catalogue of the continental mollusks of russia and - Ruthenica ...

CATALOGUE OF THE CONTINENTAL MOLLUSKS OF RUSSIA AND 

ADJACENT TERRITORIES 

YURI I. KANTOR, MAXIM V. VINARSKI, ANATOLY A. SCHILEYKO, 

ALEXANDER V. SYSOEV 

VERSION 2.3 

(published online on December 22, 2009) 1 

The current version 2.3 contains 2065 included taxa comparing to 2032 in the previous version 

2.2 (published June 9, 2009). 

Introduction to the version 1.0 

(published online on February 15, 2009) 

Inventories of molluscs of Russia (and adjacent territories) is currently becoming a priority, 

especially in last decade. Several important publications appeared recently, among others the identification 

guide to freshwater molluscs of Russia [Starobogatov et al., 2004], annotated list of the molluscs 

of Baikal Lake [Sitnikova et al., 2004; Slugina, Starobogatov, 2004], monograph of Lymnaeidae 

of Europe and northern Asia [Kruglov, 2005], as well as the first complete catalogue (although 

in abridged form and only in Russian) of all groups of Mollusca of Russia and adjacent countries 

(within the borders of former USSR) [Kantor, Sysoev, 2005]. The catalogue was compiled mostly 

from published data. Only for some groups were the data based on collections, mostly that of ZIN, 

which possesses the largest collection of molluscs from the territory of the former USSR. 

Later the illustrated catalogue of marine and brackish-water Gastropoda was published [Kantor, 

Sysoev, 2006]. It included 1240 species, being approximately 1/3 of entire fauna of molluscs. 

The catalogue included as a rule photographs of the type specimens for the species, described from 

Russia and adjacent territories. For the species, described from elsewhere, mostly the specimens 

were used for illustrations on which the record was based, that were identified by the authors in the 

corresponding publications. 

The full version of the catalogue, which includes the data on type localities and type specimens 

of each species for the remaining groups of molluscs has been in preparation since. 

After the publication of illustrated catalogue of gastropods it became obvious that such publications 

turmed to be outdated rather fast. During two years since it was completed, several important 

monographs and revisions were published, several are in preparation. 

The situation with the continental molluscs is even more complicated. Firstly, the Russian specialists 

on fresh-water molluscs appeared to be “splitters” comparing with their western colleagues. 

This sometimes leads to extreme differences in opinion on species number and taxonomy of some 

groups. For example, Graf [2007] recently analyzed the number of species of Unionoidea of 

Palearctic recognised by Russian malacologists (156 species in 34 genera) and western specialists 

(45 species in 16 genera). Most of the groups are still awaiting the critical re-assessment. 

1 Version 1.0 was published online on February 15, 2009 

version 2.0 was published online on March 23, 2009 

version 2.1 was published online on April 27, 2009 

version 2.2 was published online on June 9, 2009

In this situation I decided to publish the available to us materials on continental (fresh-water 

and terrestrial) molluscs of Russia and adjacent territories within the borders of former USSR on the 

Internet in order to make the results available to broad audience of specialists in Russia and abroad. 

Publication in this format will allow rapid corrections and additions of new materials in future. I invite 

all the specialists to contribute their experience and knowledge to build most comprehensive 

and updated inventory of continental molluscs. 

This work has been accomplished besides Yu. Kantor by A.A. Schileyko (both from 

A.N.Severtzov Institute of Ecology and Evolution of Russian Academy of Sciences) and 

A.V.Sysoev (Zoological Museum of Moscow State University). Invaluable help was provided by 

M.V. Vinarski (Omsk State Pedagogical University). 

In compiling the work all available sources were used: monographs, separate papers, as well as 

species lists and catalogues. Moreover, the original descriptions of most of the species were 

checked. This was possible through the access to several large libraries in Russia and abroad (principally 

the libraries of Muséum national d’histoire naturelle, Paris and National Museum of Natural 

History, Smithsonian Institution, Washington D.C.). As a result hundreds of mistakes in published 

sources, including the nomenclature of many species were corrected. Unfortunately, we were not 

able to find references for a few species. Still type material for many species was not traced. The 

basis of the list of terrestrial species was the unpublished manuscript of Likharev and Schileyko. It 

represented the second updated edition of the famous monograph of I.M. Likharev and E.S. Rammelmeyer 

[1952], which was supposed to be published in late 1980-s but due to long period of economic 

instability was never printed. 

In current database all terrestrial and freshwater molluscs are included. Frackish-water species 

of Gastropoda were treated in Kantor and Sysoev [2006] and therefore are not included herein. On 

the contrary, brackish water bivalves (including those of the Caspian Sea, which may be considered 

as marine) are treated in the current catalogue. 

In species entry the main habitat is included: F – stands for fresh-water species, B – for brackish-water, 

M – for marine, and T – for terrestrial species. 

Species within genera and genera within families are arranged in alphabetical order. Subgenera 

are not included. 

On the types of Ya. I. Starobogatov 

Mention should be made of the types of some species, described by Ya. I. Starobogatov (1932- 

2004), especially in co-authorship with B. M. Logvinenko from the Caspian Sea. These authors 

wrote the molluscan section in the Atlas of invertebrates of the Caspian Sea (Moscow, 1968). Logvinenko 

and Starobogatov did not mention holotypes in the text (or type localities), but specified 

them in the figure captions. Although there is no information about the depository of the types, they 

should be stored in ZIN, where Ya. I. Starobogatov worked most of his life. Nevertheless, we were 

not able to find many of the types in ZIN collection. According to personal communication of L. L. 

Yarohnovich (collection manager of fresh-water and land molluscs of ZIN), part of the collection of 

Caspian molluscs, including types of new species, was stored in the Biological Faculty of Moscow 

State University in the laboratory of B. M. Logvinenko. After his death the material was transferred

to ZIN and was kept in the personal office of Starobogatov for many years. After Starobogatov’s 

death all the numerous samples were transferred to the general molluscan storage, but until now not 

completely processed and therefore not included into type and general collections. Thus it is possible 

that many of the types that are listed in this publication as “not traced” may be available in future. 

On the inventory system in Zoological Institute of RAS 

A significant number of the types are stored in ZIN. Therefore we should briefly mention the 

system of inventory numbers used in this collection. 

Starting in the late XIX century there was a card systematic catalogue in molluscs collections 

of ZIN (probably started by S. M. Herzenstein and N. M. Knipowich). For each species a separate 

card (with several entries) was started and the locality and the author of identification were cited for 

each sample. Thus, each sample was given a number which was unique only within single species. 

In this publication we refer to such numbers in the systematic catalogue as “ZIN No. ** in the systematic 

catalogue”. Usually the card started with the type specimens and therefore most of them 

have No. 1. In the collections the lots are arranged taxonomically in accordance with the systematic 

catalogue numbers. 

Beside the systematic catalogue, in the 1960’s a system of inventory numbers was started. The 

numbers were attributed to the lots not in taxonomic order. The inventory numbers cover most of the 

samples of marine gastropods and only partially those of fresh-water and land. In this book we provide 

the inventory numbers (if available), as well as systematic catalogue numbers. The number before 

the slash is the inventory number, while that following is the number in the systematic catalogue. 

Quantitative estimate of the fauna of continental molluscs of Russia 

At present 1892 continental species are included in Version 1.0 of the catalogue. We are following 

the current number of species in Russian literature (see above about the “splitting”). 

Acknowledgements 

Many colleagues assisted us in compiling this list. We want to thank them all, but especially: 

B. I. Sirenko, P. V. Kiyashko, L. L. Yarohnovich, R. Kormushkina from ZIN; T. Ya. Sitnikova from 

Limnological Institute of RAS; V.V. Anistratenko (Institute of Zoology of National Academy of 

Sciences of Ukraine); L. A. Prozorova (Biological-Pedological Institute of Farf-Eastern Branch of 

RAS; prof. P. Bouchet, J.-P. Rocroi, V. Héros from the Muséum national d’histoire naturelle (Paris); 

J. D. Taylor, K. Way, and D. G. Reid from the Natural History Museum (London); R. Heshler and 

M. G. Harasewych (National Museum of Natural History, Smithsonian Institution, Washington 

D.C.); A. Warén (Swedish Museum of Natural History, Stokholm); M. Glaubrecht (Zoological Museum, 

Humboldt University, Berlin)

Graf D. L. 2007. Palearctic freshwater mussel (Mollusca: Bivalvia: Unionoida) diversity and the 

Comparatory Method as a species concept. Proceedings of the Academy of Natural Sciences 

of Philadelphia, 156: 71-88. 

Introduction to the version 2.0 

(published online on March 23, 2009) 

Yuri I. KANTOR 

February 15, 2009 

The new version is siginificantly corrected and updated for the fresh-water molluscs. We highlighted 

all alterations in blue. From now and on in every new version the corrections will be highlighted 

similarly, while the text of the previous version will be in black. 

The other major revision is the inclusion of the Caspian gastropods in the current version. 

They were treated in Kantor and Sysoev (2006) illustrated catalogue of Russian Marine and brackish-water 

Gastropoda. Nevertheless since the on-line catalogue includes Caspian bivalves we decided 

to include gastropods as well for the convenience of the readers. 

This brought the total number of included taxa to 2026 comparing to 1892 continental species 

included in Version 1.0. 

In addition to the acknowledgements below we would like to express our thanks to colleagues 

who assisted us in improving version 1.0 of the Catalogue: 

Peter Glöer (Hetlingen, Germany) 

Alexei V. Chernyshev (Vladivostok, Russia) 

Alexander A. Frolov (Murmansk, Russia) 

Ludvig Dorodnitsyn (Moscow, Russia). 

YURI I. KANTOR, MAXIM V. VINARSKI 

March 23, 2009

Сокращения названий музеев и институтов 

[Abbreviations of the museums and institutions] 

ANSP — Academy of Natural Sciences, Philadelphia, USA 

BMNH — Natural History Museum, London, UK 

CASIZ — California Academy of Sciences, San Francisco, USA 

IMT — Institute of Malacology, Tokyo, Japan 

IORAN — Institute of Oceanology of Russian Academy of Sciences, Moscow, Russia 

IRSNB — Institut Royal des Sciences Naturelles de Belgique 

IZU – Institute of Zoology of the National Academy of Sciences, Kiev, Ukraine 

LSL — Linnean Society, London, UK 

MA — private collection of Masao Azuma, Takarazuka, Hyõgo prefecture, Japan 

MCZ — Museum of Comparative Zoology, Harvard University, Cambridge, USA 

MHNG — Muséum d’Histoire Naturelle, Genève, Switzerland 

MNHN — Muséum national d’histoire naturelle, Paris, France 

NC — Nishinomiya Shell Museum, Hyõgo prefecture, Japan 

NMBE — Naturhistorisches Museum der Burgergemeinde Bern, Switzerland 

NMC — National Museum of Canada 

NMG – Naturhistoriska Museet Göteborg, Sweeden 

NHMB – Naturhistorisches Museum, Basel, Switzerland 

NHMV — Naturhistorisches Museum, Vienna, Austria 

NMV — National Museum of Victoria, Australia 

NMWZ — National Museum of Wales, Cardiff, UK 

NSMT — National Science Museum, Tokyo, Japan 

RAMM — Royal Albert Memorial Museum, Exeter, UK 

RBINS – Royal Belgian Institute of Natural Sciences, Brussels 

SMF — Natur-Museum Senckenberg, Frankfurt am Main, Germany 

SMNH — Swedish Museum of Natural History, Stokholm, Sweeden 

SSM — Sea and Shell Museum, Rikuzen-Takada, Iwate prefecture, Japan 

USMP — Museum of Paleontology, University of California, Berkeley, USA 

UMT — University Museum, University of Tokyo, Japan 

USNM — National Museum of Natural History, Smithsonian Institution, Washington D.C., USA 

UUZM — Uppsala University Museum, Uppsala, Sweeden 

YPM — Peabody Museum of Natural History, Yale University, New Haven, USA 

ZIN — Zoological Institute of Russian Academy of Sciences, St.-Petersburg, Russia 

ZMB — Zoologisches Museum, Museum für Naturkunde der Humboldt-Universitat, Berlin Germany 

ZMBN — Zoologisk Museum, University of Bergen, Norway 

ZMC — Zoologisk Museum, University of Copenhagen, Denmark 

ZMMU — Zoological Museum of Moscow University, Russia 

ZMUC — Zoological Museum of the University of Copenhagen, Denmark 

ZMUO — Zoologiske Museum, Universitetet i Oslo, Oslo, Norway 

ZMZ – Zoologisches Museum Zürich, Switzerland

GASTROPODA 

SYSTEMATIC LIST 

Clade NERITIMORPHA [= Neritopsina] 

HELICINOIDEA Férussac, 1822 

HELICINIDAE Férussac, 1822 

(Kantor Yu. I., Schileyko A.A., Likharev I.M., Sysoev A.V.) 

Hemipoma A. Wagner, 1905 

Type species: Helicina hakodadiense Hartman, 1890 (SD Wenz, 1938) 

Hemipoma hakodadiense (Hartman, 1890) T 

[= Hemipoma sadoense A. Wagner, 1905] 

Helicina hakodadiense Hartman, 1890: 286, pl. 3, fig. 8. 

● Type locality: «Hakodadi, Japan» (Hakodate Prefecture, Hokkaido). 

● Types: unknown. 

● Distribution: Itirup and Irup islands (Kurile Islands) (Likharev, Schileyko, MS). 

NERITOIDEA Rafinesque, 1815 

NERITIDAE Rafinesque, 1815 

(Kantor Yu. I., Vinarski M.V.) 

Theodoxus Montfort, 1810 

Type species: Theodoxus lutetianus Monfort, 1810 (= Nerita fluviatilis Linnaeus, 1758) (OD) 

● Remark: Shadin (1952) and others recorded Theodoxus transversalis (C. Pfeiffer, 1828) for 

Ukraine. Nevertheless, the area of this species is probably restricted to the middle Danube and there 

are no specimens collected on the territory of former USSR (Anistratenko et al., 1999). 

Theodoxus astrachanicus Starobogatov in Starobogatov, Filchakov, Antonova et Pirogov, 1994 F 

Starobogatov et al., 1994: 8-9, fig. 1 (1, 2). 

● Type locality: southern end of Barskaya Kosa, 1 m (Volga Delta). 

● Holotype: ZIN. 

● Distribution: basin of the Azov Sea and Volga delta, in rivers (Anistratenko, 1998b; Starobogatov et al., 

2004). 

Theodoxus danasteri (Lindholm, 1908) F 

Neritina danubialis var. danasteri Lindholm, 1908: 214-215. 

● Type locality: "Fluss Dnjestr bei Rybnitzy, Gvt. Podolien". 

● Types: whereabouts unknown. 

● Distribution: northern Black Sea maritime area, rivers (Anistratenko, 1998b; Anistratenko et al., 1999; 

Starobogatov et al., 2004). 

● Remark: some records of Th. danubialis (C. Pfeiffer, 1828) in lower Danube and its mouth probably (eg. 

Polischuk, 1974) probably are incorrectly identified specimens of Th. danasteri (Anistratenko, 1998b; Anistratenko 

et al., 1999).

Theodoxus euxinus (Clessin, 1886) F, B 

Neritina euxina Clessin, 1886: 55. 

● Type locality: "Mangalia, au der Küste des schwarzen Meeres, nördlich von Varna". 

● Types: not traced. 

● Distribution: Black Sea maritime area, lower parts of rivers (Anistratenko, 1998b; Anistratenko et al., 1999; 


Theodoxus fluviatilis (Linnaeus, 1758) F 

[= Neritina brauneri Lindholm, 1908; Neritina brauneri f. lacrymans Lindholm, 1908; Neritina brauneri f. 

alboguttata Lindholm, 1908; Neritina brauneri f. pulcherrima Lindholm, 1908; Theodoxus dniestroviensis 

Put, 1972] 

Nerita fluviatilis Linnaeus, 1758: 777. 

● Type locality: Europe. 

● Lectotype (Anistratenko et al., 1999): LSL. 566 

● Distribution: rivers of Azov-Black seas and Baltic Sea basins (Anistratenko, 1998b; Anistratenko et al., 

1999; Starobogatov et al., 2004). 

Theodoxus milachevitchi Golikov et Starobogatov, 1966 M 

Golikov, Starobogatov, 1966: 353, fig. 1 (1a, 1b). 

● Type locality: phaseoline silt, off Crimea coast, Black Sea, 44º47’15”N, 35º50’20”E, 71 fathoms. 

● Holotype: ZIN 1/6134. 

● Distribution: subfossil on phaseoline silt, off Crimea coast, Black Sea, 20-60 m (Golikov, Starobogatov, 

1966). 

Theodoxus pallasi Lindholm, 1924 F, B 

[= Theodoxus schirazensis var. major Issel, 1865] 

Lindholm, 1924c: 34 (nom. nov. pro Neritina liturata Eichwald, 1838 non Schultze, 1826). 

