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Vol. 35 – 2009 - Ecologia Mediterranea - Université d'Avignon et des ...

Vol. 35 – 2009 - Ecologia Mediterranea - Université d'Avignon et des ...

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HANADI ISMAÏL, HUSSEIN ABOU-HAMDAN, AHMAD KOUBAYSSI, STÉPHANIE FAYOLLE, GABY KHALAF, ARLETTE CAZAUBON, JACQUES HAURY<br />

Figure 3 <strong>–</strong> Seasonal variations in the taxonomic richness in the stations according to different groups (the values are the sum of<br />

species for each macrophyte type of each season). Spring = April, May, June. Summer = July, August, September. Autumn<br />

= October, December. Winter = January, March.<br />

36<br />

the macrophytes with respect to some environmental<br />

param<strong>et</strong>ers. It also made it possible<br />

to establish macrophytic associations<br />

which exist in several sectors of Litani, and to<br />

distinguish the stations by their floristic characteristics.<br />

Figure 4 <strong>–</strong> Average percentage of macrophytic cover (average + SD) among<br />

the different stations (L1, L2, L3 & L4). (a) all taxa, (b) for each<br />

type of macrophytes categories at the four stations.<br />

On the map of the CCA (axis 1 inertia 41%;<br />

axis 2 inertia 32%) L1 is separated from L2-<br />

L3 and L4 on axis 1, and L2-L3 are separated<br />

from L4 on axis 2. In addition the axis 1 (41%<br />

inertia) <strong>des</strong>cribes a gradient of mineralisation<br />

and eutrophication on its negative part and a<br />

gradient of oxygenation on its positive part.<br />

Thus, the degree of pollution effect is strongly<br />

increased through the axis 1 which opposes a<br />

sector strongly polluted on the rhithral (L2,<br />

L3, L4) to a sector lightly polluted on L1.<br />

Thus, the ecological d<strong>et</strong>erminism is clear. In<br />

one hand, we can individualize the spring,<br />

characterized by its depth and its well oxygenated<br />

water and on the other hand, the stations<br />

of the rhithral characterized by a high<br />

conductivity, nitrates and phosphates concentrations,<br />

and by a relatively faster water.<br />

From there we distinguished 3 groups:<br />

<strong>–</strong> The group (G1) positively correlated with<br />

the depth. It is made up primarily by<br />

hydrophytes colonizing the stagnant mediums<br />

and/or with weak current (Chara sp.)<br />

and of the helophytes which correspond to<br />

the rhithral of the small rivers with a very<br />

good shining and a low flow (Phalaris arundinacea,<br />

Phragmites australis).<br />

<strong>–</strong> The group (G2) inclu<strong>des</strong> macrophytes generally<br />

announced in the rhithral of large<br />

rivers (here rhithral of Litani on L4) characterized<br />

by a strong representation of<br />

hydrophytes (Ranunculus fluitans, Myriophyllum<br />

spicatum, Potamog<strong>et</strong>on crispus,<br />

Potamog<strong>et</strong>on pectinatus, <strong>et</strong>c.).<br />

ecologia mediterranea <strong>–</strong> <strong>Vol</strong>. <strong>35</strong> <strong>–</strong> <strong>2009</strong>

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