634. A. linosyris J20 Tir 

4636. Erigeron polymorphus 

4639. E. epirotica H24 Sch 

4640. E. alpinus 

4642. E. acer 

4643. E. crispus A&S B94 

4644. E. canadensis 

4649. Evax pygmaea J20 

4656. Filago germanica 

4657. F. spathulata F 

4658. F. arvensis Jav Tir 

4659. F. montana Jav 

4660. F. gallica 

4661. Antennaria dioica 

4662. Leontopodium alpinum Tir 

4665. Gnaphalium silvaticumI 

4666. G. norvegicum K48 

4667. G. hoppeanum 

4668. G. roeseri B. var. pichleri 

4669. G. supinum 

4672. Helichrysum orientale Tir 

4673. Helichrysum plicatum 

4679 II. Phagnalon rupestris subsp. graecum BB 

4681. Inula helenium 

4682. I. germanica 

4682 x 4683 II 

4683 II. I. salicina subsp. aspera 

4684. I. spiraeifolia Sch 

4685. I. hirta Sch 

4686. I. ensifolia 

4687. I. britannica 

4688. I. oculus-christii 

4690. I. conyza 

57

4692. I. bifrons A&S 

4693. I. candida var. candida BB var. acarnaniana R 

4694. I. crithmoides 

4695. I. viscosa 

4696. I. graveolens 

4697. Pulicaria sicula 

4698 B. P. vulgaris var. graeca 

4699. P. odora A&S Tir 

4700. P. dysenterica 

4703. Pallenis spinosa 

4705. Telekia speciosa 

4707. Asteriscus aquaticus 

4709. Ambrosia maritima A&S B96 

4712. Xanthium strumarium 

4713 II. X. saccharatum subsp. italicum 

4714. X. spinosum 

4715. Helianthus annuus Tir 

4716. H. tuberosus Tir 

4717. Bidens cernuus J20 

4718. B. tripartitus 

4723. Santolina chamaecyparissus Tir 

4725. Anthemis tinctoria 

4726. A. triumfetti 

4728. A. cota 

4731. A. brachycentros M32 Sch 

4733. A. austriaca H24 Jav 

4734. A. orientalis (subsp. I, II, III) 

4746. A. arvensis 

4753. A. chia 

4757. A. cotula 

* A. praecox 

4758. Ormenis mixta 

4761. Achillea ageratifolia I. subsp. ageratifolia II. subsp. aizoon H24 Jav 

4762. A. lingulata Sch 

4765 I. A. atrata subsp. multifida Jav (Korab) 

4766. A. abrotanoides 

4767. A. clavenae 

4771. A. fraasii 

4772. A. canescens Sch 

4773. A. grandifolia Jav Sch 

4776. A. nobilis 

4777. A. virescens Sch 

4778. A. asplenifolia Rohl Tir 

4779. A. crithmifolia var. crithmifolia Jav b. var. doerfleri H24 

4780 A. millefolium I. subsp. millefolium Tir II. subsp. collina 

4781. A. fililoba M27 

58

4782. A. setacea 

4783. A. tanacetifolia I. subsp. tanacetifolia Jav Sch II. subsp. stricta Sch 

4786. A. holosericea 

4787. A. chrysocoma H24 M27 

4788. A. clypeolata 

4790. A. baldaccii B96 Sch 

4791. A. coarctata 

4800. Diotis maritima A&S 

4803. Myconis myconis 

4805. Leucanthemum vulgare I. subsp. triviale II. subsp. montana A&S 

III. subsp. pallens J20 Sch 

4809. Tanacetum cinerariaefolium J20 

4810. T. larvatum 

4813. T. parthenium Tir 

4814. T. corymbosum 

4815. T. macrophyllum 

4816. T. vulgare 

4817. T. millefolium Tir 

* T. albanicum M32 

4822. Tripleurospermum caucasicum H24 

4824. T. inodorum Tir 

4825. T. tenuifolium 

4828. Matricaria chamomilla J20 

4831. Cotula anthemoides [H24] 

4832. Artemisia vulgaris 

4833. A. coerulescens 

4835. A. absinthum 

4836 B. A. paniculatum var. visianiana B96 

4838. A. lobelii (vars. A, B, C) 

4841. A. annua Tir 

4842. A. scoparia 

4843. A. campestris Tir 

4848. A. petrosa H24 Jav 

4849. Tussilago farfara 

4850. Petasites doefleri 

4851 B. P. hybridus var. ochroleucus 

4853. P. albus Tir 

4856. Homogyne alpina H24 (Koritnik) 

4859 B. Adenostyles alliariae var. kerneri 

4860. A. orientalis K48 

4863. Doronicum caucasicum H24 Penn 

4864. D. columnae 

4865. D. austriacum 

4867. D. orphanidis K48 Tir 

4868. D. grandiflorum Sch 

4873. Senecio aucheri b. korabensis 

59

4875. S. procerus 

4876. S. bosniacus 

4877. S. wagneri Tir 

4878. S. campester subsp. capitatus Sch 

4880. S. spathulifolius Tir 

4881. S. thapsoides I. subsp. thapsoides II. subsp. visianianus Bm33 Jav 

4884. S. transsilvanicus K48 

4885. S. doronicum Jav M27 

4886. S. lanatus H24 Sch 

4890. S. paludosus Tir 

4894. S. nemorensis A&S H24 

4896. S. fuchsii Sch 

4897. S. cacaliaster Tir 

4899. S. othonnae H24 Tir 

4900 II. S. abrotanifolius subsp. carpaticus H24 Jav 

4902 B. S. erucifolius var. latisectus Jav (boundary) 

4903 I. S. alpinus subsp. subalpinus Sch 

4904. S. jacobaea A&S 

4905. S. erraticus 

4906 I. S. nebrodensis subsp. rupester 

4907. S. leucanthemifolius BB 

4910. S. vernalis Penn 

4913. S. viscosus 

4914. S. vulgaris 

4916. Calendula officinalis Tir 

4918. C. arvensis BB Penn 

4920. Echinops viscosus A&S 

4924. E. microcephalus Hep Tir 

4926. E. ritro Sch 

4928. E. albidus 

4929. E. commutatus Tir 

4930. E. banaticus 

4931. E. sphaerocephalus B96 

4935. Xeranthemum annuum 

4936. X. inapertum A&S 

4937. X. foetidum 

4938. Amphoricarpus neumayeri 

4939. Carlina acanthifolia 

4940. C. acaulis 

4942. C. vulgaris I. subsp. vulgaris II. subsp. intermedia Sch 

4944. C. corymbosa 

4945. C. graeca B96 F 

4948. C. lanata 

4953. Arctium lappa A&S 

4955. A. minus Jav Tir 

4958. Staehelina uniflosculosa 

60

4966. Jurinea mollis I.subsp. glycacantha II. subsp. mollis Tir 

4972. Carduus leiophyllus 