● Type locality: Daghestan. 

● Lectotype (Starobogatov et al., 1994): ZIN 54547/63. 

● Distribution: Caspian and Aral seas; 0-35 m (Logvinenko, Starobogatov, 1968; Starobogatov, 1974), Armenia 

(Akramowski, 1976), Ural River (Pirogov et al., 1994). 

Theodoxus sarmaticus (Lindholm, 1901) F, B 

Neritina fluviatilis var. sarmatica Lindholm, 1901: 181-182. 

● Type locality: "im Oskol, in stilleren Buchten. Namentlich häufig in einer seeartigen Bucht am rechten Ufer 

des Flusses bei Novyi Oskol, ... Mühlendamm bei Golubino". 

● Syntypes: ZIN 6098/23, 6099/24, 6100/25. 

● Distribution: rivers of the northern coast of Black and Azov seas (Anistratenko, 1998b; Anistratenko et al., 


Theodoxus schultzii (Grimm, 1877) B, M 

Neritina schultzi Grimm, 1877: 77-78, pl. 7, fig. 5, pl. 8, fig. 16. 

● Type locality: two localities in the Caspian Sea: 43º17' N, 01º03' E, 40 fathoms, and 42º48' N, 01º22' E, 48 

fathoms. 

● Syntypes: ZIN, No. 5 in systematic catalogue. 

● Distribution: middle and southern Caspian Sea; 15-100 m (Logvinenko, Starobogatov, 1968). 

Theodoxus subthermalis (Bourguignat in Issel, 1865) F 

Theodoxus fluviatilis var. subthermalis Bourguignat – Issel, 1865: 22-23. 

● Type locality: not stated (Lac de Paleostomi, près de Poti, Georgie – locality of the probable syntype). 

● Probvable syntype: MHNG, Bourguignat collection. 

● Distribution: rivers and lakes of the eastern coast of Black and Azov seas (Anistratenko, 1998b; Anistratenko 

et al., 1999; Starobogatov et al., 2004).

Theodoxus velox V. Anistratenko in O. Anistratenko, Starobogatov et V. Anistratenko, 1999 F 

Theodoxus velox V. Anistratenko – O. Anistratenko, Starobogatov et V. Anistratenko, 1999: 17-18, fig. 4, 7. 

● Type locality: Dnieper delta, Zbyrjevskji liman. 

● Types: Institute of Zoology, Kiev. 

● Distribution: type locality, Kahovskoye reservoir (Anistratenko et al., 1999; V. Anistratenko, pers. comm.). 

● Remark: Zettler (2008) considers Th. velox to be a junior synonym of Th. fluviatilis. The close relationships 

between these species have been revealed by means of molecular genetical studies (Bunje & Lindberg, 2007). 

Clade CAENOGASTROPODA 

Informal group ARCHITAENIOGLOSSA 

CYCLOPHOROIDEA Gray, 1847 

COCHLOSTOMATIDAE Kobelt, 1902 


Toffolettia Giusti, 1971 

Type species: Cyclostoma striolatum Porro, 1840 (OD) 

Toffolettia lederi (O. Boettger, 1881) T 

Pomatias lederi Boettger O., 1881a: 244, pl. 9, fig. 22. 

● Type locality: "Kutais in Mingrelien" (Kutaisi, Georgia). 

● Lectotype (Zilch, 1958): SMF No. 158975. 

● Distribution: wet deciduous forests of northern branches of western (Dakhovskoie ravine) and eastern 

(Military-Osetin Road, Kurtatinskoe ravine) of Great Caucasus; southern branches of Great Caucasus from 

Fischt Mountain in the west to Rachinsky Ridge in the east (Likharev, Schileyko, MS); vicinities of Matzesta, 

Adler and Sochi (Egorov, Greke, 2003). 

CYCLOPHORIDAE Gray, 1847 


Caspicyclotus Forcart, 1935 

Type species: Cyclotus sieversi L. Pfeiffer, 1871 (monotypy) 

Caspicyclotus sieversi (L. Pfeiffer, 1871) T 

[= Cyclotus bourguignati Doumet-Adanson, 1885; Cyclotus herzi O. Boettger, 1889; Valvata brandti Westerlund, 

1897, part.; Cyclotus biggsi Schlesch, 1934; Caspicyclotus praesieversi Steklov, 1966] 

Cyclotus sieversi Pfeiffer, 1871: 69-70. 

● Type locality: «Caucasia meridionali» (Lenkoran, Azerbaijan). 


● Distribution: Eastern Transcaucasia: forests of Lenkoran lowland and Talysh, southern branches of Great 

Caucasus from Lagodekhi to Nukhi (Likharev, Schileyko, MS); one record near Mtzkheta (Rosen, 1925). 

ACICULIDAE Gray, 1850 


Acicula Hartmann, 1821

Type species: Bulimus lineatus Draparnaud, 1821 (monotypy) 

Acicula limbata Reuss, 1860 T 

Reuss, 1860: 61. 

● Type locality: «Tuchoritz und Lipen» (Czechia). 


● Distribution: Georgia: Poti, Surami, Manglisi (Boeters et al., 1989). 

Acicula moussoni O. Boettger, 1879 T 

Boettger, 1879a: 41, pl. 1, fig. 7 

● Type locality: Kazbek (Caucasus). 

● Lectotype (Boeters et al., 1989): SMF No. 4202a (not No. 4056 as designated by Zilch, 1976b – see 

Boeters et al., 1989). 

● Distribution: several records at northern and southern branches of Great Caucasus, Suram and Adzhar- 

Trialet ridges (Likharev, Schileyko, MS). 

Acicula parcelineata (Clessin, 1911) T 

[= Acme carpatica A. Wagner, 1912; Acme parcelineata var. cylindracea Sitsch, 1925; Pupula wagneri 

Hesse, 1930, nom. nud.; Acicula bakanense Steklov, 1966] 

Acme parcelineata Clessin, 1911: 165. 

● Type locality: «Dzingelau, Oestr. Schlesien» (Dziegielow near Cieszyn, Poland). 

● Syntypes: lost (Zilch, 1976b). 

● Distribution: Ukraine: Carpathians (Likharev, Schileyko, MS), Podolsk Hills (including vicinities of Lvov) 

(Sverlova, 2006). 

Platyla Moquin-Tandon, 1855 

Type species: Acme dupuyi Paladilhe, 1868 (SD Boeters, Gittenberger et Subai, 1989) 

Platyla jankowskiana (Jackiewicz, 1979) T 

Acicula jankowskiana Jackiewicz, 1979: 95, fig. 1. 

● Type locality: valley of Zhencha river, near Tatarov village, Ivano-Frankovsk Region, Ukraine. 

● Holotype: IZP. 

● Distribution: type locality. 

Platyla oedogyra (Paladilhe, 1868) T 

[= Acme similis Reinhardt, 1880; Acme perpusuilla Reinhardt, 1880] 

Acme ? var. oedogyra Paladilhe, 1868: 236, pl. 13, figs. 4-6. 

● Type locality: «Kieco». 

● Lectotype (Boeters et al., 1989): Paladilhe collection, Faculté des Sciences, Montpellier, France. 

● Distribution: Ukraine: Lvov and Zhitomir regions (Anistratenko, 1998b). 

Platyla perpusilla (Reinhardt, 1880) T 

Acme perpusilla Reinhardt, 1880: 46 

● Type locality: "beim Herkulesbade auf dem Wege nach dem Domogled zwischen abgefallenem Buchenlaub" 

[Romania]. 

● Lectotype (Zilch, 1976b): SMF No. 4058. 

● Distribution: Carpathian Biospheric Reserve (Zakarpatskaya Region, Rakhov District, "Kuzii" tract) (Sverlova, 

2006). 

Platyla polita (Hartmann, 1840) T 

[= Carychium lineatum C. Pfeiffer, 1828 non Bulimus lineatus Draparnaud, 1801, nec Carychium lineatum 

A.E.J. Férussac, 1821; Cyclostoma lubricum Held, 1847; Truncatella lubrica Held, 1847; Acme subpolita 

Gottschick, 1921]

Pupila acicularis polita Hartmann, 1840: 5, pl. 2, figs. 1, 2. 

● Type locality: "Gipfel des Schönebergs bei Hof Geismar in Hessen" (Schoneberg Mountains, Hessen, 

Germany). 


● Distribution: Kaliningrad Region, Baltic countries, vicinities of St.-Petersburg (Glint), Valdai, Moscow, 

and Central-Russian Hills, western regions of Byelorussia (Belovezhskaya Pushcha, Polesye); Ukraine – 

Carpathians, Volyno-Podolsk Hills, vicinities of Kiev, Kanevsly Natural Reserve (Kornyushin, 1988), 

Ukrainian Polesye (Zhitomir, Chernigov, Sumy regions), and in area between Prut and Dniester rivers 


DIPLOMMATINIDAE L. Pfeiffer, 1857 


Palaina Semper, 1865 

Type species: Diplommatina macgillivrayi L. Pfeiffer, 1854 (SD Iredale, 1944) 

Palaina amurensis (Mousson, 1887) T 

Diplommatina amurensis Mousson, 1887: 23-24, pl. 1, fig. 7. 

● Type locality: vicinities of Vladivostok. 

● Syntypes: ZMZ No. 526628; SMF No. 105154/2. 

● Distribution: Southern Primorye Territory (Ussurijsky and "Kedrovaya Pad'" natural reserves, vicinities of 

Vladivostok) (Likharev, Schileyko, MS). 

VIVIPAROIDEA Gray, 1847 

VIVIPARIDAE Gray, 1847 

(Kantor Yu.I., Vinarski M.V., Sysoev A.V.) 

Amuropaludina Moskvicheva, 1979 

Type species: Paludina praerosa Gerstfeldt, 1859 (OD) 

Amuropaludina chloantha (Bourguignat, 1860) F 

Vivipara chloantha Bourguignat, 1860a: 534, pl. 24, figs. 5-7. 

● Type locality: "divers affluents de l'Amour moyen". 

● Types: possibly, in MNHN or MHNG, Bourguignat collection. 

● Distribution: Amur basin, excluding upper part of Zeya basin; in rivers and running lakes (Bogatov, 

Zatrawkin, 1990 [1992]; Starobogatov et al., 2004). 

Amuropaludina pachya (Bourguignat, 1860) F 

Vivipara pachya Bourguignat, 1860a: 532-533, pl. 24, figs. 1-2. 

● Type locality: "le flueve Amour". 


● Distribution: central and lower Amur basin, in rivers and running lakes (Bogatov, Zatrawkin, 1990 [1992]; 


Amuropaludina praerosa (Gerstfeldt, 1859) F 

Paludina praerosa Gerstfeldt, 1859: 509-510, pl. 9, figs. 6a, 7. 

● Type locality: middle Amur River. 

● Lectotype (Moskvicheva, 1979): ZIN, № 205 in systematic catalogue. 

● Distribution: Amur basin, excluding upper part of Zeya basin and Amgun basin, in rivers and running lakes

(Bogatov, Zatrawkin, 1990 [1992]; Starobogatov et al., 2004). 

Cipangopaludina Hannibal, 1912 

Type species: Paludina malleata Reeve, 1863 (OD) 

Cipangopaludina kurilensis Starobogatov in Moskvicheva, 1979 F 

[= Viviparus malleatus sensu auct., non Paludina malleata Reeve, 1863] 

Starobogatov in Moskvicheva, 1979: 91. 

● Type locality: Kunashir Island [Kurile Islands]. 

● Lectotype (Bogatov, Zatrawkin, 1990 [1992]): ZIN No. 1 in systematic catalogue. 

● Distribution: southern Kurile Islands, southern and central Sakhalin, in lakes and slow-running rivers (Prozorova, 

1996[1997]a; Prozorova, 1998a; Starobogatov et al., 2004). 

Cipangopaludina sujfunensis Moskvicheva, 1979 F 

Cipangopaludina sujfunensis Moskvicheva, 1979: 89-90, fig. 3. 

● Type locality: Lake Soldatskoye near Ussuri town (basin of Razdolnaya River). 

● Holotype: ZIN No. 1 in systematic catalogue. 

● Distribution: basins of the rivers of southern Primorje, except Amur basin, in lakes (Moskvicheva, 1979; 


Cipangopaludina ussuriensis (Gerstfeldt, 1859) F 

Paludina ussuriensis Gerstfeldt, 1859: 507-508, pl. 9, figs. 1-4. 

● Type locality: "In den Seen, die mit dem mittleren und unteren Amur in Verbindung stehen, namentlich 

häufig afer in den Lachen au der Ussuri-Mündung". 

● Lectotype (Bogatov, Zatrawkin, 1990 [1992]): ZIN No. 135 in systematic catalogue. 

● Distribution: Amur basin, excluding upper and central parts of Zeya and Bureya basins, in lakes (Starobogatov 

et al., 2004). 

Cipangopaludina zejaensis Moskvicheva, 1979 F 

Cipangopaludina zejaensis Moskvicheva, 1979: 89, fig. 2. 

● Type locality: Lake Orlinoye, 120 km from mouth of Arga River (upper Zeya basin). 


● Distribution: upper and central parts of Zeya and Bureya basins, middle Amur and Ussuri basins, in lakes 

(Moskvicheva, 1979; Starobogatov et al., 2004). 

Contectiana Bourguignat, 1880 

Type species: Cyclostoma contectum Millet, 1813 (fixation of the types species not traced) 

Contectiana contecta (Millet, 1813) F 

[= Nerita fasciata Müller, 1774 (part.); Paludina crystallina Gray, 1821; Paludina inflata Villa et Villa, 

1841; Vivipara lacustris Beck, 1847] 

Cyclostoma contectum Millet, 1813: 5-7. 

● Type locality: "La Mayenne, etc. A Angers: les fossés de Tournemine; ceux de Reculée". 


● Distribution: northern part of Europe: basin of the Balhic Sea, central parts of basins of Danube and 

Dnieper, in lakes and small permanent waterbodies (Starobogatov et al., 2004). 

Contectiana fennica (Kobelt, 1909) F 

Vivipara fennica Kobelt, 1909a: 9, pl. 394, fig. 2265-2267. 

● Type locality: "Finnischer Meerbusen beim Dorfe Lachta (Gouv. St. Petersburg)". 

● Lectotype (Zilch 1955): SMF No. 47879. 

● Distribution: lakes of the basin of the Baltic Sea, upper basin of Volga River (Pleshcheevo Lake, Yaroslavl 

Region), freshwater parts of the Finish Gulf, Ukraine in Shatsk lakes (Anistratenko, 1998b; Starobogatov et

al., 2004). 

Contectiana kormosi (Kobelt, 1909) F 

Vivipara contecta kormosi Kobelt, 1909b: 315-316, pl. 62, fig. 12. 

● Type locality: "Soto in Ungarn". 

● Holotype: SMF No. 48413. 

● Distribution: lakes of the basin of the Baltic Sea, middle Danube basin (Starobogatov et al., 2004). 

Contectiana ladogensis Chernogorenko et Starobogatov, 1987 F 

Chernogorenko, Starobogatov, 1987a: 147. 

● Type locality: northern part of Ladoga Lake (NW European part of Russia). 

● Holotype: ZIN, No. 1 in systematic catalogue. 

● Distribution: type locality (Starobogatov et al., 2004). 

Contectiana listeri (Forbes et Hanley, 1849) F 

[= Vivipara communis Moquin-Tandon, 1855; Paludina moquini Roumeguere, 1858; Vivipara vera 

Frauenfeldt, 1862; Vivipara brachya Bourguignat, 1880; Vivipara carniolica Bourguignat, 1880; Vivipara 

communis Bourguignat, 1880; Vivipara isseli Bourguignat, 1880; Vivipara minutula Bourguignat, 1880; Vivipara 

paludosa Bourguignat, 1880; Vivipara taurica Bourguignat, 1880; Vivipara contecta var.russiensis 

Milachevitch, 1881; Vivipara ranarum Servain, 1884; Vivipara contecta var. rossica Westerlund, 1886; 

Paludina contecta var. gallensteini Kobelt, 1893; Vivipara contecta var. corcyrensis Kobelt, 1909; Vivipara 

contecta var. croatica Kobelt, 1909] 

Paludina listeri Forbes, Hanley, 1849 in 1848-1853: pl. 71, fig. 16; 1850: 8-11. 

● Type locality: southern half of England, Southport in Lancashire, York, the Thames, the Trent, the Ex, the 

Tawe at Swansea. 


● Distribution: Europe, except extreme south and north, Ural River, south of western Siberia (westward of 

Ob River), in lakes and small permanent waterbodies (Anistratenko, 1998b; Pirogov et al., 1994; Starobogatov 

et al., 2004). 

Contectiana turrita (Kobelt, 1909) F 

Vivipara contecta turrita Kobelt, 1909a: 8, pl. 394, fig. 2263. 

● Type locality: "Tollensee in Mecklenburg". 


