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Ecologie des foraminifères planctoniques du golfe de Gascogne ...

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tel-00480660, version 1 - 4 May 2010<br />

Geography - distribution Depth of life Species<br />

Surface, upper 60m (Schiebel et al 1997) 0-300m (Bé and Hamlin, 1967) ;<br />

Subpolar to tropical (Bé, 1977, EMIDAS). Typical species in transitionnal to polar,<br />

"Intermediate water" (Bé, 1977). They are found throughout the water column<br />

abundant in upwelling area (Hemleben et al., 1989). Waters rich in nutrients (Arabian<br />

above 400meters, but mainly in and above the thermocline. While it is not<br />

Sea) (Conan and Brummer, 2000). Abundant in Subartic and subantartic area where<br />

restricted to the photic zone in cooler waters where the non-spinose species are<br />

it represents + 50% of the assemblages (Pujol, 1980).<br />

most dominant. (Hemleben et al 1989)<br />

Tropical to transitionnal (Bé, 1977, EMIDAS) maximal <strong>de</strong>veloppemnt from 40 to 45°N<br />

(Pujol, 1980)<br />

Tropical to subtropical species, living in oligotrophic environment (Schmuker et<br />

Schiebel, 2002; EMIDAS); antycyclone eddies species (Ravula, 2004)<br />

Tropical to polar (EMIDAS). Subartic and subantarctic Provinces (Bé, 1977);<br />

becomes abundant in transitional and polar water masses (Hilbrecht, 1996); found in<br />

waters with temperatures higher than 21°C in the Arabian Sea and off the Great<br />

Barrier Reef (Brummer and Kroon, 1988; Darling et al., 2000), But in high abundance<br />

in the North Atlantic polar province of the Fram Strait, even un<strong>de</strong>r sea ice (Carsten et<br />

al., 1997). Their abundance would <strong>de</strong>crease from high latitu<strong><strong>de</strong>s</strong> to low ( Parker,<br />

1962; Vincent et Berger, 1981)<br />

Tropical to subpolar (EMIDAS). Subtropical province (Bé, 1977); In cyclonic eddies,<br />

in nutrients rich area (Kuroyanagi and Kawahata, 2004; Machain-Castillo et al 2008)<br />

Subtropical to polar (EMIDAS); subpolar (Bé and Tol<strong>de</strong>rlund,1971); Subartic (Pujol,<br />

1980); polar or subpolar (Schiebel and Hemleben, 2005)<br />

Tropical (Fairbanks et al 1982 ; Hemleben et al 1988).Tropical to temperate<br />

(EMIDAS); Subtropical province (Bé, 1977); abundance maximum around<br />

subtropicale zone, but very common in tropical area (Bé, 1977); arround 40°N in<br />

Iberic coast <strong>du</strong>ring summer 1962 (Bé and Hamlin, 1967); Signaled arround 49°59N<br />

12°W (Sime et al 2005).<br />

Tropical to subtropical (Pujol, 1980); no signaled beyond that iberic coast (Sime et al<br />

2005; Bé, 1977; Pujol, 1980)<br />

Tropical (Bé, 1977). Tropical to temperate (EMIDAS)<br />

Subtropical to transitionnal (Bé, 1977; EMIDAS)<br />

Transition zone, the only one really indigenous (Bé, 1977); temperate (EMIDAS);<br />

transitionnal to subpolar (Pujol, 1980) in oceanic front area (Rohling et al 1995);<br />

Stress habitat in the upper part of the main thermocline; preferred habitat = base of<br />

the seasonal thermocline (Cléroux et al., 2007) In oceanoic front area (Schiebel et<br />

al., 2002)<br />

Subpolar to transitionnal (Pujol, 1980); subpolar to tropical (EMIDAS). Temperate<br />

region (Hemleben et al, 1989); common in Californian current, but especially at the<br />

site 121km offshore (Ortiz et al, 1995)<br />

Subtropical in winter or transitionnal (Pujol, 1980); tropical to temperate (EMIDAS)<br />

Subtropicale d'hiver ou transitionnelle (Pujol, 1980); tropical to temperate (EMIDAS);<br />

Low abundance in South hemisphere (Parker, 1971)<br />

Subtropical to polar (EMIDAS), subartic-subantarctic Provinces; from Cap Htteras to<br />

Bay of Biscay (Bé,1977); max concentration between 45°-60°N (Pujol, 1980)<br />

Subtropical to polar (EMIDAS), alone shape to live in Arctic ocean (Bé, 1960) Artic-<br />

Antarctic Provinces (Bé,1977). Senestral more present in Arctic ocean, Labrador<br />

Sea, Greenland Sea and Norvegian Sea (Ericson, 1959); rare indivi<strong>du</strong>s beyond that<br />

40°N (Pujol, 1980)<br />

"Intermediate water" (Bé, 1977) surface to subsurface (EMIDAS), intermediate<br />

waters (Pujol, 1980)<br />

Life in surface (EMIDAS). Seem to be restricted to <strong>de</strong>eper water (Hemleben et<br />

al., 1989); 0-250m (Wilke and Peeters,in prep)<br />

Surface - subsurface (Bé, 1977, Bé and Tol<strong>de</strong>rlund, 1971, EMIDAS).<br />

Abundance max un<strong>de</strong>r 500m, often with highest frequency occasionnaly below<br />

the mixed layer, may be low sinking velocity (Schiebel et al, 2000). Life meanly<br />

in photic zone (Hemleben et al 1989). Live in surface (Pujol, 1980).<br />

"Intermediate water" (Bé, 1977) suface to subsurface (Hemleben et al., 1989;<br />