4973. C. macrocephalus Tir 

4974. C. nutans I. subsp. nutans III. subsp. micropterus J20 

4980 I. C. candicans subsp. candicans 

4985. C. armatus 

4987. C. ramosissimus Jav 

4989. C. acanthoides 

4990. C. scardicus 

4991. C. pycnocephalus 

4992. C. marmoratus A&S 

4997. Notobasis syriaca B96 

4999 II. Cirsium ligulare subsp. albanum 

5000. C. vandasii 

5001. C. eriophorum B96 Tir 

5004. C. intraspinulosum 

5012 II. C. boujarti subsp. wettsteinii H24 

5013. C. lanceolatum 

5014. C. italicum Tir 

5015. C. arvense 

5018. C. tymphaeum A&S 

5019. C. appendiculatum 

5020. C. tricephalodes Sch 

5021. C. rivulare Sch 

5027. C. eristhales Tir 

5028. C. candelabrum 

5030 B. C. canum var. subtymphaeum H24 

5031. C. heterotrichum K48 

5034. C. creticum 

5035. C. palustre Tir 

5036. C. afrum 

5037. C. strictum 

5038. C. stellatum A&S 

5041. Picnomon acarna 

5044. Silybum marianum 

5045. Tyrimnus leucographus B96 M27 

5046. Galactites tomentosa Tir 

5047. Onopordon acanthium 

5056. O. illyricum 

5057. Crupina vulgaris 

5058. C. crupinastrum Pamp 

5059. Serratula tinctoria Jav (boundary) 

5060 II. S. radiata subsp. cetinjensis H24 

5067. Centaurea triumfetti 

5068. C. epirota M31 Penn 

5073. C. tuberosa Jav 

61

5075. C. cyanus Jav Tir 

5079. C. kosanini 

5082. C. scabiosa II. subsp. fritschii Jav Sch III. subsp. spinulosa Sch 

5085. C. candelabrum Sch 

5086. C. atropurpurea 

5090. C. graeca 

5092 II. C. spruneri subsp. guicciardii B94 B96 

5104. C. salonitana 

5106. C. macedonica M27 

5107. C. rupestris Sch 

5108. C. nicolai B96 Sch 

5118. C. leucomelaena B94 M32 

5129 II. C. cuneifolia subsp. sublanata 

5133 II. C. stoebe subsp. micranthos 

5140. C. grisebachii 

5168. C. zuccariniana B96 R 

5173. C. cadmea Tir 

5176. C. ptarmicaefolia A&S 

5177. C. deustiformis 

5179. C. deusta I. subsp. deusta II. subsp. concolor 

5186. C. jacea I. subsp. angustifolia III. subsp. haynaldii H24 IV. subsp. jacea Jav Sch 

5192. C. phrygia II. subsp. pseudophrygia K48 III. subsp. stenolepis H24 

5193 II. C. nervosa subsp. nervosa Jav Sch 

5199. C. solstitialis 

5202. C. calcitrapa 

5215. Carthamus lanatus 

5221. C. coeruleus Hep 

5222. Cnicus benedictus 

5224. Scolymus hispanicus 

5229. Cichorium intybus 

5230. C. pumilum A&S 

5233 I. Lapsana communis subsp. communis 

5238. Tolpis virgata 

5243. Zazintha verrucosa 

5244. Rhagadiolus stellatus 

5245. Hedypnois rhagadioloides (subsp. I, II, III) 

5246. Hypochaeris glabra 

5247. H. radicata 

5248. H. maculata I. subsp. maculata II. subsp. pelivanovicii Sch *subsp. hispida Sch 

5250. H. cretensis 

5251. H. aethnensis A&S 

5254. Leontodon illyricus 

5255 II. L. montanus subsp. montanus Sch 

5257. L. hispidus 

5258. L. crispus (subsp. I, II, IV) 

5261. L. fasciculatus Tir 

62

5263. L. taraxacoides 

5264. L. tuberosus 

5266 III. Picris hieracioides subsp. hieracioides 

5267. P. sprengeriana Pamp 

5268. P. pauciflora A&S 

5269. P. echioides 

5270. Urospermum dalechampii Tir 

5271. Urospermum picroides 

5272. Tragopogon balcanicus 

5273. T. longirostris 

5278. T. samaritani (probably an error) 

5279. T. pratensis (subspp. I and II) Tir 

5280. T. rumelicus H24 

5282. T. tommasinii Sch 

5288. T. hybridus 

5289. Scorzonera purpurea 

5290. S. rosea 

5291. S. hispanica 

5294. S. austriaca H17 Sch 

5296. S. rumelica A&S 

5297. S. mollis M32 

5300. S. doriae 

5302. Podospermum canum 

5303. P. laciniatum Sch 

5307 I. Lagoseris sancta subsp. bifida Tir 

5312. Chondrilla juncea 

5313. Willemetia stipitate Jav 

5316. Taraxacum laevigatum I. subsp. calocephalum Jav II. subsp. pidicolum Jav Penn 

5319. T. haussknechtii A&S 

5323. T. officinale (subsp. I, II, III) 

5324. Reichardia picroides 

5325. R. macrophylla 

5330. Cicerbita alpina 

5333. C. pancicii 

5336. Sonchus maritimus A&S 

5337. S. asper I. subsp. asper A&S II. subsp. glaucescens 

5338. S. oleraceus 

5340 C. Mycelis sonchifolia var. velenovskyi A&S K48 

5341. M. glandulosa A&S 

5343. M. muralis 

5346. Lactuca graeca A&S 

5347. L. cretica A&S 

5348. L. sativa Tir 

5349. L. serriola 

5341. L. saligna 

5352. L. quercina Sch 

63

5355. L. contracta A&S 

5356. L. viminea J20 Sch 

* L. tenerrima B96 

5358. Prenanthes purpurea 

5359. Crepis bulbosa M27 

5360. C. viscidula 

5361. C. conyzifolia Sch Tir 

5363. C. baldaccii I. subsp. baldaccii II. subsp. albanica Jav Sch 

5364. C. geracioides M32 

5371. C. alpestris H24 Sch 

5372. C. jacquini 

5375. C. turcica A&S B99 

5376. C. pantocsekii Sch 

5381. C. reuteriana A&S 

5386. C. columnae 

5389 I. C. incarnata subsp. dinarica Sch 

5391. C. biennis 

5393. B. C. nicaeenis var. adenantha Jav 

5394. C. capillaris M27 Tir 

5396. C. pulchra Tir 

5398 I. C. neglecta subsp. neglecta 

5399. C. foetida I. subsp. rhoeadifolia III. subsp. glandulosa Sch 

5401. C. bertiscea Jav 

5402. C. leontodioides Tir 

5403. C. setosa 

5404. C. rutilans A&S 

5408. C. rubra A&S Penn 

5410. C. dioscoridis A&S 

Hieracium not included. Individual specimens should be checked. 