● Distribution: lakes in the basin of the Baltic Sea, Ukraine (Shatsk lakes) (Anistratenko, 1998b; Starobogatov 

et al., 2004). 

● Remark: the shell of C. turrita illustrated by Chernogorenko (1988: 652, fig. 4 в) looks different as compared 

with the holotype shell portrayed by Zilch (1955, Taf. 3, fig. 1). It raises a question about true identity 

of Contectiana turrita sensu Chernogorenko, 1988. 

Contectiana zebra (Kobelt, 1877) F 

Paludina zebra Kobelt, 1877: 74, pl. 138, fig. 1370. 

● Type locality: "Constantinopel". 

● Lectotype (Zilch, 1955, as "Typus"): SMF No. 26555. 

● Distribution: western Transcaucasia (Starobogatov et al., 2004). 

Viviparus Montfort, 1810 

Type species: Helix viviparus Linnaeus, 1758 (tautonymy) 

Viviparus ater (Christofori et Jan, 1832) F 

[= Paludina crassa Villa et Villa, 1841; Vivipara acerosa Bourgignat, 1862; Vivipara amblia Bourgignat, 

1870; Vivipara penchinati Bourgignat, 1870; Vivipara aristidis Bourgignat, 1880; Vivipara letourneuxi 

Bourgignat, 1880; Vivipara taciti Bourgignat, 1880; Vivipara tanousi Bourgignat, 1880; Vivipara thiesseana

Bourgignat, 1880; Paludina mamillata var.hazayi Westerlund, 1886; Vivipara danubialis var. banatica 

Kobelt, 1909; Vivipara mamillata var. carlovacensis Kobelt, 1909; Vivipara pyramidalis var. erjaveci 

Kobelt, 1909; Vivipara danubialis var. kormosi Kobelt, 1909; Vivipara pyramidalis var. montovana Kobelt, 

1909; Vivipara danubialis var. rumaenica Kobelt, 1909] 

Paludina atra Christofori, Jan, 1832: 3. 

● Type locality: not stated (Italia superioris – from title). 


● Distribution: rivers and lakes of the basins of Danube, Dnieper and Dniester rivers, now through entire 

Dnieper and basin of the Western Dvina River, vicinities of Tolliatti and Pyatigorsk (Anistratenko, 1998b; 


● Remark: according to the P. Glöer’s opinion (pers. communication), the true V. ater is distributed in the 

South Alps only, and all the findings of this species from the ex-USSR territory belong, in reality, to another 

species of this genus, possibly to V. acerosus (Bourgugnat, 1862). In Russian literature, the latter species is 

considered to be a junior synonm of V. ater (Chernogorenko, 1988). However, European malacologists (Falniowski 

et al., 1998; Falkner et al., 2001; Glöer, 2002) regard it as a distinct species. 

Viviparus costae (Heldreich in Mousson, 1863) F 

[Paludina nucleus (Mousson)] 

Paludina costae Heldreich – Mousson, 1863: 18. 

● Type locality: "environs de Constantinople". 


● Distribution: western Transcaucasia, north of Minor Asia, rivers (Starobogatov et al., 2004). 

Viviparus sphaeridius (Bourguignat, 1880) F 

[= Vivipara sphaeridia Bourguignat, 1880; Vivipara strongyla Bourguignat, 1880; Vivipara neptun Clessin, 

1887] 

Vivipara sphaeridia Bourguignat, 1880: 47-48. 

● Type locality: not stated. 


● Distribution: rivers and lakes of the basins of Danube, Dnieper and Dniester rivers, now through the entire 

Dnieper and the basin of the Western Dvina River (Anistratenko, 1998b; Starobogatov et al., 2004). 

Viviparus vistulae (Kobelt, 1907) F 

Vivipara fasciata vistulae Kobelt, 1907: 29. 

● Type locality: "Weichsel bei Graudenz". 

● Lectotype (Zilch 1955): SMF 47828. 

● Distribution: rivers and lakes of the basin of the Baltic Sea (Neman, Visla, Kurshsk Bay) (Starobogatov et 

al., 2004). 

● Remark: Falniowski (1989) treats this species as a intraspecific form Viviparus viviparus f. vistulae. 

Viviparus viviparus (Linnaeus, 1758) F 

[= Nerita fasciata Müller, 1774 (part.); Cyclostoma achatinum Lamarck, 1812; Viviparus vulgaris Gray, 

1850; Paludina duboisiana Mousson, 1863; Vivipara subfasciata Bourguignat, 1870; Vivipara subfasciata 

var. sequanica Bourguignat, 1870; Paludina okaensis Clessin, 1875; Vivipara forbesi Bourguignat, 1880; 

Vivipara nevilli Bourguignat, 1880; Vivipara imperialis Bourguignat, 1884; Vivipara penthica var. albisiana 

Servain, 1884; Vivipara bourguignati Servain, 1884; Vivipara paeteliana Servain, 1884; Vivipara penthica 

Servain, 1884; Vivipara strongyla Servain, 1884; Paludina duboisiana var. concis Westerlund, 1886; 

Paludina penthica var. porphyrea Westerlund, 1886] 

Helix vivipara Linnaeus, 1758: 772. 


● Lectotype (ICZN Opinion 573, 1959): LSL, No. 603. 

● Distribution: rivers and lakes of Europe except for extreme north and south, western Transcaucasia (Anistratenko, 

1998b; Starobogatov et al., 2004; Pirogov et al., 1994). Introduced in the Novosibirsk Reservoir

(Andreev et al., 2008). 

POMATIIDAE Gray, 1852 


Pomatias Studer, 1789 

Type species: Nerita elegans Müller, 1774 (SD Newton, 1891) 

Pomatias hyrcanum (Martens, 1874) T 

[= Cyclostoma caspius Mousson, 1876] 

Cyclostoma costulatum var. hyrcanum Martens, 1874: 30. 

● Type locality: «Rescht und Enzeli an der Südküste des kaspischen Meeres» (now Rasht and Bandar-e Pahlavi, 

northern Iran). 


● Distribution: forests of Talysh, Lenkoran and Khachmass lowlands (Azerbaijan) (Likharev, Schileyko, 

MS). 

Pomatias rivulare (Eichwald, 1829) T 

[= Cyclostoma costulatum Rossmässler, 1837] 

Cyclostoma rivulare Eichwald, 1829: 302. 

● Type locality: vicinities of Tarki village, near Makhachkala [Daghestan]. 


● Distribution: Crimea: foothills along upper and medium parts of the Karasu river (Puzanov, 1927); Carpathians 

(Anistratenko, Anistratenko, 2001); Moldavia (Kodry Reserve) (Coadã, 2007); forest areas of North 

Caucasus and Transcaucasia, except Talysh and forests of Lenkoran and Khachmass lowlands, where the 

species is replaced by Pomatias hyrcanum (Likharev, Schileyko, MS). 

Clade SORBEOCONCHA 

CERITHIOIDEA Fleming, 1822 

MELANOPSIDAE H. et A. Adams, 1854 

(Kantor Yu.I.; Vinarski M.V.) 

Fagotia Bourguignat, 1884 

Type species: Melanopsis esperi Férussac, 1823 (SD Wenz, 1939) 

● Remarks: the genus Fagotia is considered sometimes as a synonym of Esperiana Bourguignat, 1877 

(Falkner et al., 2001), although treated as a full genus by Starobogatov et al. (2004). Fagotia acicularis 

(Férussac, 1823) and F. esperi (Férussac, 1823), recorded from Russian territory by Shadin (1952), are 

in fact absent there (Starobogatov et al., 1992). 

Fagotia berlani Bourguignat, 1884 F 

Bourguignat, 1884: 34-35. 

● Type locality: “Le Danube а Ibralia; la Save entre Agram et Sissek”. 

● Types: possibly in MNHN or MHNG, Bourguignat collection. 

● Distribution: lower Danube and rivers of northwestern basin of the Black Sea (Anistratenko, 1998b; Starobogatov 

et al., 2004). 

● Remark: Although in 1884 Bourguignat cited the date for the species as 1880, its description was not published 

before. He might refer to the manuscript date rather that to publication date.

Fagotia danubialis Bourguignat, 1884 F 

Bourguignat, 1884 [1880]: 35-36. 

● Type locality: “Le Danube а Ibralia; la Save а Agram; la Krapina а Sused (Croatie)”. 



et al., 2004). 

● Remark: Although in 1884 Bourguignat cited the date for the species as 1880, its description was not published 

before. He might refer to the manuscript date rather that to publication date. 

Fagotia dneprensis Starobogatov, Alexenko et Levina, 1992 F 

Starobogatov et al., 1992: 64, fig. 3 (11). 

● Type locality: Dnieper River near Kherson. 

● Holotype: ZIN, № 1 in systematic catalogue. 


et al., 2004). 

Fagotia roseni Starobogatov in Starobogatov, Alexenko et Levina, 1992 F 

Starobogatov et al., 1992: 62, fig. 3 (4, 5). 

● Type locality: Quaternary deposits near Kulevi (formerly Redul-Kaleh), Georgia. 


● Distribution: Quaternary deposits of western Georgia, probably lives in rivers of South Black Sea maritime 

territory (Starobogatov et al., 1992). 

Melanopsis Férussac, 1807 

Type species: Buccinum praerosum Linnaeus, 1758 (SD Wenz, 1939) 

● Remark: Melanopsis praerosa (L., 1758), recorded for Russian territory by Shadin (1952), is absent here 

(Starobogatov et al., 1992). 

Melanopsis ashkhabadensis Izzatullaev et Starobogatov, 1984 F 

Izzatullaev, Starobogatov, 1984: 1480, fig. 1 14, 3 8 

● Type locality: Ashkhabad, rapid mountain springs — Ferjuza, Guli, etc. [former Turkmen SSR]. 


● Distribution: Ashkhabad region (Izzatullaev, Starobogatov, 1984). 

Melanopsis buccinoidea Olivier, 1801 F 

Olivier, 1801: 297, Atlas pl. 17, fig. 8. 

● Type locality: "les eaus douces de Scio, de presque toutes les оles de l'Archipel, de Crète, de Syrie". 


● Distribution: western Georgia (Izzatullaev, Starobogatov, 1984). 

Melanopsis denegabilis Férussac in Pallary, 1939 F 

Melanopsis denegabilis Férussac – Pallary, 1939: 85-86, textfig. 11, pl. 6, figs. 9-13. 

● Type locality: "La localité originale nést pas connue. La seule provenance indiquée est la Mésopotamie. ... 

Cette espèce est très commune au Levant. ... En Iraq, ... а Rahalya Springs, а Kani Sepi et а Karsi". 

● Types: BMNH. 

● Distribution: small rivers flowing to the Kura River from Lesser Caucasus (Kazakh district of Azerbaijan 


Melanopsis laevigata Lamarck, 1816 F 

[= Melanopsis ascania Bourguignat, 1880] 

Lamarck, 1816: 12, pl. 458, fig. 8.


● Syntypes: MHNG. 


Melanopsis mingrelica Bayer in Mousson, 1863 F 

Melanopsis mingrelica Bayer – Mousson, 1863: 91-92. 

● Type locality: "de l'intérieur de la Mingrélie ..., puis de Rédutkaleh". 



Melanopsis minutula Bourguignat, 1884 F 

Bourguignat, 1884: 92-93. 

● Type locality: "Fontaine froide du Hamman а Brousse (Anatolie); Nahr-Antalies dans le Liban (Syrie); puits 

artésien de Tamerna-Kedima, dans le Ziban (Algerie)". 



Melanopsis roseni Izzatullaev et Starobogatov, 1984 F 

Izzatullaev, Starobogatov, 1984: 1479, fig. 1, 11. 

● Type locality: Zakaspijsk Region (former Turkmen SSR). 


● Distribution: Zakaspijsk Region (Izzatullaev, Starobogatov, 1984). 

Melanopsis sistanica Izzatullaev et Starobogatov, 1984 F 


● Type locality: "Eastern Persia", coll. N.A.Zarudnyj, 1896 


● Distribution: “Sistan” region (Izzatullaev, Starobogatov, 1984). 

Melanopsis starostini Izzatullaev et Starobogatov, 1984 F 

Izzatullaev, Starobogatov, 1984: 1480, fig. 1 13, 3 7. 

● Type locality: Gennob, Central Kopetdag (former Turkmen SSR). 


● Distribution: Central Kopetdag (Izzatullaev, Starobogatov, 1984). 

Melanopsis transcaspia Izzatullaev et Starobogatov, 1984 F 


● Type locality: Gjaure stream, collective farm "imeni 1 Maja", Ashkhabad Region (former Turkmen SSR). 


● Distribution: type locality (Izzatullaev, Starobogatov, 1984). 

Melanopsis turkmenica Izzatullaev et Starobogatov, 1984 F 

Izzatullaev, Starobogatov, 1984: 1478-1479, fig. 1, 10. 

● Type locality: Zakaspijsk Region [former Turkmen SSR]. 



Melanopsis variabilis (v. d. Busch in Philippi, 1847) F 

Melania (Melanopsis) variabilis v. d. Busch – Philippi, 1847: 175, pl. IV, fig. 7, 8, 10. 

● Type locality: “Schiraz et Persepolis, Persiae”. 


● Distribution: small rivers flowing to the Kura River from Lesser Caucasus (Kazakh district of Azerbaijan 

(Starobogatov et al., 1992).

Melanopsis zarudnyi Izzatullaev et Starobogatov, 1984 F 

Izzatullaev, Starobogatov, 1984: 1479-1480, fig. 1 12, 3 6. 

● Type locality: "Golden spring" near Ashkhabad [former Turkmen SSR]. 



Microcolpia Bourguignat, 1884 

Type species: Melanopsis acicularis Férussac, 1823 (SD Wenz, 1939) 

● Remarks: the genus Microcolpia is considered sometimes as a subgenus of Esperiana Bourguignat, 1877 

(Glöer, 2002), although treated as a full genus by Starobogatov et al. (2004). 

Microcolpia canaliculata Bourguignat, 1884 F 

Bourguignat, 1884: 62. 

● Type locality: "Le Danube а Ibraпla". 


● Distribution: lower Danube and rivers of north-western Black Sea basin (Anistratenko, 1998b; Starobogatov 

et al., 2004). 

Microcolpia potamactebia (Bourguignat, 1870) F 

Melanopsis potamactebia Bourguignat, 1870: 67-68. 

● Type locality: "le Danube ... de Brahilov, .. des environs de Belgrade". 


● Distribution: lower Danube and rivers of north-western Balck Sea basin (Starobogatov et al., 2004). 

Microcolpia ucrainica Starobogatov, Alexenko et Levina, 1992 F 

Starobogatov et al., 1992: 69, fig. 3, 20. 

● Type locality: Yuznyj Bug River [Ukraine]. 


● Distribution: lower Danube and rivers of north-western Black Sea basin (Anistratenko, 1998b; Starobogatov 

et al., 2004). 

PACHYCHILIDAE P. Fischer et Crosse, 1892 

(Kantor Yu. I., Sysoev A.V.) 

“Parajuga” Prozorova et Starobogatov, 2003 

● Remark: this name first appeared in Starobogatov et al., 2004, as published in 2004. Neverheless, the description 

of the taxon was not published prior to the publication of the mentioned work (L.A.Prozorova, personal 

communication). According to articles 16.1 and 16.2 of ICZN the name is not available, since it was 

not explicitly indicated as intentionally new and the type species was not mentioned. 

“Parajuga” amgunica (Moskvicheva in Zatrawkin, 1986) F 

Juga (Hua) chankensis Moskvicheva – Zatrawkin, 1986: 35, fig. 1 (7), 2 (7). 

● Type locality: mouth of Nimilen River, tributary of Amgun’ River. 


● Distribution: basin of Amgun and Tugur rivers (northern Khabarovsk Territory, rivers (Bogatov, 

Zatrawkin, 1990[1992]; Starobogatov et al., 2004). 

“Parajuga” amurensis (Gerstfeldt, 1859) F 

[= Semisulcospira cancellata sensu Shadin, 1952, part. non Benson, 1833] 

Melania amurensis Gerstfeldt, 1859: 512-514, figs. 14-24. 

● Type locality: "im Amur, im mittleren Laufe und einem Theile des unteren dieses Stromes".

● Lectotype (Bogatov, Zatrawkin, 1990[1992]): ZIN, No. 1 in systematic catalogue. 

● Distribution: entire Amur basin, in rivers (Starobogatov et al., 2004). 

“Parajuga buettneri” (Ehrmann in Buettner et Ehrmann, 1927) F 

Melania buettneri Ehrmann – Buettner, Ehrmann, 1927: 246-247. 

● Type locality: Ussuri River, Spasskoje. 

● Types: not traced. According to Dance (1986) information, the Ehrmann collection is housed in SMF. 

● Distribution: Amur below Bureinskyi Ridge, in rivers and flowing parts of lakes (Starobogatov et al., 

2004). 

