Fauna of NZ 45 - Landcare Research

32 Kuschel (2003): Nemonychidae, Belidae, Brentidae (Insecta: Coleoptera)
ventrites combined, in female tapering to a slightly
upcurved apex and only briefly passing end of abdomen;
base sloping obliquely towards mesonotum; striae not or
hardly sulcate, deeply punctate; interstriae not costate;
inferolateral flange parallel to side margin as far as end of
striae 6 and 7 and ending there, not followed by an oblique
carina or flange. Hind wing as in Fig. 57, conspicuously
pigmented, dark, densely microtrichious, but folds pale,
lacking microtrichia; about 4x as long as wide, without anal
lobe, distinctly sinuous on hind margin just beyond middle;
apical field equal or subequal to venational area basad of
stigma; radial window distinct, small, triangular, with short
Rr; r-m distinct, inserted on Rr well basad of radial window,
transverse, slightly curved apicad in middle; m-cu strong,
straight; Cu strongly, irregularly sclerotised at junction with
m-cu; 3 anal veins present; hind margin ciliate, longest cilia
12 µm long.
Prosternum in male 3x as long as a coxal diameter, in
female 2x as long as a coxal diameter, area behind coxae in
both sexes half a coxal diameter long; sternopleural suture
distinct at least in part, well marked with puncta and short
transverse rugae; prosternal process between coxae narrow,
half the width of funicle. Femora weakly clavate,
pedunculate, not reaching end of abdomen in both sexes;
tibiae straight; fore tibiae with a broad, finely pubescent
grooming device between outer tooth and spur, lacking an
ascending comb on lower edge; spurs 1-2-2, strong; tibiae
and tarsi shiny, lacking a grey coating; tarsite 1 longer than
2 and 3 combined, tarsite 3 deeply bilobed, wider than 2;
onychium at apex ventrally subtruncate, not extending
below the claws, when not showing in dorsal view; claws
with with one or more fine, inconspicuous setae.
Male. Tergite 7 elongate, strongly convex across, with
rather small, entirely lateral pruinose patch; spiracle in a
pale patch. Tergite 8 in lateral view largely exposed beyond
7, in ventral view only briefly visible beyond ventrite 5.
Sternite 8 as in Fig. 100, transverse, undivided, not
emarginate at apex. Sternite 9 (Fig. 100) with long, rather
narrow arms and an elongated bladal sclerite with a slight
head at posterior end. Tegmen (Fig. 101) as long as aedeagus,
articulated on sides, ventral part with long anular arms,
dorsal part very long beyond connective membrane, with
long, diverging parameral lobes, these setose dorsally and
ventrally, with a pale median line beyond connective
membrane. Aedeagus as in Fig. 102, 103, pedon parallelsided,
tapering to a point, briefly upturned at apex; tectum
continuous with apodemal arms, extended to ostium;
apodemes shorter than aedeagal body, pedal arms articulated
with aedeagal body; internal sac twice as long as aedeagus,
with a flagelliform basal sclerite, this not quite as long as
apodemes, with a pair of median sclerites inside aedeagal
body.
Female. Tergite 7 elongate, compressed to some extent,
deep in lateral view; intersegmental membrane close to
base of tergite 8 with a pair of large, 2.7 mm long by 1.9
mm wide glands (Fig. 104), the glands applied with their
broadest face against inner walls of tergite, this face with a
distinct indentation where touching tracheae of spiracle 7.
Tergite 8 as in Fig. 104–105, with few, usually 5 long teeth,
with a strongly downcurved apex. Sternite 8 as in Fig. 106,
blade subtruncate, with about 20 minute setiferous teeth
on apical margin, with an apically split median arm extending
to a little beyond middle of blade; apodeme a little more
than twice as long as blade; membrane connecting with
sternite 7 (ventrite 5) with a large glandular pouch on either
side extending to near apex of apodemes. Genitalia as in
Fig. 108, hemisternites (Fig. 107) long, undivided, with
lateroapical stylus and a few setae; the well pigmented
genital sheath with a deeply coloured sclerite near base of
hemisternites; vagina about 4x as long as hemisternites,
more or less sacciform towards end, continuous with a
rather indistinct bursal part; spermatheca (Fig. 109) slender,
falciform, duct nearly as long as hemisternites and vagina
combined, pigmented and tapering as far as first bend,
widening before inserting at bursal end; gland distinctly
sclerotised, a little paler than spermathecal body; two long
cylindrical vaginal appendages present; colleterial gland
absent.
Distribution. Endemic to New Zealand.
Host plants. Woodborer in various dying or freshly dead
trees.
Remarks. One species varying greatly in size, some males
exceeding the length of any other brentine in the world.
The genus is currently placed in the tribe Ithystenini. It is
only remotely related to Ithystenus Pascoe of Australia and
Vanuatu, Mesetia Blackburn of Australia, and Bulbogaster
Lacordaire of Fiji, but may be a little more closely related
to Prodector Pascoe of Sulawesi.
Lasiorhynchus appears to be unique in many ways to
all other Brentinae, in particular in having:
(1) a distinct, normal, well-exposed scutellum instead
of an entirely concealed one;
(2) an elytral base that slopes down to the mesonotum
instead of falling vertically;
(3) no oblique carina or flange (the ‘submarginal ridge’
of Lyal & King, 1998) beyond the inferolateral flange on
the underside of the elytral declivity;
(4) no protruding apical lobe on the ventral surface of
the claw segment instead of a distinct one amply visible in
dorsal view between the claws;