64

Part 5: Flora Europaea vol. 5 

This list was originally prepared for volume 5 of Flora Europaea but for various reasons 

was never completed and utilized. Unlike the lists for volumes 1-4 of Flora Europaea this 

list contains all records for Albania that could be traced circa 1965, but is certainly not 

complete, the most obvious omission being Flora of Tirana which was not directly 

available to me when this list was compiled. There are about 90 species reported here 

which are not given for Albania in Flora Europaea and there are about 50 species 

reported for Albania in Flora Europaea vol. 5 which are not in this list. The latter 

probably represent records which were not traced in 1965, particularly Flora of Tirana, 

some reidentification of specimens reported under another name here, and some which 

occur in areas that were included in “Albania” as defined by Hayek but which are not 

known from present-day Albania. Each of these has to be checked independently. 

The nomenclature and sequence within families follows Hayek with a few minor 

discrepancies, but some species are reported both under the name used in Hayek and 

under a synonym (especially in Gramineae). The family sequence follows Flora 

Europaea. 

Alismataceae 

Alisma plantago-aquatica J20, Jav., Sch. 

A. lanceolatum A & S 

Butomaceae 

Butomus umbellatus J20, Penn., Sch 

Hydrocharitaceae 

Vallisneria spiralis Sch 

Juncaginaceae 

Triglochin bulbosa M 27 

T. palustris Jav, A & S 

Potamogetonaceae 

Potamogeton pectinatus Sch 

P. perfoliatus J20, M27, Roh, Sch 

P. lucens J20, M27 

P. crispus J20, M27, Sch 

P. natans Jav 

P. fluitans Sch (?= P. nodosus) 

P. gramineus Sch 

Ruppiaceae 

Ruppia maritima subsp. spiralis Sch 

Posidoniaceae 

Posidonia oceanica M? 

65

Zosteraceae 

Zostera nana B96 

Zanichelliaceae 

Zanichellia palustris J20 

Najadaceae 

Najas marina J20 Sch 

N. minor J20 Sch 

Liliaceae 

Narthecium scardicum K48, Sch 

Veratrum album var. album Jav, Sch 

var. flavum M32 var. viride H17 H24 M32 M27 Jav Sch 

Colchicum autumnale J20, M32 

C. parnassicum BB 

C. kochii M32 

C. lingulatum M32, A & S 

Asphodelus albus H17, M32, M27, Jav, K48, Sch 

A. microcarpus Gr, H24, J20, M32, M27, A & S, Hep, Sch 

A. fistulosus Gr, H24, A & S, Sch 

Asphodeline lutea A & S, Sch, H24, J20 

A. liburnica M32, M27, Jav (Koritnik), Penn, Form, Sch 

A. taurica A & S 

Anthericum liliago H24, Jav, Sch 

A. ramosum M27, Hep, Sch 

Gagea pusilla A & S, Sch 

G. minima H24 

Allium ampeloprasum M27, Penn, Sch (A. leucanthum var. tiranense Roh) 

A descendens H24, A&S, Sch 

A. margaritaceum A&S , Roh, Form 

A. dalmaticum B01, H24, J20, Jav, A&S, Sch, Roh 

A. arvense A&S, Sch 

A. sphaerocephalum J20, Jav, A&S, Hep, B94, Sch 

A. chamaespathum A&S 

A. ursinum Sch 

A. vineale var. vineale A&S var. compactum M32 

A. schoenoprasum subsp. sibiricum H24 

A. meteoricum. M32, Bm 33, Sch 

A. moschatum M27, A&S, Hep, B94 

A. cupani H24, A&S 

A. neapolitanum Gui 

A. rotundatum var. preslianum Pamp 

A. suaveolens A&S, Jav (Cafa Morins) 

A. subhirsutum A&S, Penn, Sch 

A. roseum var. roseum M32, M27, Pamp, Penn, Sch var. javeriense M27 

66

A. nigrum A&S 

A. pulchellum H24, M27, Jav 

A. guiccardii H24, J20, Sch 

A. flavum M32, H24, J20, Jav (Koritnik), A&S, K48, Hep, Roh, Sch 

A. carinatum H24, M32, A&S, Sch 

A. fuscum A&S 

Lilium carniolicum var. albanicum H24, M32, M27, Jav, K48, Gui, Sch 

L. heldreichii A&S 

L. martagon M27, Jav, Gui, Sch 

Fritillaria tenella H24 

F. gracilis H17, H24, M27, Jav, Penn, Gui, Sch 

F. neglecta Bm33 

F. macedonica M32, M27, M42 

F. ionica M32, A&S, Sch 

Tulipa silvestris M27 subsp. grisebachiana H24, M32, B99 subsp celsiana R, M27, A&S 

T. australis Gui 

Erythronium dens-canis M32, M27, Hep, Sch 

Urginea maritima A&S, Ster! (1954) 

Scilla bifolia H17, M31, Hep, Penn, Sch var. nivalis M27, Gui 

S. autumnalis J20, M32, Jav (Cafa Morins), A&S, BB, Sch 

Ornithogalum comosum H24, A&S 

O. tenuifolium H24, M32, M27, Sch 

O. umbellatum Jav, 

O. nanum Penn, BB 

O. exscapum M32, M27 

O. pyramidale subsp. narbonense M27, A&S, Penn, Sch subsp. pyramidale Sch 

O. sphaerocarpum H17, H24, Jav, Sch 

Hyacinthus romanus Penn 

Bellevalia romana H24, M27, A&S, BB 

Muscari tenuiflorum M27, A&S, H80, Sch 

M. comosum H24, J20, M27, Penn, Sch 

M. weissii var. karpatanum Jav 

M. racemosum H24, M32, M27, Jav(Koritnik), A&S, Sch 

M. neglectum Pamp 

M. botryoides H24, M32, M27, Sch var. kerneri Penn 

M. commutatum var. albanicum BB 

Asparagus tenuifolius J20 

A. verticillatus Form 

A. maritimus M27 

A. acutifolius H24, H17, J20, M32, M27, Jav, A&S, Hep, Roh, Form, Sch, Weiss 

Ruscus aculeatus H24, J20, M32, Jav, Hep, Roh, Sch 

Polygonatum multiflorum Jav, A&S 

P. verticillatum M32, K48 

P. officinalis Penn 

Convallaria majalis H24 

Paris quadrifolia A&S 

67

Smilax aspera J20, M27, Penn, Form, Sch 

Amaryllidaceae 

Galanthus nivalis Penn, Sch 

Leucojum aestivum J20, M27, A&S, Penn, Sch 

Pancratium maritimum M27, A&S, Hep, Hep, Form, Sch 

Narcissus radiiflorus H24, A&S, Sch 

N. tazetta BB, Hep 

N. serotinus BB 

N. poeticus Hep 

Sternbergia lutea BB 

S. sicula BB 

Dioscoreaceae 

Tamus communis H24, J20, M32, H17, M27, Jav, Pamp, Sch 

Dioscorea balcanica M32, H24, Jav, M42 

Iridaceae 

Crocus albiflorus A&S, Sch 

C. scardicus A&S 

C. longiflorus M32, M27 

C. dalmaticus M32, M27 

C. nivalis M32, M27 

C. veluchensis M32, M27, Bm33 

C. neapolitanus H24(Koritnik), Sch 

C. hadriaticus BB 

C. olivieri R 

C. boryi BB 

C. cancellatus BB 

C. heuffelianus Penn, Sch 

Romulea bulbocodium M32, BB, Hep 

R. linaresii M32 

(R. ramiflora var. parlatoris BB R. rollii BB) 