“Parajuga” arsenievi Prozorova et Starobogatov, 2004 F 

Prozorova et Starobogatov, 2004: 281, pl. 106, fig. 5. 

● Distribution: Zerkalnaya River basin (eastern slopes of Sikhote-Alin, central Primorje) (Starobogatov et al., 

2004). 


of the species was not published prior to the publication of the mentioned work (L.A.Prozorova, 

personal communication). According to articles 16.1 of ICZN the name is not available, since it was not explicitly 

indicated as intentionally new and the type material was not mentioned. 

“Parajuga” chankensis (Moskvicheva in Zatrawkin, 1986) F 

Juga (Hua) chankensis Moskvicheva – Zatrawkin, 1986: 33, 34, fig. 1,5, 2,5. 

● Type locality: middle Amur river. 


● Distribution: entire Amur basin except for Amgun and upper reaches of Zeya and Bureya; in rivers and 

lakes (Bogatov, Zatrawkin, 1990[1992]; Starobogatov et al., 2004). 

“Parajuga” czerskii (Moskvicheva in Zatrawkin, 1986) F 

Juga (Hua) czerskii Moskvicheva – Zatrawkin, 1986: 34, 35, fig. 1,6, 2,6. 

● Type locality: valley of Odarka River, Khanka Lake basin. 


● Distribution: Ussuri basin, Amur basin (from Bureya Ridge to Bogorodskoe village); in rivers (Bogatov, 


“Parajuga” heukelomiana (Reeve, 1859) F 

[= Semisulcospira cancellata sensu Shadin, 1952, part. non Benson, 1833] 

Melania heukelomiana Reeve, 1859: pl. 17, sp. 123. 

● Type locality: not known to author. 


● Distribution: entire Amur basin; in rivers and large lakes (Bogatov, Zatrawkin, 1990[1992]; Starobogatov 

et al., 2004). 

“Parajuga” nodosa (Westerlund, 1897) F 

[= Melania amurensis var. laevigata Gerstfeldt, 1859, non Melania laevigata Lamarck, 1815; Melania 

amurensis sensu Shadin, 1952, part. non Gerstfeldt, 1859] 

Baicalia (Maackia) nodosa Westerlund, 1897b: 128 

● Type locality: "Sibirien. Fluss Argunj". 


● Distribution: entire Amur basin except for Amgun and upper reaches of Zeya; in rivers and lakes (Bogatov, 

Zatrawkin, 1990[1992]; Starobogatov et al., 2004).

“Parajuga” popovae Prozorova et Starobogatov, 2004 F 

Prozorova et Starobogatov, 2004: 136 (nom. nov. pro Juga (Hua) starobogatovi Zatrawkin et Moskvicheva in 

Zatrawkin, 1986, non Popova, 1981). 

● Distribution: Tugur River (northern Khabarovsk Territory); in slowly flowing rivers (Bogatov, Zatrawkin, 

1990[1992]; Starobogatov et al., 2004). 





“Parajuga” sihotealinica Prozorova et Starobogatov, 2004 F 



2004). 





“Parajuga” subcalculus Prozorova et Starobogatov, 2004 F 

Prozorova et Starobogatov, 2004: 281, pl. 106, fig. 8 (nom. nov. pro Juga calculus sensu Zatrawkin et Moskvicheva 

in Zatrawkin, 1986, non Melania calculus Reeve, 1859). 

● Distribution: southern Primorje Territory, rivers and lakes of basins of Amur and Ussuri bays (Starobogatov 

et al., 2004). 





“Parajuga” subextensa Prozorova et Starobogatov, 2004 F 

Prozorova et Starobogatov, 2004: 282-283, pl. 106, fig. 13 (nom. nov. pro Juga extensa sensu Zatrawkin et 

Moskvicheva in Zatrawkin, 1986, non Melania extensa Martens). 

● Distribution: southern Primorje Territory, rivers and lakes of basins of Amur and Ussuri bays (Starobogatov 

et al., 2004). 





“Parajuga” subtegulata Prozorova et Starobogatov, 2004 F 

Prozorova et Starobogatov, 2004: 282, pl. 106, fig. 11 (nom. nov. pro Juga tegulata sensu Zatrawkin et 

Moskvicheva in Zatrawkin, 1986, non Melania tegulata Martens, 1905). 

● Distribution: rivers and lakes of southern Primorje (basins of Anur and Ussuri bays) (Starobogatov et al., 

2004). 





“Parajuga” tugurensis (Zatrawkin et Moskvicheva in Zatrawkin, 1986) F 

Juga (Hua) tugurensis Zatrawkin, Moskvicheva – Zatrawkin, 1986: 36, fig. 1,11. 

● Type locality: Tugur River, 35 km upstream from the mouth, near Tugur settlement.


● Distribution: Tugur River basin (northern Khabarovsk Territory); in rivers (Bogatov, Zatrawkin, 


“Parajuga” zerkalnensis Prozorova et Starobogatov, 2004 F 



2004). 





THIARIDAE Gill, 1871 (1823) 

(Kantor Yu.I.) 

Melanoides Olivier, 1804 

Type species: Melanoides fasciolata Olivier, 1804 (monotypy) 

Melanoides kainarensis Starobogatov et Izzatullaev, 1980 F 

Starobogatov, Izzatullaev, 1980: 26-28, fig. 2 1, 2 8, 

● Type locality: warm spring KhodzhaKajnar, Kugitang Ridge, Chardzhou region, Turkmen SSR. 


● Distribution: Charshangin District, Chardzhou Region, Turkmenia (Starobogatov, Izzatullaev, 1980). 

Melanoides pamiricus Lindholm, 1930 F 

Melanoides tuberculatus pamiricus Lindholm, 1930: 69, figs. 1, 2. 

● Type locality: (hot spring Dzhaushangoz, upper reaches of Shakhdara River, southeastern Tajikistan). 

● Types: probably lost (Starobogatov, Izzatullaev, 1980). 

● Distribution: hot spring Dzhaushangoz, southeastern Tajikistan (Starobogatov, Izzatullaev, 1980). 

Melanoides shahdaraensis Starobogatov et Izzatullaev, 1980 F 

Starobogatov, Izzatullaev, 1980: 29-30, fig. 2 7, 13. 

● Type locality: hot spring Dzhaushangoz, 3360 m above the sea level, Shakhdar River basin, western Pamir. 


● Distribution: type locality (Starobogatov, Izzatullaev, 1980). 

CLADE LITTORINIMORPHA 

RISSOOIDEA Gray, 1847 

AMNICOLIDAE Tryon, 1863 

AMNICOLINAE Tryon, 1863 (including Terrestribythinellidae) 

(Kantor Yu. I., Vinarski M.V., Sysoev A.V.) 

Akiyoshia Kuroda et Habe, 1954 

Type species: Akiyoshia uenoi Kuroda et Habe (OD)

Akiyoshia sachalinensis Zatrawkin et Bogatov, 1988 F 

Zatrawkin, Bogatov, 1988: 129, textfig. (7). 

● Type locality: southern Sakhalin, near Yablochnyi settlement. 


● Distribution: southern Sakhalin (Bogatov, Zatrawkin, 1990[1992]). 

Bythinella Moquin-Tandon, 1855 

Type species: Bulimus viridis Poiret, 1801 (fixation of the types species not traced) 

Bythinella adsharica Lindholm, 1913 F 

Lindholm, 1913: 67. 

● Type locality: vicinities of Batum, spring near river Adsharis-Zchali at its entrance to Tschoroch. 

● Syntypes: ZIN? 

● Distribution: Transcaucasia (Adzharia) (Shadin, 1952). 

Bythinella austriaca (Frauenfeld, 1857) F 

Paludinella austriaca Frauenfeld, 1857: 574, fig. 2. 

● Type locality: “Quellen von Dornbach nächst Wien”, Austria. 

● Types: Vienna Museum? 

● Distribution: central part of Danube basin and in Karpatian area of Ukraine (Starobogatov, Zatrawkin, 

1987; Anistratenko, 1998b). 

● Remark: outside the territory of former USSR the species is also recorded in Sudety and Tatry, as well in 

western Romania and Southern Germany (Glöer, 2002). This species includes at least 4 subspecies (Glöer, 

2002) and it is not clear which one is found in Russian territory. 

Bythinella hungarica Hazay, 1881 F 

Hazay, 1881: 93, pl. 14, fig. 1. 

● Type locality: Budapest, St. Margaretr Island. 


● Distribution: central part of Danube basin and in Karpatian area of Ukraine (Starobogatov, Zatrawkin, 

1987; Anistratenko, 1998b). 

Kolhymamnicola Starobogatov et Budnikova, 1976 

Type species: Amnicola kolhymensis Starobogatov et Streletzkaja, 1967 (OD) 

Kolhymamnicola kolhymensis (Starobogatov et Streletzkaja, 1967) F 

Amnicola kolhymensis Starobogatov, Streletzkaja, 1967: 228-229, fig. 12. 

● Type locality: Hood lands of the Kolyma River, near the small settlement Karetovo. 


● Distribution: basins of Kolyma river, rivers of Chukotka and Kamchatka, in small permanent waterbodies 

(Starobogatov, Streletzkaya, 1967; Starobogatov et al., 2004); Lena, Indigirka and Yana river basins (Dolgin, 

2001). 

Kolhymamnicola miyadii (Habe, 1942) F 

Amnicola miyadii Habe, 1942: 25, 27, figs. 1, 2. 

● Type locality: Simusyu-tх of the North Kurile Islands [Simushir?]. 

● Holotype: not traced. 

● Distribution: North Kurile Islands, Kamchatka, in lakes (Prozorova, 1998a; Starobogatov et al., 2004). 

Kolhymamnicola ochotica Zatrawkin et Bogatov, 1988 F 

Zatrawkin, Bogatov, 1988: 128, textfig. (5). 

● Type locality: Magadan Region, Okhotsk District, Bolshoye Ueginskoye Lake. 

● Holotype: ZIN, No. 1 in systematic catalogue.

● Distribution: basin of Okhota River (north of Khabarovsk Region), in lakes (Bogatov, Zatrawkin, 


Kolhymamnicola wasiliewae Zatrawkin et Bogatov, 1988 F 

Zatrawkin, Bogatov, 1988: 127-128, textfig. (3, 4). 

● Type locality: small inlet on the river Bolin (tributary of the lower Amur River) near Amursk city. 


● Distribution: Amur basin, in small rivers (Bogatov, Zatrawkin, 1990[1992]; Starobogatov et al., 2004); 

Sakhalin Island (Prozorova et al., 2004). 

Marstoniopsis Regteren Altena, 1936 

Type species: Hydrobia steinii Martens, 1858 (OD) 

Marstoniopsis scholtzi (Schmidt, 1856) F 

Hydrobia scholtzi Schmidt, 1856: 158. 

● Type locality: “sumpfigen Wiesengräben zwischen Breslau und Marienau”. 


● Distribution: basin of Baltic Sea, upper part of Volga basin, in rivers on slow stream and in lakes (Starobogatov 

et al., 2004); Zapadnyi Bug (Anistratenko, 1998b – as Marstoniopsis steinii (Martens, 1858)). 

Outside former USSR also in Great Britain, Netherlands and Belgium (Glöer, 2002; Anderson, 2005) 

Marstoniopsis steinii (von Martens, 1858) F 

Hydrobia steinii von Martens, 1858: 183, Taf. V, fig. 5. 

● Type locality: “Lake Tegel between Berlin and Spandau [nowadays in Berlin]”. 

● Types: not traced. Possibly, in ZMB (fide Dance, 1986). 

● Distribution: basin of Baltic Sea, upper part of Volga basin, in rivers on slow stream and in lakes (Starobogatov 

et al., 2004), Zapadnyi Bug (Anistratenko, 1998b). 

● Remark: European malacologists (Glöer, 2002; Anderson, 2005) consider M. steinii as a junior synonym 

of M. scholtzii. 

Terrestribythinella Sitnikova, Starobogatov et Anistratenko, 1992 

Type species: Terestribythinella baidashnikovi Sitnikova, Starobogatov et Anistratenko, 1992 (OD) 

Terrestribythinella amphibiotica Anistratenko, 1995 T 

Anistratenko, 1995: 66-68. 

● Type locality: «Zachrovana Dolina» area, Ilyinetzkoje Forestry, Irshava District, Zakarpatskaya Region. 

● Holotype: IZU. 

● Distribution: type locality (Anistratenko, 1998b). 

Terrestribythinella baidashnikovi Sitnikova, Starobogatov et Anistratenko, 1992 T 

Sitnikova et al., 1992: 10-11, fig. 2 (3). 

● Type locality: Ugolskoe Forestry, Karpatsky Natural Reserve, Tyachev District, Zakarpatskaya Region. 



Terestribythinella carpathica Sitnikova, Starobogatov et Anistratenko, 1992 T 

Sitnikova et al., 1992: 11, fig. 2 (4). 

● Type locality: Ugolskoe Forestry, Karpatsky Natural Reserve, Tyachev District, Zakarpatskaya Region. 



BAICALIINAE P. Fischer, 1885


Baicalia Martens, 1876 

Type species: Ligea carinata W. Dybowski, 1875 (SD Dall, 1877) 

Baicalia carinata carinata (W. Dybowski, 1875) F 

[= Trachybaikalia carinata martensiana subvar. orthos B. Dybowski et Grochmalicki, 1914; Trachybaikalia 

carinata martensiana elencha B. Dybowski et Grochmalicki, 1914] 

Ligea carinata W. Dybowski, 1875: 45-46, pl. 4, fig. 1, pl. 6, fig. 7, a–d. 

● Type locality: originally “Baikal, 300-350 m”, Kultuk (SW coast of southern Baikal) (restricted Sitnikova 

et al., 2004). 

● Syntypes: ZIN. 

● Distribution: southern and east-central Baikal, 50-100 m (Sitnikova et al., 2004). 

Baicalia carinata fuchsiana (B. Dybowski, Grochmalicki, 1915) F 

Trachybaicalia carinata var. fuchsiana B. Dybowski, Grochmalicki, 1915a: 299, pl. 2, fig. 5 a–d. 

● Type locality: not stated (Balkal – from title). 

● Syntypes: not traced. 

● Distribution: southern Baikal, 12-20 m (Sitnikova et al., 2004). 

Baicalia carinata hoernesiana (B. Dybowski et Grochmalicki, 1915) F 

[= Baicalia carinata carinata, part. sensu Kozhov, 1936, non W. Dybowski, 1875] 

Trachybaicalia carinata var. hoernesiana B. Dybowski, Grochmalicki, 1915a: 295-296, pl. 2, fig. 4 a–b. 

● Type locality: "Baikalsee". 

● Syntype: ZIN. 

● Distribution: northern Baikal and Maloe More, 10-100 m (Sitnikova et al., 2004). 

Baicalia dybowskiana dybowskiana (Lindholm, 1909) F 

[= Trachybaikalia dybowskiana lindholmi B. Dybowski et Grochmalicki, 1914] 

Baikalia (Trachybaikalia) dybowskiana Lindholm, 1909: 71, texfig. 3. 

● Type locality: Berhin (W coast of central Baikal). 


● Distribution: Baikal, 3-80 m (Sitnikova et al., 2004). 

Baicalia dybowskiana carinatoides Kozhov, 1936 F 

Baicalia (Baicalia) dybowskiana var. carinatoides Kozhov, 1936: 120, pl. 4, fig. 12. 

● Type locality: Chivyrkyiskij Bay (northern Baikal). 

● Type material: lost (Sitnikova et al., 2004). 

● Distribution: type locality, 1.5-2 m (Kozhov, 1936; Sitnikova et al., 2004). 

Baicalia carinatocostata (W. Dybowski, 1875) F 

[= Trachybaikalia carinato–costata, T. carinato–costata bittneri subvar. clara, opaca, pyramidalis, micronella; 

T. carinato–costata credneri subvar. inflatella, T. carinato–costata sanbergeri, T. carinato–costata 

moussoni – B. Dybowski, Grochmalicki, 1914] 

Ligea carinato–costata W. Dybowski, 1875: 46, pl. 3, figs. 24-26, pl. 6, fig. 8. 

● Type locality: originally “Baikal, 300-350 m”, Kultuk (SW coast of southern Baikal) (restricted Sitnikova 

et al., 2004). 



Baicalia rugosa (Lindholm, 1909) F 

[= Trachybaicalia carinata var. martensiana subvar. rudis B. Dybowski, Grochmalicki, 1914] 

Baikalia (Trachybaikalia) carinata forma rugosa Lindholm, 1909: 69.

● Type locality: Svyatoi Nos (northern Baikal). 


● Distribution: northern and central Baikal, 3-40 m (Sitnikova et al., 2004). 

Baicalia turriformis (W. Dybowski, 1875) F 

Ligea turriformis W. Dybowski, 1875: 50-51, pl. 3, figs. 4-7; pl. 6, fig. 9 a-d. 

