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Ecologie des foraminifères planctoniques du golfe de Gascogne ...

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tel-00480660, version 1 - 4 May 2010<br />

Below the surface mixed layer, Eastern North Atlantic Central Water (ENACW) is present<br />

down to 600 m water <strong>de</strong>pth. Between about 600 and 1300 m, Mediterranean Outflow Water<br />

(MOW) is characterised by relatively high salinity and low oxygen concentration (van Aken,<br />

2000). Northeast Atlantic Deep Water (NEADW) occurs from 1300 to 3000 m water <strong>de</strong>pth<br />

(e.g., Durrieu et al., 1999).<br />

The SW French continental shelf is characterized by freshwater input from the Giron<strong>de</strong> and<br />

Adour rivers (Fig.II-1). The Giron<strong>de</strong> has an average annual flux of 1100 m 3 s −1 with a seasonal<br />

maximum runoff of ~5000 m 3 s −1 in February, ~3250 m 3 s −1 in spring, and a minimum of<br />

~100 m 3 s −1 in summer (Schäfer et al., 2002). The Adour river has a mean discharge of<br />

350 m 3 s −1 (Brunet and Astin, 1999). The Adour and other rivers draining the Pyrenees are<br />

expected to have a high seasonal variability in runoff (Coynel et al., 2005). Peak flow may<br />

exceed 1000 m 3 s −1 <strong>du</strong>ring few days in winter and spring (Gil and Sanchez, 2000). In general,<br />

the fresh water plumes of both Giron<strong>de</strong> and Adour rivers flow northwards along the coast, but<br />

winds may force plumes to the south (Froi<strong>de</strong>fond et al., 1996, 2002; Puillat et al., 2004).<br />

Increased water discharge of the Giron<strong>de</strong> and Adour rivers may cause significant increase of<br />

nutrients triggering primary pro<strong>du</strong>ctivity in waters of the Bay of Biscay (Puillat et al., 2004;<br />

Kelly-Gerreyn et al., 2006a).<br />

In the Bay of Biscay, thermal stratification occurs between May and mid September, resulting<br />

in a pycnocline at about 50 m <strong>de</strong>pth, whereas the upper water column between January and<br />

the beginning of April is completely mixed (Puillat et al. 2004, and references therein).<br />

During mid September to December, progressive erosion of the thermocline is caused by<br />

wind-driven mixing of the surface ocean. Stratification as a result of river runoff and<br />

freshwater input peaks between March and June. In fall, stratification <strong>de</strong>creases because of<br />

<strong>de</strong>creased river discharge and increased wind driven mixing (Puillat et al. 2004).<br />

Chlorophyll-a concentration is highest in March, April, September and October in the Bay of<br />

Biscay (Fernan<strong>de</strong>z et al., 1991). During stratified periods, maximal chlorophyll-a<br />

concentration is recor<strong>de</strong>d at the bottom of the euphotic zone near the pycnocline (Labor<strong>de</strong> et<br />

al., 1999). During winter, wind-driven mixing causes <strong>de</strong>epening of the seasonal pycnocline,<br />

and nutrient concentrations in surface waters increase progressively (Tréguer et al., 1979).<br />

48

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