Iris aphylla M32, M27 

I. pallida M32, J20, M27, Sch 

I. sintenisii M32, M27, Penn, B94 

I. variegata H24(S. Pastrik) 

I bosniaca H24(Pastrik) 

I. pseudacorus H24, J20, Hep, Sch 

I. sisyrinchium Hep 

I. graminea(?I. sintenisii) J20 

Gladiolus segetum A&S 

G. paluster var. paluster H17, H24, Penn, Bm33, Sch var. reuteri M32, Sch 

G. communis Gr, M27, ?H24 

G. illyricus Gr, M27, Sch, ?H24 

G. byzantinus Pamp, Penn 

68

Hermodactylus tuberosus BB, Hep 

Juncaceae 

Juncus alpinoarticulatus A&S, Jav, K48, Sch 

J. anceps subsp. hercegovinus A&S 

J. longicornis A&S 

J. maritimus A&S, H24, B94 

J. tommasinii A&S 

J. acutiflorus B94(sylvaticus) Sch 

J. trifidus M32, K48, Jav(border) 

J. bufonius J20, M27, Jav, Sch, H24 

J. articulatus J20, H24, Jav, A&S, Roh, Sch, Weiss(lamprocarpa) 

J. subulatus B96 

J. inflexus(glaucus) J20, M27, B94, Sch, B01 

J. compressus J20, M27, Sch 

J. tenageia Sch 

J. atratus Sch 

J. acutus M27, A&S, BB, Roh, Sch, Form, Weiss 

J. effusus Jav 

J. alpigenus Jav Sch(thomasii) 

Luzula spicata H24, H17(border), M27, Sch, M32 var. pindica A&S, K48 

L. campestris M32, Sch 

L. silvatica H24, A&S, M27, K48 (maxima) 

L. forsteri M27, A&S, Sch 

L. pilosa H17 (Vuti) 

L. multiflora var. alpestris H17, M27 

L. luzulina Sch 

Gramineae 

Bromus ramosus subsp. ramosus A&S, K48 

B. cappadocicus A&S 

B. condensatus var. condensatus H17 var. microtichus H17, J20 

B. fibrosus var. fibrosus Sch 

B. erectus var. erectus Sch var. transsilvanicus H24 

B. pannonicus Sch 

B. inermis Sch 

B. madritensis H24, A&S, Sch 

B. tectorum H24 

B. japonicus(patulus) B99, Sch 

B. sterilis H17, H24, J20, Sch 

B. arvensis A&S, Roh(turcicus), Sch, Jav 

B. racemosus J20, Sch 

B. mollis J20, M27, A&S, Sch 

B. commutatus M27 

B. squarrosus J20, A&S, Roh, Sch 

B. intermedius J20, A&S, Jav 

69

B. molliformis A&S, Sch, Jav 

B. scoparius A&S 

Brachypodium pinnatum var. pinnatum A&S, K48, Sch, Jav 

var. rupestre H24, J20, Sch 

B. silvaticum J20, A&S, Roh, Sch, Jav 

B. distachyum A&S, J20, Form, Sch 

B. ramosum Gr, M32, M27, A&S 

B. serpentinii A&S, M27(as ramosum), M42, K48 (albanicum) 

Agropyron litorale J20, A&S, Sch 

A. pungens Form 

A. intermedium J20, Jav 

A. trichophorum A&S(=intermedium) 

A. junceum A&S 

Triticum ovatum J20, Sch 

T. ovatum J20, Sch var. triaristatum H24, J20, M32, M27, A&S, Roh, Sch, Jav 

var. biuncinale Sch 

T. triuncinale Gr., A&S, Roh, Sch 

T. uniaristatum A&S 

T. monococcum M32, M42, Roh 

T. villosum H24, J20, M27, A&S, Roh, Form, Sch, Jav 

Secale montanum M42 

Hordeum europaeum M32, A&S, K48(silvaticum) 

Hordeum asperum H24 

H. crinitum A&S 

H. bulbosum J20, Sch 

H. hystrix A&S 

H. gussoneanum A&S Sch 

H. maritimum M27 

H. murinum subsp. leporinum J20, Sch, Jav 

Sesleria nitida H24, J20, M32, M27, Sch, Jav (skanderbeggii) 

S. heufleriana M27, A&S, K48 

S. coerulea subsp. varia H17, J20, Sch 

S. tenerrima A&S, B99 

S. tenuiflora H17, Sch, Jav 

S. argentea var. cylindrica Sch subsp. robusta Sch 

S. coerulans Sch 

S. korabensis Jav 

S. comosa Jav 

Echinaria capitata A&S 

Arundo plinii J20, Weiss, B94 

Phragmites communis A&S, J20, Sch 

Molinia coerulea A&S, J20 var. arundinaria J20 

Eragrostis megastachya J20, B01, Roh, A&S (as cilianensis) 

