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THÈSE - Université de Franche-Comté

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Fig. 3 shows the results of a principle component analysis carried out on the matrix “13<br />

farming practice variables” x “46 parcels”. The two first components represented here<br />

accounted for 49.8 % of the total inertia. Nitrogen fertilisation and mowing variables (yield<br />

and number of annual cuttings) were the variables most correlated to Axis 1 (Fig. 3). The first<br />

principal component was thus interpreted as a gradient of intensification. This gradient<br />

opposed two types of parcel management: one where fertilisation and mowing are the main<br />

features and the other grazing.<br />

A. terrestris dynamics in relation to farming practices and to T. europaea relative abundance<br />

Among all variables of practices and T. europaea relative abundance, only those correlated to<br />

an A. terrestris population dynamic variables at a risk lower than 0.15 were kept (Table 3).<br />

Figs 4a and 4b are based on a redundancy analysis and show the overall canonical relationship<br />

according to the two first axes between the table “farming practices and T. europaea relative<br />

abundance” and the table “A. terrestris population dynamic variables”. This relationship was<br />

found to be globally significant (P < 0.01 after 999 permutations). The canonical axes<br />

explained 18.7 % and 5.5 % of the variance of the A. terrestris population dynamics,<br />

respectively (total 24.2 %). Fig. 4a represents the maximum values for each parcel in the<br />

canonical space: they were clearly larger in the parcels where the relative abundance of T.<br />

europaea was higher at the beginning of the study period (spring 2002) and where grazing<br />

took place in the fall. Furthermore, the parcels for which the increase of A. terrestris<br />

populations began later in a commune were those with the lowest number of cuttings and<br />

addition of organic nitrogen fertiliser (Fig. 4b). The four variables (tesp2, autgraz, organ and<br />

cutting) account for less than 30 per cent of total variance, suggesting another explanatory<br />

variable also affected the A. terrestris population kinetics.<br />

THE HIGH DENSITY PHASE OF AN A. TERRESTRIS POPULATION CYCLE<br />

The A. terrestris relative abundance <strong>de</strong>creased with increasing grazing intensity (P < 0.001)<br />

and soil work intensity (P < 0.001).<br />

Influence of livestock grazing on A. terrestris relative abundance<br />

Significant differences were found between intensive pastures and permanent grassland and<br />

between intensive pastures and traditional pastures (Fig. 5a).<br />

Thèse C. Morilhat 2005 105

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