● Type locality: originally “Baikal, 300-350 m”; Kultuk (SW coast of southern Baikal) (restricted Sitnikova 

et al., 2004). 


● Distribution: southern and central Baikal, 10-500 m (Sitnikova et al., 2004). 

Godlewskia Crosse et Fisher, 1897 

Type species: Leucosia godlewskii W. Dybowski, 1875 (OD) 

Godlewskia bacilliformis (Kozhov, 1936) F 

Baicalia (Gerstfeldtia) bacilliformis Kozhov, 1936: 110, 310, pl. 3, figs. 31-33. 

● Type locality: near village Syhaya, 39 m (E coast of southern Baikal); Oto-Khushun, 72 m (Maloe More); 

Barguzin Bay, 20-40 m. 


● Distribution: southern and central Baikal, 20–100 m (Sitnikova et al., 2004). 

Godlewskia columella (Lindholm, 1909) F 

[= Gerstfeldtia columella, var. typica, tumida, spicata, rufula – B. Dybowski et Grochmalicki, 1913] 

Baikalia (Gerstfeldtia) columella Lindholm, 1909: 67, pl. 1, figs. 46-48. 

● Type locality: off Baranchiki (W coast of southern Baikal). 



Godlewskia godlewkii godlewkii (W. Dybowski, 1875) F 

[= Gerstfeldtia godlewskii ladislavi, G. g. felixi, G. g. henrici, G. g. stanislavi, G. g. constantinae, G. g. antoninae, 

G. g. medialia, G. g. parvula, G. g. victori, G. g. kasimirae – B. Dybowski, Grochmalicki, 1913] 

Leucosia godlewskii W. Dybowsky, 1875: 40-41, pl. 3, figs. 10-14, pl. 6, fig. 2a-d. 

● Type locality: not stated in original description, Kultuk (SW coast of southern Baikal) (restricted Sitnikova 

et al., 2004). 



● Remark: possibly, some of 10 infraspecific forms described by Dybowski and Grochmalicki (1913) may 

apper to be separate species and subspecies differing in the shell size and the whorl convexity. The subspecies 

Godlewskia godlewkii speciosa (Kozhov, 1936) may thus appear to be a junior synonym of one of these 

varieties (Sitnikova et al., 2004). 

Godlewskia godlewkii speciosa (Kozhov, 1936) F 

Baicalia (Gerstfeldtia) godlewskii var. speciosa Kozhov, 1936: 114, 310. 

● Type locality: Maloe More (border between central and northern Baikal). 


● Distribution: Maloe More and northern Baikal, 10-250 m (Sitnikova et al., 2004). 

Godlewskia pulchella pulchella (W. Dybowski, 1875) F 

[= Gerstfeldtia pulchella pulchella, G. p. conoidalis, G. p. columnalis, G. p. fuscata, G. p. fusca – B. Dybowski 

et Grochmalicki, 1913] 

Leucosia godlewskii var. pulchella W. Dybowski, 1875: 41, pl. 3, figs. 15-19. 

● Type locality: not stated in original description; Kultuk (SW coast of southern Baikal) (restricted Sitnikova 

et al., 2004).



Godlewskia pulchella intermedia (Kozhov, 1936) F 

Baicalia (Gesrtfeldtia) pulchella var. intermedia Kozhov, 1936: 115-116, 311, pl. 4, figs. 37- 40. 

● Type locality: Ongureny (northern Baikal, slightly northward of Maloe More) and the entire northern half 

of Baikal, 8-180 m. 


● Distribution: northern Baikal, Maloe More 10-180 m (Sitnikova et al., 2004). 

Godlewskia wrzesniowskii wrzesniowskii (W. Dybowski, 1875) F 

Ligea wrzesniowskii W. Dybowski, 1875: 47, pl. 3, figs. 44-46, pl. 6, fig. 14. 


et al., 2004). 



Godlewskia wrzesniowskii olchonensis (Kozhov, 1936) F 

Baicalia (Gerstfeldtia) wrzesniowskii var. olchonensis Kozhov, 1936: 108-109, 310, pl. 3, figs. 29-30. 

● Type locality: Olhonskiye Vorota, 30-40 m (border between central and northern Baikal); Barguzin Bay. 


● Distribution: central Baikal, 20–40 m (Sitnikova et al., 2004). 

Godlewskia wrzesniowskii profunda (Lindhlm, 1924) F 

Baicalia (Baicalia) herderiana profunda Lindholm, 1924a: 23. 

● Type locality: Tukalaragda (NE coast of northern Baikal). 


● Distribution: northern Baikal, 15–25 m (Sitnikova et al., 2004). 

Korotnewia Kozhov, 1936 

Type species: Baikalia (Godkewskia) korotnevi Lindholm, 1909 (OD) 

Korotnewia angigyra (Lindholm, 1909) F 

Baikalia (Godlewskia) angigyra Lindholm, 1909: 64, pl. 1, figs. 56-58. 

● Type locality: Boguchanskaya Bay (W coast of northern Baikal). 


● Distribution: northern Baikal, Maloe More, 11-200 m (Sitnikova et al., 2004). 

Korotnewia korotnevi korotnevi (Lindholm, 1909) F 

Baikalia (Godlewskia) korotnevi Lindholm, 1909: 65, pl. 1, figs. 22-24. 

● Type locality: Dagarskaya Bay (northern Baikal). 

● Syntypes: ANSP, ZIN. 


Korotnewia korotnevi gracilis (Kozhov, 1936) F 

[= Baikalia (Godlewskia) korotnevi Lindholm, 1909, part.] 

Baicalia (Korotnewia) korotnewi var. gracilis Kozhov, 1936: 78, 82-83, 308, pl. 5, figs. 31-33. 

● Type locality: Olhonskiye Vorota (border between central and northern Baikal), Mukhor Bay (Maloe 

More), 2-5 m. 


● Distribution: Maloe More [Baikal], 2-20 m (Sitnikova et al., 2004).

Korotnewia korotnevi selengensis (Kozhov, 1936) F 

Baicalia (Korotnewia) korotnewi var. selengensis Kozhov, 1936: 78, 83-84, 308, pl. 5, figs. 40-42; pl. 8, fig. 

9. 

● Type locality: off Selenga River mouth, Posolskyi Sor (E coast of central Baikal), 8-20 m. 


● Distribution: central Baikal, 10-120 m (Sitnikova et al., 2004). 

Korotnewia semenkewitschi semenkewitschi (Lindholm, 1909) F 

Baikalia (Parabaikalia) semenkewitschi Lindholm, 1909: 30; pl. 1, figs. 30, 31. 




Korotnewia semenkewitschi nitida (Kozhov, 1936) F 

Baicalia (Korotnewia) semenkewitschi forma nitida Kozhov, 1936: 79, 80. 

● Type locality: Selenga shallows near Mysovaya and Boyarskaya (SE coast of southern Baikal). 



Liobaicalia Martens, 1876 

Type species: Leucosia stiedae W. Dybowski, 1875 (SD Dall, 1877) 

Liobaicalia stiedae (W. Dybowski, 1875) F 

Leucosia stiedae W. Dybowski, 1875: 38, pl. 3, figs. 20-23; pl. 6, fig. 1, pl. 7, figs. 9-10. 

● Type locality: originally “"Baikalsee"”; Kultuk (SW coast of southern Baikal) (restricted Sitnikova et al., 

2004). 



● Remark: Sitnikova (1991) suggested, that Liobaicalia stiedae polita Dybowski et Grochmalicki, 1920 is a 

synonym of L. stiedae stiedae; L. s. hispida polita Dybowski et Grochmalicki, 1920 should be considered as 

a subspecies; while L. stiedae parvula polita Dybowski et Grochmalicki, 1920 should be a separate species, 

although still there are no proofs for that (Sitnikova et al., 2004). 

Maackia Clessin, 1880 

Type species: Ligea costata W. Dybowski, 1875 (SD Westerlund, 1880) 

Maackia angarensis (Gerstfeldt, 1859) F 

Hydrobia angarensis Gerstfeldt, 1859: 7, figs. 12-13. 

● Type locality: Angara, Baikal. 


● Distribution: Angara River and SW littoral of Baikal in the area of headwaters of Angara, 1-20 m (Starobogatov 

et al., 2004; Sitnikova et al., 2004). 

Maackia bythiniopsis (Lindholm, 1909) F 

Baikalia (Baikalia) bythiniopsis Lindholm, 1909: 45, pl. 1, figs. 1-4. 

● Type locality: Baranchuk (W coast of southern Baikal). 



Maackia costata (W. Dybowski, 1875) F 

Ligea costata W. Dybowski, 1875: 46-47, pl. 3, figs. 34-37; pl. 6, fig. 12a-d. 


et al., 2004).



Maackia herderiana herderiana (Lindholm, 1909) F 

Baikalia (Baikalia) herderiana Lindholm, 1909: 47, pl. 1, figs. 11-14. 

● Type locality: Baranchuk, Listvennichnoye (SW coast of Baikal). 


● Distribution: southern and central Baikal (Sitnikova et al., 2004). 

● Remark: Starobogatov et al. (2004) list this taxon as Maackia raphidia herderiana. 

Maackia herderiana parvula Kozhov, 1936 F 

Baicalia (Eubaicalia) herderiana var. parvula Kozhov, 1936: 57, 60, 307, pl. 3, fig. 14. 

● Type locality: Proval Bay, shallow water (east-central Baikal). 



Maackia herderiana semicostulata (Lindholm, 1924) F 

Baicalia (Baicalia) angarensis semicostulata Lindholm, 1924a: 22. 

● Type locality: Kultuk (SW coast of southern Baikal). 



Maackia pusilla (Lindholm, 1909) F 

Baikalia (Baikalia) pusilla Lindholm, 1909: 49, pl. 1, figs. 49-51. 

● Type locality: Haransa Bay in Maloe More (border between central and northern Baikal). 


● Distribution: Baikal Lake: Maloe More and Bolshoi Ushkanii Island, 5-20 m (Sitnikova et al., 2004). 

Maackia raphidia (Bourguignat, 1860) F 

Bithynia raphidia Bourguignat, 1860a: 537, pl. 23, figs. 20, 21. 

● Type locality: “lac Baikal”. 


● Distribution: Angara River to Irkutsk (Starobogatov et al., 2004). 

Maackia umbilicifera (Starostin, 1926) F 

Baikalia (Eubaikalia) umbilifera Starostin, 1926: 23, textfigure 23. 

● Type locality: off Bolshoi Ushkanyi Island, 50 m [Baikal]. 

● Holotype: ZMMU No. Lc-1397. 

● Distribution: type locality, 10-50 m (Sitnikova et al., 2004). 

Maackia variesculpta (Lindholm, 1909) F 

Baikalia (Baikalia) variesculpta Lindholm, 1909: 48, pl. 3, figs. 5-10. 




Maackia werestschagini (Kozhov, 1936) F 

Baicalia (Eubaicalia) werestschagini Kozhov, 1936: 55, 63-64, 307, pl. 3, figs. 41-42. 

● Type locality: 4 localities in Maloe More (border between central and northern Baikal), 2-3 m. 


● Distribution: Baikal: Maloe More, 2-3 m (Sitnikova et al., 2004).

Parabaikalia Lindholm, 1909 

Type species: Leucosia florii W. Dybowski, 1875 (OD) 

Parabaikalia dubiosa (Kozhov, 1936) F 

Baicalia (Parabaicalia) elata var. dubiosa Kozhov, 1936: 76-77, 307, pl. 5, fig. 10, 11. 

● Type locality: Boguchanskaya, Frolikha and Ayaya inlets, near Kurla and Urbikan capes, Barguzin and 

Chivyrkuiskij bays (northern Baikal), 5-15 m. 


● Distribution: northern Baikal, 4-20 m (Sitnikova et al., 2004). 

Parabaikalia elata (W. Dybowski, 1875) F 

Leucosia angarensis var. elata W. Dybowski, 1875: 43-45, pl. 4, figs. 18-25, pl. 6, fig. 5a-d. 

● Type locality: originally "Baikal-See ... 10 Metern"; Kultuk (SW coast of southern Baikal) (restricted Sitnikova 

et al., 2004). 



Parabaikalia florii (W. Dybowski, 1875) F 

Leucosia florii W. Dybowski, 1875: 39, pl. 3, figs. 1-3; pl. 6, fig. 3a-d; pl. 7, fig. 8. 

● Type locality: originally “"Baikals"; Kultuk (SW coast of southern Baikal) (restricted Sitnikova et al., 

2004). 


● Distribution: southern and central Baikal, 1.5-100 m (Sitnikova et al., 2004). 

Parabaikalia kobeltiana (Lindholm, 1909) F 

Baikalia (Parabaikalia) kobeltiana Lindholm, 1909: 51, text-figure 1. 



● Distribution: western coast of southern and central Baikal, 2-30 m (Sitnikova et al., 2004). 

Parabaikalia milaschevitschi (Lindholm, 1909) F 

[= Baikalia florii sensu Milaschewitsch, non Dybowski, 1875) in Korotnev, 1901, part.] 

Baikalia (Parabaikalia) milaschevitschi Lindholm, 1909: 51, text-figure 2. 

● Type locality: Bezymannaya Bay near Goryachinsk (E coast of central Baikal). 



Parabaikalia oviformis (W. Dybowski, 1875) F 

Leucosia oviformis W. Dybowski, 1875: 42, pl. 3, figs. 8–9. 


et al., 2004). 



Pseudobaikalia Lindholm, 1909 

Type species: Baikalia jentteriana Linholm, 1909 (OD) 

Pseudobaikalia cancellata (Lindholm, 1909) F 

Baikalia (Pseudobaikalia) cancellata Lindholm, 1909: 59, pl. 1, figs. 33-34. 

● Type locality: Dagarskaya Guba (NE coast of northern Baikal). 


● Distribution: type locality, 19-98 m (Sitnikova et al., 2004).

Pseudobaikalia contabulata (W. Dybowski, 1875) F 

Ligea contabulata W. Dybowski, 1875: 47-48, pl. 3, figs. 38-43; pl. 6, fig. 13a-d. 


et al., 2004). 

● Syntypes: Zoological Museum of Lvov University. 

● Distribution: Baikal, 1.5-200 m (Sitnikova et al., 2004). 

Pseudobaikalia elegantula (Lindholm, 1909) F 

Baikalia (Pseudobaikalia) elegantula Lindholm, 1909: 57, pl. 1, figs. 41-42. 

● Type locality: off Kurma, Maloe More (Baikal). 


● Distribution: Maloe More, northern Baikal, 1.5-15 m (Sitnikova et al., 2004). 

Pseudobaikalia jentteriana Lindholm, 1909 F 

Baikalia (Pseudobaikalia) jentteriana Lindholm, 1909: 55, pl. 1, figs. 35-37. 



● Distribution: northern Baikal, Maloe More, 15-100 m (Sitnikova et al., 2004). 

Pseudobaikalia pulla pulla (W. Dybowski, 1875) F 

Leucosia angarensis var. pulla Dybowski, 1875: 44-45, pl. 4, fig. 26 – 34, pl. 6, fig. 6 a-d. 

● Type locality: origianally "Baikal-See ... 300 Metern"; Kultuk (SW coast of southern Baikal) (restricted 

Sitnikova et al., 2004). 



Pseudobaikalia pulla tenuicosta (Lindholm, 1909) F 

Baikalia (Pseudobaikalia) tenuicosta Lindholm, 1909: 58, pl. 2, figs. 38-40. 




Pseudobaikalia subcilindrica (Lindholm, 1909) F 

Baikalia (Pseudobaikalia) subcilindrica Lindholm, 1909: 56, pl. 1, figs. 43-45. 



● Distribution: type locality, 3.5 m (Sitnikova et al., 2004). 

Pseudobaikalia zachwatkini (Kozhov, 1936) F 

Baicalia (Pseudobaicalia) zachwatkini КKozhov, 1936: 86, pl. 3, fig. 43-48; pl. 12, fig. 6-7. 

● Type locality: Bolshiye Koty (W coast of southern Baikal). 



Pyrgobaicalia Starobogatov, 1972 

Type species: Pyrgobaicalia aenigma Starobogatov, 1972 (OD) 

● Remark: this genus was originally (but provisionally) assigned to the family Baicaliidae (= Baicaliinae), 

though its actual taxonomic position is still unclear (T. Sitnikova, pers. comm.). 

Pyrgobaicalia aenigma Starobogatov, 1972 F 

Starobogatov, 1972a: 169-170, fig. 6. 

● Type locality: Tajik SSR, Gorno-Badakhshan Autonomous Region, Dunkulduk Lake (probably carried out

from a nearby spring). 



Teratobaikalia Lindholm, 1909 

Type species: Baikalia (Teratobaikalia) macrostoma Lindholm, 1909 (OD) 

Teratobaikalia ciliata (W. Dybowski, 1875) F 

[= Dybowskia ciliata, var. typica (major, minor, pulla), angarensis – B. Dybowski, Grochmalicki, 1923] 

Ligea ciliata W. Dybowski, 1875: 49, Taf. 3, fig. 27-29, Taf. 4, fig. 10. 