E. minor J20 

E. pilosa B01, J20, Sch 

Melica transsilvanica J20, M32, A&S, Sch, Jav 

70

M. ciliata Sch, Jav(nebrodensis), H17, J20 (both as subaristata) 

M. minuta A&S, 

M. uniflora H24, M32, K48, Jav 

Briza maxima J20, M27, Sch 

B. media A&S 

B. minor J20, M27 

B. spicata A&S 

Aeluropus litoralis B96, M27, A&S 

Dactylis glomerata subsp. glomerata H24, K48, Jav 

subsp. hispanica J20, Sch, M27 var. abbreviata Jav, Roh 

Cynosurus cristatus J20, Sch 

C. echinatus J20, M27, Sch 

Catabrosa aquatica J20, Sch 

Poa annua H17, J20, BB, M27 var. parviflora Sch 

P. bulbosa H17, H24, J20, M27, A&S, BB, Penn, Sch, Jav 

P. pumila H17, A&S, Sch, Jav 

P. compressa Jav, Roh 

P. pannonica A&S (as sterilis) 

P. alpina subsp. badensis H17, J20, M27, Sch subsp. alpina H17, J20, B94, Sch, Jav 

P. caesia B96, Sch 

P. cenisia Sch, Jav var. dolosa B94 

P. ursina Sch, Jav 

P. nemoralis H17, A&S, K48, Sch, Jav 

P. trivialis H17, J20, M27, Sch 

P. silvicola A&S, Jav 

P. pratensis subsp. pratensis Sch, H24(strigosa) subsp. angustifolia Sch 

P. chaixii A&S (check against next species in Hayek) 

Glyceria fluitans J20, Sch, Jav 

G. plicata A&S, K48 

Festuca ovina M27, B96 (as glauca), 

F. koritnicensis H24 

F. panciciana H17, H24, J20, Sch, Jav 

F. heterophylla K48 

F. dalmatica J20, K48, Sch 

F. vallesiaca J20, K48 (as pseudovina) 

F. sulcata Sch 

F. halleri subsp. halleri Sch 

F. laevis Sch 

F. violacea H24, M32, M27, Sch, Jav (as korabensis) 

F. rubra H17, Sch 

F. varia subsp. varia R, H24 subsp. pungens R, H24, Sch, Jav 

F. xanthina M32, 

F. paniculata H24, J20, M32, Sch 

F. affinis var. coarctata A&S 

F. gigantea K48 

F. spadicea K48, Jav (boundary) 

71

F. arundinacea A&S 

F. elatior J20, A&S, Sch 

Vulpia myurus H24, J20, Roh, Sch, Jav, M27 

V. sciuroides M27 

V. uniglumis M27 

V. fasciculata Roh 

V. ciliata J20, Sch 

V. dertonensis Roh, Sch, Jav 

Scleropoa rigida J20, M27, Roh, Form, Sch, Jav 

Lolium temulentum J20, Sch 

L. strictum H24, J20, Sch, B94(rigidum) 

L. perenne H24, J20, M32, Sch, Jav 

L. multiflorum subsp. italicum Sch 

L. remotum Sch 

Lepturus cylindricus J20, 

Psilurus aristatus Gr, A&S 

Nardus stricta H24, K48, Jav 

Sieglingia decumbens A&S 

Danthonia calycina M27, Sch 

Koeleria splendens H24, A&S, J20, Bm33 (subcaudata), Sch, Jav, Jav (macedonica) 

K. pyramidata A&S 

K. gracilis J20 (var. glabra), Sch, Jav 

K. phleoides A&S, M27, J20, Roh, Sch 

Trisetum flavescens A&S, K48 (pratense), Jav (tenuiformis) 

T. myrianthemum J20 

T. distichophyllum subsp. albanicum Sch, Jav 

Ventenata dubia H24, A&S 

Avena convoluta A&S 

A. compacta B96, Jav 

A. sterilis M32 

A. barbata H24, J20, A&S, B94, Sch 

Arrhenatherum elatior A&S 

Gaudinia fragilis B96, H17, J20, A&S, Sch 

Holcus lanatus J20, Jav, Sch 

Deschampsia caespitosa H24, A&S, K48, Jav 

D. media B01 

D. flexuosa H17, K48, Sch 

Aira capillaris H24, J20, M27, Jav, Sch 

A. caryophyllea Sch 

Calamagrostis pseudophragmites Jav, Bm33 

C. epigeos A&S, Sch 

C. varia H24, Jav, K48, Sch 

C. arundinacea M32, Sch 

Ammophila arenaria M27 

Agrostis alba H24, J20, Jav, A&S, K48, Sch 

A. capillaris J20, A&S 

72

A. canina J20, Sch 

A. castellana subsp. byzantina B96, H24, J20, Jav 

A. rupestris H17, H24 

A. semiverticillata A&S 

Gastridium ventricosum J20 

Polypogon monspeliensis J20, M32, A&S, Sch 

P. maritimus Weiss 

Lagurus ovatus M32, M27, Penn, Weiss 

Sporobolus arenarius A&S 

S. pungens (Agrostis pungens)Weiss 

Alopecurus gerardii H17, M27, Jav, A&S, K48, B94, Sch 

A. pratensis A&S 

A. myosuroides H24, BB(agrestis) 

A. utriculatus H17, H24, J20, M27, BB, Sch 

Helechloa alopecuroides B96, J20 

Crypsis aculeata B96, A&S, Weiss 

Phleum echinatum Gr, A&S 

P. alpinum var. alpinum K48, Sch var. commutatum H17, B94, Sch 

P. pratense A&S var. debile J20, Sch, Roh, Form 

P. phleoides A&S. 

P. paniculatum H24, Jav 

P. montanum H24, Jav, A&S 

P. michellii var. michellii Sch var. subincrassatum H24, Jav, Sch 

Lasiagrostis calamagrostis H24, J20, Jav, Roh, Sch 

Stipa bromoides B96, H24, J20, Roh, Sch, Form 

S. tortilis A&S 

Stipa pennata var. pennata A&S, K48, B99 var. joannis M27, Jav. ("afa Morins) 

subsp. tirea H24, H17, J20, R, M32, M27 

S. capillata K48 subsp. mediterranea Sch 

Oryzopsis miliacea A&S 

O. holciformis A&S 

Milium effusum A&S 

M. vernale Sch 

Anthoxanthum odoratum H17, J20, K48, Sch 

A. aristatum M27 

A. gracile A&S 

Echinochloa crus-galli Form 

Phalaris bulbosa J20 

P. brachystachys M32 

Oryza sativa B96 (cult.) 

Leersia oryzoides A&S 

Beckmannia eruciformis M32, J20 

Cynodon dactylon A&S, Form, Weiss, Sch 

Digitaria sanguinalis J20, A&S 

D. paspalodes Sch 

Panicum crus-galli A&S, J20 

73

P. repens A&S 

P. miliaceum J20 

Setaria glauca A&S, J20, Sch 

S. verticillata Form 

S. viridis J20, Sch 

Imperata cylindrica J20, M27, BB, Sch 

Erianthus ravennae B96, J20, A&S, Weiss 

E. hostii J20, M32, Roh, Sch 

Phacelurus digitatus B96, M27, A&S, B94 

Andropogon ischaemum H24, J20, M27, Jav, Roh, Sch 

A. distachyum A&S 

Chrysopogon gryllus H24, J20, M32, M27, Jav, Roh, Form, Sch 

Sorghum halepensis J20, M32, A&S, Sch 

Cymbopogon hirtus Gr, J20, M32, A&S, Roh, Form, BB 

Tragus racemosus J20, A&S, B94 

Araceae 

Arum italicum H24, J20, M27, A&S, Sch 

A. maculatum subsp. vulgare Sch subsp. angustatum Sch 

Arisarum vulgare BB 

Lemnaceae 

Lemna minor source unclear 

Sparganiaceae 

Sparganium erectum J20 (=. neglectum fide Hayek) 