EMIDAS); 50-100m (Pujol, 1980). Shallow dwelling sp (Wilkes and Peeters, in<br />

prep)<br />

G. bulloi<strong><strong>de</strong>s</strong><br />

G. siphonifera<br />

G. calida<br />

T. quinqueloba<br />

G. glutinata<br />

Surface to subsurface (EMIDAS); profon<strong>de</strong> (Pujol, 1980) G. uvula<br />

Life in surface (Bé, 1977; Hemleben et al 1989; EMIDAS). Superficial waters<br />

(Jones, 1967); life upper 25m in equatorial Atl (Boltovskoy an Wright, 1976)<br />

G. ruber (white)<br />

Life in surface (Bé, 1977; Pujol, 1980; Hemleben et al., 1989) G. trilobus trilobus<br />

Life in surface (Bé, 1977; Hemleben et al., 1989, EMIDAS); surface to<br />

intermediate waters (Pujol, 1980)<br />

Subsurface to <strong>de</strong>ep (EMIDAS), repro<strong>du</strong>ction like G. truncatulinoi<strong><strong>de</strong>s</strong> in surface<br />

waters (Hemleben et al 1989)<br />

"Deep-Water" (Bé, 1977). Life in subsurface, up to 100m (Schiebel et al 2002) ;<br />

intermediate <strong>de</strong>ep water <strong>de</strong>pths (Wilke and Peeters, in prep.); When the Base of<br />

the summer thermocline is warmer than 16°C, T°C iso of Inflata indicate a<br />

<strong>de</strong>eper habitat, <strong>du</strong>ring winter, inflata lives in the mixed-layer (Cléroux et al.<br />

2007). "We found maximum concentrations of G. inflata within the thermocline,<br />

which is in agreement with observations by Fairbanks et al. (1982)" (Wilke et al.<br />

2006)<br />

"Deep-Water" Bé, 1977 ; mostly in subsurface (EMIDAS). Deep dweller (Erez<br />

and Honjo, 1981). Schiebel 1995 has found them from surface to 1000m (Bé,<br />

1960 ; Bé, 1977, Ortiz et al 1996). At Spring, would life close to surface and 'ld<br />

sink in <strong>de</strong>ep water(Hemleben et al, 1989); present (50-100m) and present in<br />

shallower water (Oberhansli et al, 1992); need not be adapted to a subsurface<br />

habitat is not entirely ruled out by our data (Ortiz et al 1996). G scitula occurs in<br />

surface water may have been the result of vertical transportation (Ottens, 1992).<br />

G.scitula is a <strong>de</strong>ep dwelling species that lives more abundantly than other<br />

species in the export zone and was found there in situ (Erez and Honjo, 1981;<br />

Hemleben et al., 1989).<br />

"Deep-Water" Bé, 1977; zone epipélagic from Nov to May, then un<strong>de</strong>rt more<br />

than 300m <strong>de</strong>pth (Pujol, 1980). Deep, surface <strong>du</strong>ring repro<strong>du</strong>ction (Hemleben et<br />

al., 1989; Schiebel et Hemleben, 2005; EMIDAS)<br />

"Deep-Water" Bé, 1977; zone epipélagic from Nov to May, then un<strong>de</strong>rt more<br />

than 300m <strong>de</strong>pth (Pujol, 1980); Deep, surface <strong>du</strong>ring repro<strong>du</strong>ction (Hemleben et<br />

al., 1989; Schiebel et Hemleben, 2005; EMIDAS)<br />

"Deep-Water" (Bé, 1982); max abondance from 100-400m but could go until<br />

1500m(Vergnaud-grazzini, 1973). His vertical distribution is related with<br />

pycnocline (Kuroyanagi and Kawahata, 2004); Surface to subsurface (EMIDAS)<br />

Arround 200m <strong>de</strong>pth (Bé, 1960), "Deep-Water" (Bé, 1982) ; surface to<br />

subsurface (EMIDAS)<br />

Thermocline dweller (Peeters, 2000; Fairbanks et al., 1982); Surface to<br />

subsurface (EMIDAS)<br />

G. trilobus sacculifer<br />

G. hirsuta<br />

G.inflata<br />

G. scitula<br />

G. truncatulinoi<strong><strong>de</strong>s</strong> senestre<br />

G. truncatulinoi<strong><strong>de</strong>s</strong> <strong>de</strong>xtre<br />

N. pachy<strong>de</strong>rma <strong>de</strong>xtre<br />

pachy<strong>de</strong>rma senestre<br />

P-D intergra<strong>de</strong>-<strong>du</strong>tertrei<br />

Tropical to subpolar (Hemleben et al., 1989), transitionnal (Pujol, 1980) "intermediate Water" (Bé, 1977); Surface (EMIDAS) O. universa<br />

Tropical to subtropical extremely rare on the sea bed (Parker and Berger, 1971),<br />

Subtropical, important in Atlantic W from Caraibean Sea to south of Québec, and in<br />

South Atlantic, between Brasil and Cap Town (Bé and Tol<strong>de</strong>rlund, 1971)<br />

Likes <strong>de</strong>ep waters as G.truncatulinoi<strong><strong>de</strong>s</strong> and crassaformis (Jones, 1967; Bé and<br />

Tol<strong>de</strong>rlund, 1971); consi<strong>de</strong>rate like intetrmediate species (Pujol, 1980); >60m<br />

(Schiebel et al 2002)). Between 50-100m (Bé, 1977); Surface to subsurface<br />

(EMIDAS)<br />

H. pelagica

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