● Type locality: Kultuk (SW coast of southern Baikal) (restricted Sitnikova et al., 2004). 


● Distribution: Baikal, 2-100 m; Angara River down to Irkutsk (Sitnikova et al., 2004, Starobogatov et al., 

2004). 

● Remark: This highly variable species may consist of two separate species and several subspecies, inhabiting 

different biotopes. Thus it is possible, that some of the varietes, described by B. Dybowski, Grochmalicki, 

1923 will appear to be valid taxa (Sitnikova et al., 2004). 

Teratobaikalia clandestina (Beckman et Starobogatov, 1975) F 

[= Teratobaikalia (Baikaliella) clandestina sensu Sitnikova, 1991, non Beckman, Starobogatov, 1975] 

Baicalia (Baicaliella) clandestina Beckman, Starobogatov, 1975: 97, fig. 2А. 

● Type locality: near Sljudjanka city [southern Baikal Lake], depth 240 m. 


● Distribution: type locality (Beckman, Starobogatov, 1975; Sitnikova et al., 2004). 

Teratobaikalia duthiersii duthiersii (W. Dybowski, 1875) F 

Ligea duthiersii W. Dybowski, 1875: 48-49, pl. 3, figs. 30-33, pl. 6, fig. 11a-d. 


et al., 2004). 


● Distribution: southern Baikal, 15-50 m, Angara River down to Irkutsk (Sitnikova et al., 2004, Starobogatov 

et al., 2004). 

Teratobaikalia duthiersii angarensis (B. Dybowski, 1913) F 

Baikalia (Trichiobaikalia) duthiersii var. angarensis B. Dybowski, 1913: 211, pl. 5, fig. 31a. 

● Type locality: Angara River. 


● Distribution: Angara River (Sitnikova et al., 2004). 

Teratobaikalia duthiersii pachypleura (Lindholm, 1924) F 

Baicalia (Trichiobaikalia) duthiersii var. pachypleura Lindholm, 1924а: 24. 



● Distribution: Baikal (Sitnikova et al., 2004). 

Teratobaikalia humerosa (Beckman et Starobogatov, 1975) F 

[= Teratobaikalia (Baikaliella) clandestina clandestina sensu Sitnikova, 1991, non Beckman, Starobogatov, 

1975] 

Baicalia (Baicaliella) humerosa Beckman, Starobogatov, 1975: 97, fig. 2D 

● Type locality: near Jazovka River (northeastern Baikal), depth 20-24 m. 


● Distribution: type locality, Chivyrkyiskyi bay (Sitnikova et al., 2004).

Teratobaikalia macrostoma (Lindholm, 1909) F 

[Baicalia (Teratobaicalia) macrostoma forma lyogyra Lindholm, 1924] 

Baikalia (Teratobaikalia) macrostoma Lindholm, 1909: 43, pl. 1, figs. 25-29. 



● Distribution: northern and central Baikal, Maloe More, 1-40 m (Sitnikova et al., 2004). 

Teratobaikalia nana (Lindholm, 1909) F 

Baikalia (Baikaliella) nana Lindholm, 1909: 44, pl. 1, figs. 52-55. 

● Type locality: Maloe More (border between central and northern Baikal) and Boguchanskaya Guba (northern 

Baikal) – to be checked in ZIN. 

● Lectotype (Beckman, Starobogatov, 1975): ZIN. 


Teratobaikalia producta (Kozhov, 1936) F 

Baicalia (Baicaliella) nana var. producta Kozhov, 1936: 97, pl. 5, figs. 48-49. 

● Type locality: Ayaa Bay (NE coast of northern Baikal). 

● Lectotype (Sitnikova et al., 2004): ZIN. 

● Distribution: type locality and Maloe More, 13-100 m (Sitnikova et al., 2004). 

BITHYNIIDAE Gray, 1857 

BITHYNIINAE Gray, 1857 


Bithynia Leach in Abele, 1818 

Type species: Helix tentaculata Linnaeus, 1758 (OD) 

Bithynia curta (Garnier in Picard, 1840) F 

[= Bithiynia codia Bourgignat, 1862] 

Paludina impura var. curta Garnier – Picard, 1840: 304. 



● Distribution: entire Europe, except extreme north-east, south of Western Siberia, in rivers and lakes (Starobogatov 

et al., 2004; Lazutkina, 2004). 

Bithynia decipiens (Millet, 1843) F 

Paludina decipiens Millet, 1843: pl. 63. 

● Type locality: ??? 


● Distribution: entire Europe, except extreme north-east, south of Western Siberia, in rivers and lakes (Starobogatov 


Bithynia producta Moquin-Tandon, 1855 F 

[= Paludina impura var. producta Menke, 1830, nom. nud.] 

Bithynia tentaculata var. producta Moquin-Tandon, 1855: 528, pl. 39, fig. 41. 

● Type locality: "а Agen" (France). 


● Distribution: rivers and lakes of Baltic Sea basin, upper Volga basin, southern European Russia (Starobogatov, 

Zatrawkin, 1987; Starobogatov et al., 2004).

Bithynia tentaculata (Linnaeus, 1758) F 

Helix tentaculata Linnaeus, 1758: 774. 


● Probable syntypes: UUZM Nos. 1174 a-c. 

● Distribution: entire Europe except extreme north-east, Western Siberia, Ural River, in rivers and lakes 

(Starobogatov, Zatrawkin, 1987; Pirogov et al., 1994; Starobogatov et al., 2004; Lazutkina, 2004). 

Codiella Monterosato in Locard, 1884 

Type species: Turbo leachi Sheppard, 1823 (monotypy) 

Codiella leachii (Sheppard, 1823) F 

Turbo leachii Sheppard, 1823: 152. 

● Type locality: “habitat in rivis – Found paringly in Campsey Mere, and Holbrook stream and mill-ponds”. 


● Distribution: Baltic Sea basin and upper part of Volga basin (Starobogatov, Zatrawkin, 1987; Starobogatov 

et al., 2004). 

Codiella kickxii (Westendorp, 1835) F 

[= Bithynia ventricosa Gray in Turton, 1840; Bithynia celtica Paladilhe, 1870] 

Paludina kickxii Westendorp, 1835: 337. 

● Type locality: ?? 


● Distribution: Baltic Sea basin (Starobogatov et al., 2004). 

Digyrcidum Locard, 1882 

Type species: Bythinia bourguignati Paladilhe, 1869 (monotypy) 

Digyrcidum bourguignati (Paladilhe, 1869) F 

Bythinia bourguignati Paladilhe, 1869: 225-227, pl. 19, figs. 1-3. 

● Type locality: "dans les basins du jardin Picos, près de Perpignan (Pyrénées Orientales)". 


● Distribution: south of Europe, eastward to Volga delata, in rivers and lakes (Starobogatov et al., 2004). 

Opisthorchophorus Beriozkina, Levina et Starobogatov, 1995 

Type species: Paludina troscheli Paasch, 1842 (OD) 

Opisthorchophorus abakumovae Andreeva et Starobogatov, 2001 F 

Andreeva, Starobogatov, 2001: 77-78, fig. 1. 

● Type locality: western coast of Krivoye Lake, Tarskyi area of Omsk District [Western Siberia]. 

● Holotype: ZIN, No. 1 in systematic catalogue.. 

● Distribution: Western Siberia, southern Ural, probably northern Kazakhstan, in temporary and semipermanent 

waterbodies (Andreeva, Starobogatov, 2001; Starobogatov et al., 2004). 

Opisthorchophorus baudonianus (Gassies, 1859) F 

[= Bithynia proxima Frauenfeld, 1862; Bithynia blanci Locard, 1894] 

Bithinia baudoniana Gassies, 1859: 289-290. 

● Type locality: "les fossés de la grande Lande voisine des prés salés, au Teich" (France). 


● Distribution: southern Europe, in Russia soutward from the Oka River, south of Western Siberia, in temporary 

waterbodies (Starobogatov et al., 2004; Lazutkina, 2004). 

Opisthorchophorus hispanicus (Servain, 1880) F 

[= Paludina inflata Hansen, 1845 non Villa et Villa, 1841]

Bythinia Hispanica Servain, 1880: 147. 

● Type locality: “Alluvions du Guadalaviar, près de Valence”. 

● Types: Servain’s collection has been merged with the collection of Bourguignat (G. Falkner, pers. communication). 

Possibly, types are either in MNHN or in NHMV. 

● Distribution: southern Europe and southern Western Siberia, northern Kazakhstan (Irtysh basin), in temporary 

waterbodies (Pirogov et al., 1994; Starobogatov et al., 2004). 

Opisthorchophorus troschelii (Paasch, 1842) F 

Paludina troscheli Paasch, 1842: 300-301, textfigs a-d. 

● Type locality: "bei Berlin gefundenen". 

● Lectotype (Glöer, 2003): SMF 4063/1. 

● Distribution: Europe, except northern, entire Western Siberia, Kazakhstan, Ural River (Starobogatov, 

Zatrawkin, 1987; Pirogov et al., 1994; Anistratenko, 1998b; Starobogatov et al., 2004). 

● Remark: Falkner (2003) believes that Paludina troschelii is a junior synonym of Turbo leachii Scheppard, 

1823, whereas the large form of what is treated as troschelii belongs to Paludina transsilvanica E. Bielz, 

1853. Nevertheless, Glöer (2004) denied the synonymy of Paludina troschelii and Turbo leachii and considered 

restoration of Paludina transsilvanica premature. 

Opisthorchophorus valvatoides Beriozkina et Starobogatov in Beriozkina, Levina et Starobogatov, 1995 

F 

Beriozkina, Levina, Starobogatov, 1995: 34-35, figs. 1I, 4F. 

● Type locality: Sergievka, Balashov district, Saratov Region. 

● Holotype: ZIN? In spite of the authors’ statement, there is no holotype in the ZIN collection and its current 

whereabouts is unknown. 

● Distribution: temporary waterbodies of south of eastern Europe, northern Kazakhstan (Irtysh basin) (Starobogatov 

et al., 2004). 

“Paraelona” Beriozkina et Starobogatov in Anistratenko et Stadnichenko, 1995 

Type species: Bithynia majewsky Frauenfeld, 1862 (OD) 

● Remark: Glöer & Maassen (2009: 47) have shown that the type species of the genus Paraelona Beriozkina 

et Starobogatov in Anistratenko et Stadnichenko, 1995 has been described on the base of a single specimen 

that is nothing but a juvenile specimen of Bithynia tentaculata. Thus, the genus Paraelona has no type species 

and is not a valid genus name. 

“Paraelona” boissieri (Charpentier in Küster, 1852) F 

[= Bythinia ilysaeca Locard, 1894] 

Paludina boissieri Charpentier – Küster, 1852: 35-36, pl. 7, figs. 30-32. 

● Type locality: "die Umgegend von Rom". 


● Distribution: lower part of the Dnieper, Stavropol Territory, in permanent waterbodies (Anistratenko, 

1998b; Starobogatov et al., 2004). 

“Paraelona” fausseki Beriozkina et Starobogatov in Anistratenko et Stadnichenko, 1995 F 

Paraelona fausseki Beriozkina et Starobogatov – Anistratenko et Stadnichenko, 1994[1995]: 152, 153, fig. 

130. 

● Type locality: Kuma River. 

● Holotype: ZIN? In spite of the authors’ statement, there is no holotype in the ZIN collection and its current 

whereabouts is unknown. 

● Distribution: Ciscaucasia, Rostov Region, in temporary waterbodies (Anistratenko, 1998b; Starobogatov et 

al., 2004). 

“Paraelona” majewsky (Frauenfeld, 1862) F 

[= Bythinia hellenica Kobelt, 1892]

Bithynia majewsky Frauenfeld, 1862: 1153. 

● Type locality: "Dalmatien, Kroatien, ... Plattensee in Ungarn". 

● Lectotype: NHMV. 

● Distribution: south of Europe, eastward to Volga delta, in semipermanent waterbodies (Anistratenko, 


● Remark: According to to Glöer & Maassen (2009: 47), Bithynia majewsky, has been described on the basis 

of juvenile specimen that is nothing but a juvenile specimen of Bithynia tentaculata. It is not clear which species 

Beriozkina, Levina and Starobogatov (1995) when describing “Paraelona majewsky”. 

“Paraelona” milachevitchi Beriozkina et Starobogatov in Anistratenko et Stadnichenko, 1995 F 

Paraelona milachevitchi Beriozkina et Starobogatov – Anistratenko et Stadnichenko, 1994[1995]: 151, 152, 

fig. 129. 

● Type locality: village Balki, Konka river (one of branches in the delta of Dnieper), Kherson Region, 

Ukraine. 


● Distribution: Dnieper delta and lower Ob basin waterbodies (Starobogatov et al., 2004; Andreeva, Abakumova, 

2003). 

“Paraelona” socialis (Westerlund, 1886) F 

[= ? Digyreidum renei Letourneux, 1887] 

Bythinia socialis Westerlund, 1886: 19. 

● Type locality: "Sicilien, Oreto bei Palermo". 

● Types: not traced, possibly in NMG. 

● Distribution: southern Europe, lower Ob basin waterbodies, northern and western Kazakhstan, in temporary 

waterbodies (Anistratenko, 1998b; Starobogatov et al., 2004; Andreeva, Abakumova, 2003). 

“Paraelona” sphaerica (Bourguignat in Locard, 1894) F 

Bithynia sphaerica Bourguignat – Locard, 1894:102, pl. VI, fig. 9. 

● Type locality: “le Minho près Valença” (Portugal). 

● Types. Possibly, im MNHN (fide Dance, 1986). 

● Distribution: south-eastern Europe, Ukraine (Anistratenko, Stadnichenko, 1994[1995]). 

MYSORELLINAE Annandale, 1920 


Allocinma Annandale et Prashad, 1919 

Type species: Amnicola sistanica Annandale et Prashad, 1919 (OD) 

Allocinma caspica (Westerlund, 1902) F 

Bythinia (Digycidum) servainiana var. caspica Westerlund, 1902: 45. 

● Type locality: "Kaspisches Meer an der Ostseite". 


● Distribution: described, as it was said, from the Caspian Sea off Krasnovodsk; probably lives in waterbodies 

of Bolshoi Balkhan (Turkmenistan) (Starobogatov et al., 2004). 

Boreoelona Starobogatov et Streletzkaja, 1967 

Type species: Bithynia contortrix Lindholm, 1909 (OD) 

Boreoelona caerulans caerulans (Westerlund, 1896) B 

Bithynia (Elona) caerulans Westerlund, 1896: 197. 

● Type locality: “Dshungarei” [nothern Balkhash Lake].

● Types: ZIN? 

● Distribution: Balkhash Lake (Shadin, 1952). 

Boreolona caerulans moltschanovi (Lindholm, 1915) F 

Bythinia moltschanovi Lindholm, 1915: 344-345. 

● Type locality: "Beim See Karateren" (Amu Darya Delta). 


● Distribution: type locality (Shadin, 1952; Izzatullaev, 1982). 

Boreoelona contortrix (Lindholm, 1909) F 

Bithynia contortrix Lindholm, 1909: 30, pl. 1, fig. 61, 62. 

● Type locality: Angarskyi sor of Baikal Lake. 

● Lectotype (Starobogatov, Streletzkaja, 1967): ZIN No. 1 in systematic catalogue. 

● Distribution: eastern Siberia, Amur basin, Primorje (except the north), in permanent and semipermanent 

waterbodies (Starobogatov, Streletzkaja, 1967; Starobogatov et al., 2004). 

Boreoelona ehrmanni Prozorova et Starobogatov, 1991 F 

Prozorova, Starobogatov, 1991: 137, fig. 1. 

● Type locality: lake near railway station Khabarovsk-2, Khabarovsk Territory. 


● Distribution: basin of Amur and Primorje, in temporary waterbodies (Starobogatov et al., 2004). 

Boreoelona lindholmiana (Starobogatov et Streletzkaja, 1967) F 

Bithynia lindholmiana Starobogatov, Streletzkaja, 1967: 227-228, fig. 11. 

● Type locality: Semipalatinsk Region, water bodies near the Lake Kos-Kul. 


● Distribution: Altai, in lakes and back-waters of rivers (Starobogatov, Streletzkaja, 1967; Starobogatov et 

al., 2004). 

Boreoelona sibirica (Westerlund, 1886) F 

Bythinia troscheli var. sibirica Westerlund, 1886: 18. 

● Type locality: "Sibirien". 


● Distribution: Western and eastern Siberia, north of the Far East, in lakes and smaller permanent wasterbodies 


Boreoelona ussuriensis (Ehrmann in Buettner et Ehrmann, 1927) F 

Bithinia ussuriensis Ehrmann – Buettner, Ehrmann, 1927: 242-244, pl. 12, fig. 3. 