S. neglectum A&S, Jav 

Typhaceae 

Typha angustata B96, Roh, A&S 

T. latifolia A&S, Jav 

T. shuttleworthii A&S 

Cyperaceae 

Pycreus flavescens B96, J20, A&S, Sch 

P. longus H24, J20, M32, Jav, A&S, Roh, Sch 

P. rotundus B96, J20, M27, B94, Sch 

Acerellus pannonicus B96, A&S 

Galilea mucronatus A&S 

Fimbristylis dichotomum B96, J20, Hep, Roh 

Cyperus glaber Sch 

C. fuscus B96, J20, Jav, A&S, Hep, Sch 

C. difformis B94 

C. schoenoides B94 

Eriophorum latifolium Jav(Korab), A&S 

E. angustifolium Sch 

74

Schoenoplectus setaceus B96, A&S, Bm 33 

Scirpus cernuus B96, 

S. flavescens Hep 

S. lacuster B96, J20, Form, Sch 

S. tabernaemontani A&S 

Holoschoenus vulgaris subsp. vulgaris Sch, B96, J20, A&S, Jav ("afa Morins) 

var. australis M27, Roh, Sch 

subsp. romanus M27, B96, J20, Weiss, Sch 

Blymus compressus B96, H17, Jav, A&S, Sch 

Bulboschoenus maritimus B96, J20, M27, Jav, A&S, Roh, Sch 

Eleocharis palustris B96, J20, Sch 

E. pauciflora Jav, A&S 

E. acicularis B01, Sch 

E. uniglumis A&S 

E. ovata Sch 

E. carniolica Sch 

Schoenus nigricans H17, J20, M27, Jav("afa Morins), A&S, BB, Sch, Weiss 

(Incl. S. karpatii Penzes fide Sandwith) 

S. mucronatus Weiss 

Cladium mariscus A&S 

Carex curvula B24, M27 

C. divisa M27, A&S, BB, Sch 

C. pairaei Sch 

C. divulsa B96, H17, J20 

C. remota Sch 

C. vulpina B96, J20, K48, A&S 

C. leporina K48, Sch 

C. paniculata B96, A&S, K48, Sch 

C. stellulata Jav, A&S, K48 

C. canescens Jav (Korab) 

C. gracilis A&S 

C. dacica Jav 

C. goodenoughii A&S, K48, Gui, Sch 

C. markgrafii M27, M32 

C. verna H24, A&S, K48 (caryophyllea) 

C. praecox BB 

C. tomentosa A&S 

C. halleriana M27, A&S 

C. humilis M27 

C. ornithopoda A&S 

C. panicea A&S 

C. glauca M31, A&S, BB, K48, J20, H17, M27 

C. pallescens K48, Sch 

C. sempervirens H24, M27, Sch 

C. laevis B96, H17, M32, Jav, A&S, K48, Bm33, Sch 

C. macrolepis M32, M27 

75

C. hirta B96, J20, Gr, M27, Sch 

C. distans H24, H17, J20, M27, A&S, B99, Sch 

C. hostiana Sch 

C. flava Jav 

C. lepidocarpa M32, Sch 

C. oederi A&S 

C. silvatica B96, H17, M27 

C. pseudocyperus A&S 

C. riparia B96, Gr 

C. rostrata B96, Gr, Jav(Korab) 

Orchidaceae 

No attempt has been made to resolve synonymy in this family nor to resolve the names 

vis-à-vis Flora Europaea. 

Ophrys apifera M32, A&S, Penn 

O. fusca M27, BB 

O. bertolonii M27 

O. ferrum-equinum A&S, BB, Penn 

O. lutea H24, A&S 

O. atrata H24 

O. cornuta H24, A&S, Penn 

O. bombyliflora H24, BB, Penn 

O. mammosa ?source 

Orchis coriophora var, coriophora A&S, Sch 

var. fragrans H24, J20, M27, Penn, Sch,, M32, H17 

O. tridentata H17, M27, A&S, M32 

O. provincialis M32, M27 subsp. pauciflora M27, Penn 

O. quadripunctata M32, H17, M27, A&S, H80 

O. laxiflora M32, H24(?), J20, H17, M27, A&S, Pamp, Penn, Sch, Penn(palustris) 

O. cordigera M32, M27, Sch, Jav 

O. sambucina M32, M27, Sch, Penn subsp. lagotis A&S 

O. incarnata M27, A&S 

O. maculata var. macrostachys M32, M27, K48, Jav subsp.saccifera A&S, Hep, B94 

O. bosniaca M32, Sch, Jav 

O. morio subsp. morio Sch subsp. picta M27, A&S 

O. ustulata M27, A&S, Sch 

O. traunsteineri A&S ? 

O. papilionacea A&S 

O. italica Penn 

O. rubra H17, H24 

O. pallens H24(Pastrik) 

O. purpurea H24, A&S 

O. spitzelii H24, A&S 

O. mascula H24, A&S, Penn, Sch, H17 (Vuci) 

Serapias lingua M32, M27, Penn 

S. cordigera M32, M27 

76

S. vomeracea J20, H17, M27, A&S, M32 

S. longipetala H17, M27, Pamp 

S. laxiflora var. columnae M27 

Himantoglossum hircinum A&S, Sch, Jav, H24 

Anacamptis pyramidalis H24(?), M32, J20, H17, M27, A&S, Penn, Sch 

var. brachystachya M32 

A. X weberi M32 

Coeloglossum viride M27, K48, H24, Jav, Sch 

Nigritella nigra M32, M27, Jav(boundary) 

Gymnadenia frivaldii M32 (G. nigra Sch) 

G. conopea M27, A&S, K48, H24, Sch 

G. albida H24, Sch 

Platanthera bifolia M27, Jav, H24 

P. chlorantha M27, A&S 

Epipactis latifolia M32, M27, A&S, Hep, Jav 

E. microphylla A&S, K48 

E. atropurpurea Sch 

E. palustris A&S, Jav 

E. atrorubens K48 (rubiginosa) 

Cephalanthera longifolia M32, M27, Penn 

C. pallens M27 

C. rubra M27, A&S, Jav, H24 

Limodorum abortivum M32, H24, M27, A&S 

Listera ovata M27, A&S 

Spiranthes spiralis H24, J20 

Neottia nidus-avis M32, M27, K48 

Corallorhiza trifida M27 

Aceras hircina H80 

77

Pflanzensammlungen aus Nordalbanien 

Bruno Schütt 

Editor’s note. A copy of this manuscript was made available to me by the Curator of 

Botany at the Uebersee Museum, Bremen (BREM), in about 1961. It appears to have 

been a preliminary listing of Schütt’s collections from northern Albania covering 

approximately the years 1927-1935. Most of his collection came from the area north of a 

line drawn from Shkodra to the point where the river Drin crosses the Albanian border 

into Jugoslavia (Serbia). He also collected a small number of plants north of the river 

Drin, south west of Shkodra. The list consists of about 1500 species and about 4000 

specimens. Many of his collections were the first reliable record of a species from 

Albania, and some are still the only record of the species. It seems that Schütt was unable 

to collect early in the season so the lowland spring flora is not well represented nor is the 

late season flora. Schütt was a schoolteacher by profession so probably collected mainly 

in July and August. Although Schütt was primarily responsible for the identifications he 

appears to have been in contact with Markgraf (working at B at the time) as in one or 

two small papers published by Schütt he acknowledges help by Markgraf and Markgraf 

(1961) mentions Schütt as a contributor to the flora of Albania. His collections of 

Minuartia were examined and determined by Mattfeld. 

Anyone wishing to examine specimens collected by Schütt should contact Dr. Monika 

Steinhof, the Curator of Botany at BREM. The plants from Albania form only a small part 

of Schütt’s herbarium which is not yet fully incorporated into the herbarium at BREM. 

Although Schütt’s manuscript contained a list of localities they were not precisely 

localized and many are in the former Jugoslavia. Also a significant number of localities 

mentioned in the manuscript are not included in Schütt’s list of localities so, for 

convenience, an appendix of localities is provided at the end of the manuscript. 

78

100

101

102

103

104

105

106

107

108

109

110

111

112

113

114

115

116

117

118

119

120

121

122

123

124

125

126

127

128

129

130

131

132

133

134

135

136

137

138

139

140

141

142

143

144

145

146

147

148

149

150

151

152

153

154

155

156

157

158

159

160

161

162

163

164

165

166

167

168

Index of localities in Schütt’s list of plants. 

Schütt included a list of localities in his manuscript, but it includes localities in adjacent 

areas of the former Jugoslavia, and for Albania gives minimal information about the 

precise position of the locality. Also his list is very incomplete so there are many 

localities not mentioned in his list. In a few cases the modern spelling of the name is 

different so it may be difficult to locate the place except by studying good quality maps. 

Places in brackets after a locality are modern Albanian spellings of names or alternative 

names for places which may appear on 20 th century maps A few localities are ambiguous 

and this is pointed out in this index. It is likely that these ambiguities can be resolved by 

examining the specimens concerned and trying to relate these to dates when Schütt was 

collecting in nearby places. A longitude and latitude to the nearest 30” is provided for 

convenience, most localities can be found on the internet if a more precise position is 

needed, although it is doubtful if such precision is meaningful given the circumstances 

under which the collections were made. A longitude and latitude described as 

approximate is one which has been estimated based on collection localities which are 

combinations of two or more places. 

I wish to acknowledge much help from Marash Rakaj (University of Shkodra) and Zoltan 

Barina (National Museum, Budapest) in providing me with up-to-date spellings of placenames 

and finding localities which I had been unable to trace. 

Abatë 42° 18.5’N 19° 49.5” E 

Alessio (Lezhë) 41° 47’N 19° 19.5’E 

Agripa e Gurit (Kripa e Gurit) 42° 15.5’N 19°56’E 

gr. Bardanjolt (? Bardhaj or Bardhanjor) 42° 04.5’N 19° 35.5’E 

kl. Bardanjolt (? Bardhaj) 42° 06’N 19° 36.5’E 

Bërzhet (Bëzhet) 42° 19’N 19° 33.5’E 

Bëstriq (Pashtrik) 42° 13’N 20° 31.5’E (he was not able to visit the summit) (another 

mountain with this name further south, not visited by Schütt) 

Biograd 42° 38.5’N 19° 45.5’E 

Bityq (Bytyçit) 42° 17’N 20° 13’E 

Bogë 42° 24’N 19° 39’E 

Boks 42° 17.5’N 19° 41’E 

Bolanice 42° 36.5’ N 19° 40’E 

Bridzhe (Grexhë) 42° 23’N 19° 26’E 

Brize 42° 14.5’N 19° 52.5’E 

Brzhat = Bërzhet 

Budaç 42° 33.5’N 19° 44’E 

Bunë (Bojana) 41° 51.5’N 19° 22’E 

Bun i Grebenit 42° 34’N 19° 42.5’E 

Bun i Markofçit 42° 29’N 20° 00’E 

Bun i Nikçit 42° 28.5’ N 19° 42’E 

Bun i Peraj 42° 21’N 19° 52.5’E (approximate) 

Bun i Thores not traced 

Bun i Trokus 42° 31’N 20°01’E 

Cardak i Kakinjes 42° 20.5’N 19° 53’E 

169

Çerek (Merturi) 42° 21.5’N 19° 55’E 

Çerem 42° 30’N 19° 55.5’E 

Cokaj (Kokaj) 42° 13.5’N 19°35.5’E 

Curaj i epër (Epërm) 42° 21’N 19° 55’E 

Dardhë 42° 12’N 20° 09.5’E 

Dedaj 42° 18’N 19° 32.5’E 

Dobrodol (Doderdol) 42° 33’N 20° 04’E 

Drabogijë (Dragobi)42° 26’N 19° 59’E 

Drinasa (Drin i Madh) 42° 02’N 19° 31.5’E 

Drisht 42° 07.5’N 19° 36’E 

Ducaj (Prekaj) 42° 20.5’N 19° 39’E 

Dushman 42° 09.5’N 19° 52.5’E 

Fushë Rudnicës (between Shala and Gussinje, at Jugoslav border) 

Gemsiqë (=Gomsiqe) 42° 02’N 19° 40.5’E 

Gjoani (Xhani near Kiri) approx. 42° 14’N 19° 42.5’E 

Gjonpepaj 42° 18.5’N 19° 55.5’E 

Gjuraj b. Plandë 42° 17’ N 19° 40’E 

Gomsiqë (river) = Gemshic 42° 01’N 19° 51.5’E 

Gryka Rapsh (Jugoslav (Crna Gora) border near Han i Grabonit) 

Gur i Kuq ( nw of Gur i Leks) 42° 11.5’N 19° 48’E (near Ndregijnaj, and Kiri) 

Han i Grabonit 42° 25.5’N 19° 31’E 

Han i Gropës 42° 32’N 19° 42’ N 

Han i Hot 42° 21’N 19° 26.5’E 

Han i Sakalit- Han i Spashit (near Kukes) 

Ibaljë (Jballje, Iballë) 42° 12’N 20° 00’E 

Jezerce 42° 26’N 19° 49’5’E 

Kakinje 42° 20.5’N 19° 50’E 

Kirit Cukalit 42° 08’N 19° 42.5’E 

Kçirë 42° 01.5’N 19° 54’E 

Kiri 42°14’N 19°42.5’E 

Kishaj 42° 12’N 20° 08.5’E 

Kolcegaj ( Kolgecaj)42° 21’N 19°26.5’E 

Kirit e Fushës (Kroiet e Fushës) 42° 10.5’N 19° 44’E 

Krumë 42° 11.5’N 20° 25’E 

Kurila 42° 17’N 19° 34’E 

Lesh (Lezhë) 41° 47’N 19° 38.5’E 

Lipushë (Lepushë) 42° 31.5’N 19° 44.5’E 

Ljesnice (? Jeshnicë) 42° 34’N 19° 39.5.’E 

Maja (Mali) e Berdashes (Bridashes) 42° 25’N 19° 39.5’E 

Maja e Bëris doles (Berizhdolit) 42° 30.5’N 19° 42.5’E 

Maja e Beshit 42° 23’N 19° 49’E 

Maja e Cukalit 42° 08.5’N 19° 45’E 

Maja e Dreles 42° 22’N 19° 54’E 

Maja e Derit (Drerit) 42° 15’N 19° 37.5’E 

Maja e Gurikulës 42° 32.5’N 19° 45.5’E (border peak) 