● Type locality: Ussuri near Spasskoje. 

● Types: possibly, in SMF (fide Dance, 1986). 

● Distribution: basin of Amur and Primorje (except the north), in temporary waterbodies (Bogatov, 


Gabbiella Mandahl-Barth, 1968 

Type species: Bithynia stanleyi var. humerosa Martens, 1879 (OD) 

Gabbiella araxena Akramowski, 1970 F 

Akramowski, 1970: 255. 

● Type locality: Araks basin. 


● Distribution: Araks basin and lower Kura river (Akramowski, 1976; Starobogatov, Zatrawkin, 1987).

Gabbiella cyrea Starobogatov in Starobogatov et Zatrawkin, 1987 F 

Starobogatov, Zatrawkin, 1987: 151. 

● Type locality: Syukh-bulag River, Azerbaijan. 


● Distribution: type locality (Starobogatov, Zatrawkin, 1987). 

Digoniostoma Annandale, 1920 

Type species: Paludina cerameopoma Benson, 1830 (OD) 

Digoniostoma kashmirense (Nevill, 1884) F 

Bithynia tentaculata var. kashmirensis Nevill, 1884: 39. 

● Type locality: Kashmir, Sri Nagar. 

● Syntypes: Indian Museum, Calcutta. 

● Distribution: central Amu-Darya (Izzatullaev, 1982; Starobogatov, Zatrawkin, 1987). 

● Remark. In the mentioned publications no diagnosis, not description was provided. Therefore the 

name is nomen nudum. We were not able to trace, where the name was validated. 

Digoniostoma oxiana Izzatullaev, 1982 F 

Izzatullaev, 1982: 337-338, textfig. (B). 

● Type locality: deposits of Amu-Darya River near settlement Kerkichi, Khadzhambosky district of Turkmen 

SSR. 


● Distribution: southeast of Turkmenia, southeast of Tajikistan (Izzatullaev, 1982; Starobogatov, Zatrawkin, 

1987). 

Parafossarulus Annandale, 1924 

Type species: Paludina striatula Benson, 1842 (OD) 

● Remark: Shadin (1952) recorded Parafossarulus striatulus (Benson, 1842) from Amur basin, Primorye. 

Nevertheless, according to the authors of later publications (Bogatov, Zatravkin, 1990 [1992]; Starobogatov 

et al, 2004) the species does not inhabit the territory of Russia. Therefore we exclude it from the current version 

of the catalogue. 

Parafossarulus manchouricus (Gerstfeldt in Bourguignat, 1860) F 

Bythinia manchourica Gerstfeldt – Bourguignat, 1860a: 535, pl. 24, figs. 11-13. 

● Type locality: "le fleuve Amour et divers cours d'eau de la Sibérie méridionale". 


● Distribution: Amur basin, in rivers and lakes (Starobogatov et al., 2004). 

Parafossarulus spiridonovi Zatrawkin et Starobogatov in Zatrawkin, Dovgalev et Starobogatov, 1989 

F 

Zatrawkin, Dovgalev, Starobogatov, 1989: 77-78, fig. 3 

● Type locality: near village Kirovo, basin of Bidzan River (Jewish Autonomous Region), Khabarovsk district. 


● Distribution: type locality (Zatrawkin, Dovgalev, Starobogatov, 1989; Starobogatov et al., 2004). 

Parafossarulus sungariensis Moskvicheva in Starobogatov, Zatravkin, 1987 F 

Parafossarulus sungariensis Moskvicheva – Starobogatov, Zatravkin, 1987: 152. 

● Type locality: Sungari basin, small lake near Harbin (China). 


● Distribution: known from Sungari River, may be found also in Amur near mouth of Sungari (Starobogatov 

et al., 2004).

COCHLIOPIDAE Tryon, 1866 

LITTORIDININAE Thiele, 1928 


Kainarella Starobogatov, 1972 

Type species: Kainarella minima Starobogatov, 1972 (OD) 

Kainarella minima Starobogatov, 1972 F 

Starobogatov, 1972a: 168, fig. 4. 

● Type locality: Turkmen SSR, Charshanginskij district, hot spring Khodzha-Kajnar. 


● Distribution: type locality (Starobogatov, 1972a). 

Nurekia Izzatullaev, Sitnikova et Starobogatov, 1985 

Type species: Nurekia triculiformis Izzatullaev, Sitnikova et Starobogatov, 1985 (OD) 

Nurekia triculiformis Izzatullaev, Sitnikova et Starobogatov, 1985 F 

Izzatullaev, Sitnikova, Starobogatov, 1985: 59, fig. 3 3. 

● Type locality: South Tajikistan (southern spurs of the Vakhsh Ridge, Danga-rinskij district, vicinities 

ofAlimtaj, southward from Nurek water reservoir, rotten aryk. 


● Distribution: South Tajikistan (Izzatullaev, Sitnikova, Starobogatov, 1985). 

Pseudocaspia Starobogatov, 1972 

Type species: Caspia issykkulensis Clessin, 1894 (OD) 

Pseudocaspia issykkulensis (Clessin, 1894) F 

Caspia issykkulensis Clessin, 1894: 66. 

● Type locality: Issyk-Kul Lake. 

● Types: ZIN. 

● Distribution: Issyk-Kul Lake (Shadin, 1952) 

Pseudocaspia (?) kainarensis Starobogatov, 1972 F 





Pseudocaspia ljovuschkini Starobogatov, 1972 F 


● Type locality: Turkmen SSR, Charshanginskij district, Kaptar-Khana Cave, underground brackish-water 

pond. 



Pseudocaspia narzikulovi (Izzatullaev, 1972) F 

Pseudamnicola narzikulovi Izzatullaev, 1972: 45-46, fig. 1(2). 

● Type locality: deposits of Vaksh River in the natural Reserve “Tigrovaya balka” near jetty. 


● Distribution: type locality (Izzatullaev, 1972).

Pseudocaspia starostini Starobogatov, 1972 F 

Starobogatov, 1972a: 166-167, fig. 2. 




Pseudocaspia tadjikistanica (Izzatullaev, 1972) F 

Caspia tadjikistanica Izzatullaev, 1972: 44, fig. 1(1 а-б). 

● Type locality: deposits of Vaksh River in the natural Reserve “Tigrovaya balka” near jetty. 


● Distribution: type locality (Izzatullaev, 1972). 

HYDROBIIDAE Stimpson, 1865 

(Kantor Yu.I.; Vinarski M.V.) 

BELGRANDIINAE DE STEFANI, 1877 

Andrusovia Brusina in Westerlund, 1903 

Type species: Andrusovia dybowskii Brusina in Westerlund, 1903 (OD) 

Andrusovia andrusovi Starobogatov, 2000 M 

Starobogatov, 2000: 39, 41, fig. 1B. 

● Type locality: eastern part of the South Caspian Sea, 39°05’N, 52°35’E, 39 m. 


● Distribution: middle and southern Caspian Sea; 21-115 m (Starobogatov, 2000). 

Andrusovia brusinai Starobogatov, 2000 M 

Starobogatov, 2000: 41, fig. 1C. 

● Type locality: eastern part of the Middle Caspian Sea, 42°42,5’N, 51°32,5’E, 80 m. 



Andrusovia dybowskii Brusina in Westerlund, 1903 M 

Westerlund, 1903: 89. 

● Type locality: Caspian (probably southern part), no precise data. 



Andrusovia marina (Logvinenko et Starobogatov, 1968) M 

Horatia marina Logvinenko, Starobogatov, 1968: 382, fig. 367 (18). 

● Type locality: northern slope of the Middle Caspian Basin, 43°32,5’ N, 49°17,5’ E, 60 m. 


● Distribution: middle and southern Caspian Sea; 21-406 m (Logvinenko, Starobogatov, 1968; Starobogatov, 

2000). 

Belgrandiella A. J. Wagner, 1927 

Type species: Paladilhia (Belgrandiella) kusceri A. J. Wagner, 1914 (OD) 

Belgrandiella abchasica Starobogatov, 1962 F 

Starobogatov, 1962 : 48, fig. 1 E.

● Type locality: Nizhnjaja Shakuranskaja cave, Caucasus. 


● Distribution: type locality (Starobogatov, 1962). 

Belgrandiella caucasica Starobogatov, 1962 F 

Starobogatov, 1962 : 48, fig. 1 D. 

● Type locality: Krasnoaleksandrovskaja cave, Caucasus. 


● Distribution: type locality (Starobogatov, 1962). 

Bucharamnicola Izzatullaev, Sitnikova et Starobogatov, 1985 

Type species: Pseudamnicola bucharica Shadin, 1952 (OD) 

Bucharamnicola bucharica (Shadin, 1952) F 

Pseudamnicola bucharica Shadin, 1952: 233-234, fig. 162. 

● Type locality: springs of Hissar Ridge (Tajikistan). 

● Syntypes: ZIN. ??? 

● Distribution: springs of Hissar Ridge (Tajikistan) (Shadin, 1952; Izzatullaev et al., 1985). 

Geyeria A. J. Wagner, 1914 

Type species: Geyeria plagiostoma A. J. Wagner, 1914 (OD) 

Geyeria horatieformis Starobogatov, 1962 F 

Starobogatov, 1962: fig. 1 3 [Z]. 




Geyeria valvataeformis Starobogatov, 1962 F 

Starobogatov, 1962: 48-49, fig. 1 Ж [Zh]. 

● Type locality: Krasnoaleksandrovskaja cave, Caucasus 



Horatia Bourgignat, 1887 

Type species: Horatia klecakiana Bourgignat, 1887 (SD Westerlund, 1902) 

Horatia borutzkii Shadin, 1932 F 

Shadin, 1932: 12-13, pl. 1, fig. 1. 

● Type locality: Riogesa Cave near Kutaisi (Georgia). ??? 


● Distribution: type locality (Shadin, 1952). 

Horatia ljovuschkini Starobogatov, 1962 F 

Starobogatov, 1962: 44, fig. 1 г [G]. 




Horatia sokolovi Starobogatov, 1962 F 

Starobogatov, 1962: 43-44, fig. 1 б [B]. 


● Holotype: ZIN.


Martensamnicola Izzatullaev, Sitnikova et Starobogatov, 1985 

Type species: Hydrobia brevicula Martens, 1874 (OD) 

Martensamnicola brevicula (Martens, 1874) F 

Hydrobia (Amnicola) brevicula Martens, 1874: 30-31, pl. 2, fig. 28 

● Type locality: puddle near Samarkand fortress. 

● Types: not specified. 

● Distribution: Central Asia (particularly Tajikistan), springs (Shadin, 1952; Izzatullaev et al., 1985). 

Martensamnicola hissarica (Shadin, 1950) F 

Amnicola hissarica Shadin, 1950: 62, fig. 1. 

● Type locality: spring in Gazhni ravine in Hissar Ridge, 32 km north of Stalinabad, 1200 m above sea level. 

● Syntypes: ?? 

● Distribution: Tajikistan (spring on right bank of Kondar River), Uzbekistan (spring at Bakhmal, Dzhzakh 

Region) (Izzatullaev et al., 1985). 

Martensamnicola kazakhstanica Izzatullaev, Sitnikova et Starobogatov, 1985 F 

Izzatullaev et al., 1985: 55-56, fig. 3 (1). 

● Type locality: Kazakhstan, Chimkent Region, spring near Vannovka village. 


● Distribution: type locality (Izzatullaev et al., 1985). 

Paladilhiopsis Pavlović, 1913 

Type species: Paladilhia robiciana Clessin, 1882. 

Paladilhiopsis aculeus Starobogatov, 1962 F 

Starobogatov, 1962: 47, fig. 2 G. 




Paladilhiopsis carpathica L. Soos, 1940 F 

Soos, 1940: ??? 

● Type locality: Vicinity of Hoverla Mt., Carpathian Mts. (fide Grossu, 1956). 

● Holotype: National Hungarian Museum, Budapest. 

● Distribution: Ukraine – Goverly caves, Ivano-Frankovsk district (Anistratenko, 1998b). 

Paladilhiopsis orientalis Starobogatov, 1962 F 

Starobogatov, 1962: 46-47, fig. 2 D. 




Paladilhiopsis pulcherrima Starobogatov, 1962 F 

Starobogatov, 1962: 47, fig. 2 A-B. 




Paladilhiopsis schakuranica Starobogatov, 1962 F 

Starobogatov, 1962: 47, fig. 2 Zh.




Paladilhiopsis shadini Starobogatov, 1962 F 

Starobogatov, 1962: 45-46, fig. 2V. 

● Type locality: Tzebelda cave, Caucasus. 



Paladilhiopsis subovata Starobogatov, 1962 F 

Starobogatov, 1962: 46, fig. 2E 




Sogdamnicola Izzatullaev, Sitnikova et Starobogatov, 1984 

Type species: Hydrobia pallida Martens, 1874 (OD) 

Sogdamnicola pallida (Martens, 1874) F 

Hydrobia (Amnicola) pallida Martens, 1874: 31-32, pl. 2, fig. 27. 

● Type locality: little pond in the garden in Uzgun (Zarafshan valley). 

● Syntypes: possibly, in ZMB (fide Dance, 1986). 

● Distribution: springs of Central Asia (Shadin, 1952). 

Sogdamnicola shadini Izzatullaev, 1984 F 

Izzatullaev, 1984b: 173, text. fig. a, b. 

● Type locality: Urgut near Samarkand, spring [Central Asia]. 


● Distribution: springs of Central Asia (Izzatullaev, 1984b). 

“Tadzhikamnicola Izzatullaev, 2002” 

Type species: ??????? 

● Remark: this name appeared in the list of taxa, described by Z.I. Izzatullaev in a brochure dedicated to his 

anniversary. Nevertheless we were not able to find the description of this genus in any of his publications. 

Therefore it is not impossible that the taxon has not been formally described. 

“Tadzhikamnicola” likharevi (Izzatullaev, 1973) F 

Pseudamnicola likharevi Izzatullaev, 1973b: 77, fig. 1B. 

● Type locality: flood plain of the river Surhob, vicinities of the soviet farm “Muminabad”, Kulyabskij district, 

spring [Tajikistan]. 


● Distribution: Tajikistan (Izzatullaev, 1973b). 

“Tadzhikamnicola” pavlovskii (Izzatullaev, 1973) F 

Pseudamnicola pavlovskii Izzatullaev, 1973b: 76-77, Fig. 1A. 

● Type locality: flood plain of the river Surhob, vicinities of the soviet farm “Muminabad”, Kulyabskij district, 

spring [ Tajikistan]. 


● Distribution: Tajikistan (Izzatullaev, 1973b). 

Turkmenamnicola Izzatullaev, Sitnikova et Starobogatov, 1985 

Type species: Pseudamnicola lindholmi Shadin, 1952 (OD)

Turkmenamnicola lindholmi (Shadin, 1952) F 

Pseudamnicola lindholmi Shadin, 1952: 231-232, fig. 162. 

● Type locality: Shar-Arab spring near Kushka (Turkmenia). 


● Distribution: type locality (Shadin, 1952; Izzatullaev et al., 1985). 

Turkmenamnicola raddei (Boettger, 1889) F 

Pseudamnicola lindholmi Boettger, 1889: 970, pl. 2, fig. 6 a-c. 

● Type locality: “bei Chodsha-Kala”. 

● Types: "Lectotype" SMF No. 4160, "paralectotypes" SMF No.s 4161/1, 93402/4. 

● Distribution: Turkmenia (Shadin, 1952). 

Turkmenamnicola smaragdovae (Abrikosov et Tzvetkov, 1945) F 

Pseudamnicola smaragdovae Abrikosov, Tzvetkov, 1945: 207-209, fig. 1. 

● Type locality: sources of the Sekiz-yab River near Germab (Mikhailovskoe) village, south of Geok-tepe. 

● Holotype: ZMMU (not found). 

● Distribution: upper reaches of Sekib-Yab [sic] (vicinities of Germab village) and Sherlok rivers, Turkmenia 

(Shadin, 1952). 

● Remark: The holotype was mentioned in the original description (as "type") but without any distinguishing 

characters. The ZMMU collection contains 9 specimens from the type series (out of 44 originally mentioned), 

and there is no indication of the type status on the label. Therefore, we cannot exclude that one of them is in 

fact the holotype, though there is no possibility to identify it as such. 

Valvatamnicola Izzatullaev, Sitnikova et Starobogatov, 1984 

Type species: Pseudamnicola archangelskii Shadin, 1952 (OD) 

Valvatamnicola archangelskii (Shadin, 1952) F 

Pseudamnicola archangelskii Shadin, 1952: 232-233, fig. 161. 

● Type locality: spring at the road from Vozadil village to Shakhmardan village (Alai mountain system, Uzbekistan). 

● Syntypes: ZIN. ??? 


Valvatamnicola schahimardanica Izzatullaev, 1984 

Izzatullaev, 1984b: 174, textfig. v (letter). 