Maja e Hekuravë 42° 23’N 19° 58’E 

170

Maja e Kolatës 42° 29’N 19° 54.5’E (border peak) 

Maja e Leshnicës 42° 12’N 19° 52’E 

Maja e Marlulis (Marlulës) 42° 37.5’N 19° 42’E 

Maja e Metohijës 42° 34’N 19° 40’E 

Maja e Pajës 42° 29.5’N 19° 47.5’E (border peak) 

Maja e Shënikut 42° 28.5’E 19° 45’E 

Maja Strebiscit (Parun) approx. 42° 18’N 19° 37.5’E 

Majan (Mojan) 42° 37.5’N 19° 41’E (boundary peak) 

Mal i Rebës 42° 38’N 20° 11’E 

Medua (=San Giovanni de Medua = Shën Gjin) 41° 49’N 19° 35.5’E 

Mgulë 42° 13’N 19° 41’E 

Miliskau 42° 13’N 20° 20.06’E 

Müselini (Müselim, Myselim) 42° 06’N 19° 34’N 

Nicaj 42° 18.5’N 19° 48.5’E 

Nikç 42° 28.5’N 19° 42’E 

Ndrejaj 42° 24.5’N 19°46.5’E 

Nikçit = Bun i Nikçit 

Okol 42° 25’N 19° 46’E 

Palcë 42° 15.5’N 19° 54’E 

Parun approx. 42° 14’N 19° 38.5’E 

Pashtrik = Beshtriq 

Patkoje (Potkaj) not traced 

Peraj 42° 18’N 19° 55.5’E 

Plandë (Plani, Plan) 42° 18’ N 19° 43’E 

Pogaj 42° 10.5’N 20° 32’E 

Ponar 42° 18’N 20° 03.5’E 

Pront i Sath (Prroni i Thatë, stream) approx. 42° 19’N 19° 36.5’E 

Qaf e Bishkashit 42° 14’N 19° 37’E 

Qaf e Boshit 42° 18.5’N 19° 45.5’E 

Q. Borit = Q. e Valbones 

Qaf e Dershanes 42° 30’N 19° 55’E 

Qaf e Droçës 42° 24’N 19° 56’E 

Qaf Gur i Kuq approx. 42° 11’5’N 19° 48’E 

Qaf e Kirit 42° 13.5’N 19° 43’E (approximate) 

Qaf e Koprishit 42° 31’N 19° 43’E 

Qaf e Markofçes 42° 28.5’N 19° 20’E 

Qaf e Menules 42° 16.5’N 19° 26.5’E 

Qaf e Nermaja 42° 16.5’N 19° 48.5’E 

Qaj e Pejës 42° 26.5’N 19° 47’E 

Qaf e Shtogut 42° 18’N 19°40.5’E 

Qaf e Valbonës 42° 24’N 19° 49.5’E (approximate) 

Qaf e Vranices 42° 31’N 19° 59.5’E 

Quelzë (Qielzë) 42° 06’N 19° 54.5’E 

Rajë 42° 16.5’N 20°00’E 

Rapsh 42° 23.5’N 19° 30.5’N (approximate) (border region) 

171

Riolli.(Rjodhi) There are two localities of this name which Schütt could have visited. One 

is north west of Lezhë and Schütt made a few collections from the vicinity of Lezhë and 

Shën Gjin. The second is close to the river valley north east of Shkodra which is an area 

from which he made many collections so it is likely most collections from this locality 

are from Riolli(Shkodra) 

Riolli (Rrjoll, Rjodhi) (Shkodra) 42° 10’N 19° 12.5’E to 42° 13’N 19° 35.5’E 

Riolli (Rrjoll, Rjodhi) (Lezhë) 41° 51’N 19° 30.5’E 

Rosafa (Rozafa) Fortified hill in Shkodra 

Selcë 42° 30.5’N 19° 37’E 

Sermë 42° 13.5’N 19° 51.5’E 

Shala valley approx. 42° 18.5’N 19° 48.5’E 

Shën-Gjin (St. Giovanni di Medua) 41° 48’N 19° 36’E 

Shkelzen 42° 27.5’N 20° 08’E 

Skrobotusha 42° 35’N 19° 38.5’N 

Skutari (Skodra, Shkodér) 42° 04’N 19° 19°31’E 

Smutirog 42° 37’N 19° 31’E (approximate) 

Spash (Spas) 42° 11’N 20° 15’E 

Stan i Rebës 42° 29.5’N 20° 10’E 

Stan i Rupës 42° 28.5’N 20° 00’E 

Stare 42° 23’E 19° 27.5’E 

Tarabosh ambiguous; Schütt collected at two places with this name usually distinguishing 

them as follows, but rarely he does not; these are most likely Tarabosh in Parun 

Tarabosh (Skutari) 42° 03’ N 19° 27.5’E 

Tarabosh in Parun (Tarabosh i Bëzhetes) 42° 18’5’N 19° 36.5’E 

Thethi 42° 24’N 19° 44’E 

Toplanë 42° 13’N 19° 53.5’E 

Trojan 42° 33’N 19° 45.5’E (boundary peak) 

Trokus 42° 31’N 20°01’E 

Tropojë (Ljum Tropojës) 42° 24’N 20° 20.5’E 

Ura e Tamarës 42° 27.5’N 19° 34’E 

Ura e Mesit perhaps at 42° 08.5’N 19° 39.5’N (first river crossing above Kiri) 

Valbonë (Drin) tributary of the Drin; (Schütt seems to have used this for the area near the 

source of the river near the northern boundary) 

Van i Dejës (Vau i Dejës) 42° 01’N 19° 37.5’E 

Veleçik (near Rapsh) 42° 22’N 19° 32’E 

Velgjas 42° 18.5’N 19° 41’E 

Vermoshë 42° 35.5’N 19° 43.5’E to 42° 36’N 19° 41.5’E 

Vilzë 42° 07’N 19° 49.5’E 

Vlahën (near Bestriq) 42° 15’N 20° 28.5’E 

Vukaj, near Maja e Cukolit 42° 07.5’N 19° 43’E 

Vukli 42° 29.5’N 19° 39’E 

Zahrishtë 42° 12.5’N 20° 21.5’E 

Zogaj 42° 17.5’N 20° 21.5’E 

172

Sources of records for Albania - University of Toronto Mississauga

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