● Type locality: spring between Shahimardan and Vyadil, Fergan Area, UzbekSSR. 

● Holotype: ZIN, No.1 in systematic catalogue. 

● Distribution: type locality (Izzatullaev, 1984b). 

HYDROBIINAE Stimpson, 1865 

Pseudamnicola Paulucci, 1878 

Type species: Paludina macrostoma Küster, 1852 (SD Wagner, 1927) 

Pseudamnicola brusiniana (Clessin et W. Dybowski in W. Dybowski, 1888) M 

[= Lithoglyphus caspius “Kryn” Grimm, 1876 (partim); Zagrabica sphaerion “Mouss” Lindholm, 1924] 

Zagrabica brusiniana Clessin, W. Dybowski — W. Dybowski, 1888: 52-53, pl. 2, fig. 7. 

● Type locality: “Kaspi-See”. 


● Distribution: southern and middle Caspian Sea; 0-200 m (Logvinenko, Starobogatov, 1968).

Pseudamnicola depressispira Logvinenko et Starobogatov, 1968 M 

Logvinenko, Starobogatov, 1968: 381, fig. 367 (14). 

● Type locality: western part of southern Caspian Sea, northward of Kuraginsky Kamen Island, 81 m. 

● Holotype: not traced, may be in ZIN. 

● Distribution: type locality, only empty shells (Logvinenko, Starobogatov, 1968). 

● Remark: no identified material was found in ZIN. 

Pseudamnicola exigua (Eichwald, 1838) M 

Paludina exigua Eichwald, 1838: 152-153. 

● Type locality: “cum reliquis Paludinis, fossilis”. 


● Distribution: northern and eastern Caspian Sea, only empty shells (Logvinenko, Starobogatov, 1968). 

● Remark: no identified material was found in collections of ZIN. 

Pseudamnicola prasina Rosen, 1903 F 

Rosen, 1903: 182. 

● Type locality: Obi-Garm (Tajikistan). 

● Syntypes: ZMMU. 


Pseudamnicola sphaerion (Mousson, 1863) M 

[= Lithoglyphus caspius “Kryn” Clessin et W. Dybowski in W. Dybowski, 1888; Lithoglyphus exiguus sensu 

Lindholm, 1924, non Paludina exigua Eichwald, 1838] 

Bithynia sphaerion Mousson, 1863: 89-30. 

● Type locality: “mer Caspienne, а Talysch”. 


● Distribution: southern Caspian Sea; 0-35 m, only empty shells (Logvinenko, Starobogatov, 1968). 


PYRGULINAE Brusina, 1882 (1869) 

Caspia Clessin et W. Dybowski, 1888 

Type species: Caspia baerii Clessin et W. Dybowski in W. Dybowski, 1888 (SD Wenz, 1926) 

● Remark: The genus contains both marine, brackishwater and few freshwater species. In order, not to split 

the genus, we provide description of all the species. 

Caspia baerii Clessin et W. Dybowski in W. Dybowski, 1888 M 

W. Dybowski, 1888: 36-37, pl. 3, fig. 4a-b. 


● Syntypes: not traced, not in ZIN. 



Caspia brotzkajae Starobogatov in Anistratenko et Prisjazhnjuk, 1992 M 

Caspia (Clathrocaspia) brotzkajae Starоbogatov — Anistratenko, Prisjazhnjuk, 1992: 18, fig. 2, a. 

● Type locality: Dagestan shore of Caspian Sea. 

● Holotype: not traced, should be in ZIN. 

● Distribution: Caspian Sea — type locality; about 60 m (Anistratenko, Prisjazhnjuk, 1992). 

Caspia gaillardi (Tadjalli-Pour, 1977) M 

Pyrgula (Caspia) gaillardi Tadjalli-Pour, 1977: 107, fig. 8. 

● Type locality: Astara-Hachtpar, 0-30 m. 

● Holotype: Faculté de Médecine Vétérinaire de l’Université de Téhéran.

● Distribution: mentioned by Sitnikova and Starobogatov (1998) among material for morphological study 

collected in southern and middle Caspian Sea. 


Caspia gmelinii Clessin et W. Dybowski in W. Dybowski, 1888 M 

W. Dybowski, 1888: 37-38, pl. 3, fig. 7a-b. 





Caspia knipowitchi Makarov, 1938 M,B 

Caspia gmelini var. knipowitchi Makarov, 1938: 1058, fig. 1. 

● Type locality: Dnieper River near Kherson City. 


● Distribution: middle Caspian Sea, 50-75 m; mouth of Dnieper river, limans of Black Sea, delta of Don 

River (Logvinenko, Starobogatov, 1968; Anistratenko, 1998a). 

Caspia logvinenkoi (Golikov et Starobogatov, 1966) F 

Pyrgula logvinenkoi Golikov, Starobogatov, 1966: 354, fig. 1 (7). 

● Type locality: delta of Don River. 


● Distribution: delta of Don River (Golikov, Starobogatov, 1966; Anistratenko, 1998a; Starobogatov et al., 

2004). 

Caspia makarovi (Golikov et Starobogatov, 1966) F,B 

[= Pyrgula gmelini aluschtensis Golikov et Starobogatov, 1972] 

Pyrgula makarovi Golikov, Starobogatov, 1966: 353- 354, fig. 1 (5). 

● Type locality: liman of Dnieper. 


● Distribution: off mouths of Dnieper, Don, Dniester rivers and limans of the Black Sea; 0-3 m (Golikov, 

Starobogatov, 1966; Anistratenko, 1998a). 

Caspia stanislavi Alexenko et Starobogatov, 1987 F,B 

Caspia gmelini stanislavi Alexenko, Starobogatov, 1987: 33, fig. 1. 

● Type locality: delta of Dnieper River, sta. 1. 

● Holotype: should be in ZIN, not traced. 

● Distribution: lower reaches of Dnieper river and Dnieper-Bug liman (Anistratenko, 1998a; Starobogatov et 

al., 2004). 


Caspiohydrobia Starobogatov, 1970 

Type species: Pyrgohydrobia eichwaldiana Golikov et Starobogatov, 1967 (OD) 

Caspiohydrobia aralensis Starobogatov et Andreeva, 1981 B 

Starobogatov, Andreeva, 1981: 31, fig. 5. 

● Type locality: Aral Sea, Sary-Chaganak Bay, 2.5 m. 


● Distribution: Aral Sea, Shukuroi Lake, Mnogosopochnoe Lake (Kokchetav Region), brackish-water bodies 

of Turgai depression (upper Ubagan River Basin, Kustanai Region, Kazakhstan) (Starobogatov, Andreeva, 

1981; Andreeva, 1987).

Caspiohydrobia behningi Starobogatov et Andreeva, 1981 B 


● Type locality: Aral Sea, central part of Bolshoye More, 21 m. 


● Distribution: Aral Sea, brackish-water bodies of Turgai depression (upper Ubagan River Basin, Kustanai 

Region, Kazakhstan) (Starobogatov, Andreeva, 1981; Andreeva, 1987). 

Caspiohydrobia bergi Starobogatov et Andreeva, 1981 B 


● Type locality: Aral Sea, north-east from Vozrozhdenie Island, 8 m. 


● Distribution: Aral Sea, Mnogosopochnoe Lake (Kokchetav Region), brackish-water bodies of Turgai depression 

(upper Ubagan River Basin, Kustanai Region) (Starobogatov, Andreeva, 1981; Andreeva, 1987). 

Caspiohydrobia borealis Andreeva et Frolova, 1989 B 

Andreeva, Frolova, 1989: 134-135, fig. a. 

● Type locality: Ubagan River, near exit from the Lake Kushmurun (Kustanaiskaya District) [Kazakhstan]. 


● Distribution: northern and central Kazakhstan (Andreeva, Frolova, 1989). 

Caspiohydrobia chrysopsis (Kolesnikov, 1947) M 

Hydrobia chrysopsis Kolesnikov, 1947: 106, 110. 

● Type locality: not specified (Caspian Sea – from the title). 


● Distribution: southern Caspian Sea; 0-10 m; Aral Sea (Logvinenko, Starobogatov, 1968; Andreeva, 1987). 

Caspiohydrobia conica Logvinenko et Starobogatov, 1968 M 

Logvinenko, Starobogatov, 1968: 346-348, fig. 358 (3). 

● Type locality: northern, middle and southern Caspian Sea, 0-35 m. 

● Possible syntypes: ZIN 4314/5, 4315/6. 

● Distribution: Caspian Sea, down to 35 m; Aral Sea, 10-12 m (Starobogatov, 1974). 

● Remark: Sysoev, Kantor (1992) erroneously cited holotype of the species. Since holotype was never mentioned 

in the original description, their statement was false. 

Caspiohydrobia coniformis Starobogatov et Izzatullaev, 1974 B 

Starobogatov, Izzatullaev, 1974: 933-934, fig. 1. 

● Type locality: river sediments near Laur village, Dangarinskij district, Tajik SSR, coll. Z. Izzatullaev, 

25.6.1971. 


● Distribution: brackish waters of Tajikistan (Andreeva, 1987). 

Caspiohydrobia convexa (Logvinenko et Starobogatov in Golikov et Starobogatov, 1966) M,B 

Pyrgohydrobia convexa Logvinenko et Starobogatov — Golikov, Starobogatov, 1966: 353, fig. 1 (2, 3). 

● Type locality: Great Kirov Bay, Caspian Sea. 


● Distribution: southern Caspian Sea, limans of Black and Azov seas, Aral Sea; 0-10 m (Golikov, Starobogatov, 

1966; Logvinenko, Starobogatov, 1968; Andreeva, 1987). 

Caspiohydrobia curta (Logvinenko et Starobogatov, 1968) M,B 

Pyrgohydrobia curta Logvinenko, Starobogatov, 1968: 348, fig. 358 (7). 

● Type locality: not specified. 


● Distribution: Caspian Sea, 11-40 m; Aral Sea (Logvinenko, Starobogatov, 1968; Andreeva, 1987).


Caspiohydrobia cylindrica (Logvinenko et Starobogatov, 1968) M,B 

(Plate 42 K) 

Pyrgohydrobia cylindrica Logvinenko, Starobogatov, 

1968: 350, fig. 358 (13). 


● Syntypes: not traced, may be in ZIN 

● Distribution: Caspian Sea; 0-35 m; Aral Sea (Logvinenko, Starobogatov, 1968; Andreeva, 1987). 


in the original description, their statement was false. No identified material was found in 

ZIN. 

Caspiohydrobia dubia (Logvinenko et Starobogatov, 1968) M,B 

Pyrgohydrobia dubia Logvinenko, Starobogatov, 1968: 348, fig. 358 (6). 

● Type locality: not specified, Caspian Sea (from the title). 

● Holotype: not traced in ZIN. 

● Distribution: Caspian Sea, 0-40 m; Aral Sea (Logvinenko, Starobogatov, 1968; Andreeva, 1987). 

● Remark: there is a single sample of the species in ZIN collections, not from a Caspian Sea. 

Caspiohydrobia eichwaldiana (Golikov et Starobogatov, 1966) M,B 

[= Paludina pusilla Eichwald, 1830, non Basterot, 1825; Littorinella acuta “Al. Br.” Eichwald, 1855; Hydrobia 

pusilla (“Eichwald”) W. Dybowski, 1888; Hydrobia ventrosa auctt., non Montagu, 1803; Hydrobia ventrosa 

var. Evanescens Kolesnikov, 1947, non Hydrobia evanescens Guerne, 1880] 

Pyrgohydrobia eichwaldiana Golikov, Starobogatov, 1966: 353, fig. 1 (4). 

● Type locality: Kuyalnik liman near Odessa, Black Sea. 


● Distribution: southern Caspian Sea, limans of Black and Azov seas; 0-35 m (Golikov, Starobogatov, 1966; 

Logvinenko, Starobogatov, 1968). 

Caspiohydrobia elongata Starobogatov et Izzatullaev, 1974 B 

Starobogatov, Izzatullaev, 1974: 934, fig. 2. 

● Type locality: river sediments near Laur village, Dangarinskij district, Tajik SSR, coll. Z. Izzatullaev, 

25.6.1971. 



Caspiohydrobia gemmata (Kolesnikov, 1947) M,B 

Hydrobia grimmi var. gemmata Kolesnikov, 1947: 106, 110. 


● Types: not traced, not found in ZIN. 

● Distribution: northern, middle and southern Caspian Sea; 0-50 m; Aral Sea (Logvinenko, Starobogatov, 

1968; Andreeva, 1987). 

Caspiohydrobia grimmi (Clessin in W. Dybowski, 1888) M,B 

Hydrobia grimmi Clessin — W. Dybowski, 1888: 55-56, pl. 3, fig. 2. 


● Syntypes: not traced, not in ZIN. 

● Distribution: southern Caspian Sea, 25-40 m; Aral Sea (Logvinenko, Starobogatov, 1968; Andreeva, 1987). 

Caspiohydrobia husainovae (Starobogatov, 1974) B 

Pyrgohydrobia husainovae Starobogatov, 1974: 257, fig. 227. 

● Type locality: about 20 km northwestward of Obruchev Island (Aral Sea), 20 m.


● Distribution: Aral Sea, Shukuroi Lake, waterbodies of Karasu-Shiderty River (Pavlodar Region, Kazakhstan) 

(Starobogatov, 1974; Andreeva, 1987). 

Caspiohydrobia johanseni Frolova, 1984 B 

Frolova, 1984: 46. 

● Type locality: lower stream of Karasu-Shiderty River, Pavlodarskaja district, Kominternovskij region, 5 km 

from Zagotskot settlement. 

● Lectotype (sic!): ZIN, No. 6 in systematic catalogue. 

● Distribution: northern and central Kazakhstan, brackish-water bodies of Turgai depression (upper Ubagan 

River Basin, Kustanai Region, Kazakhstan) (Frolova, 1984; Andreeva, 1987). 

● Remark: shell of lectotype is nearly dissolved in acid alcohol and is unrecognisable. 

Caspiohydrobia kazakhstanica Starobogatov et Andreeva, 1981 B 


● Type locality: Aral Sea, central part of Bolshoye More, 19 m. 


● Distribution: Aral Sea; waterbodies of Karasu-Shiderty River (Pavlodar Region), brackish-water bodies of 

Turgai depression (upper Ubagan River Basin, Kustanai Region) (Starobogatov, Andreeva, 1981; Andreeva, 

1987). 

Caspiohydrobia ljaurica Starobogatov et Izzatullaev, 1974 B 


● Type locality: river sediments near Laur village, Dangarinskij district, Tajik SSR. 



Caspiohydrobia nikitinskii Starobogatov et Andreeva, 1981 B 


● Type locality: Aral Sea, off Baigubekmuryn Cape, 4.8 m. 


● Distribution: Aral Sea, brackish-water bodies of Turgai depression (upper Ubagan River Basin, Kustanai 

Region, Kazakhstan) (Starobogatov, Andreeva, 1981; Andreeva, 1987). 

Caspiohydrobia nikolskii Starobogatov et Andreeva, 1981 B 

Starobogatov, Andreeva, 1981: 33-34, fig. 8. 

● Type locality: Aral Sea, Arktumsuk Cape, 10 m. 


● Distribution: Aral Sea (Starobogatov, Andreeva, 1981). 

Caspiohydrobia obrutchevi Starobogatov et Andreeva, 1981 B 

Starobogatov, Andreeva, 1981: 31-32, fig. 4. 

● Type locality: Aral Sea, southeastern part of Bolshoye More, 20 km northwestward from Obruchev Island, 

20 m. 


● Distribution: Aral Sea (Starobogatov, Andreeva, 1981). 

Caspiohydrobia oviformis (Logvinenko et Starobogatov, 1968) M 

Pyrgohydrobia oviformis Logvinenko, Starobogatov, 1968: 349, fig. 358 (9). 



● Distribution: southern Caspian Sea; 0-30 m (Logvinenko, Starobogatov, 1968). 

● Remark: no identified material was found in ZIN.

Caspiohydrobia parva (Logvinenko et Starobogatov, 1968) M,B 

Pyrgohydrobia parva Logvinenko, Starobogatov, 1968: 348, fig. 358 (4). 



● Distribution: Caspian and Aral seas; 5-40 m (Logvinenko, Starobogatov, 1968; Andreeva, 1987). 


Caspiohydrobia pavlovskii Starobogatov et Izzatullaev, 1974 B 


● Type locality: river sediments near Lyaur village, Dangarinsky region, Tajikistan. 


● Distribution: type locality and Aral Sea (Starobogatov, Izzatullaev, 1974; Andreeva, 1987). 

Caspiohydrobia sidorovi Starobogatov et Andreeva, 1981 B 


● Type locality: Aral sea, Sary-Chegonak Bay, 2,5 m. 


● Distribution: Aral Sea, waterbodies of Karasu-Shiderty River (Pavlodar Region), brackish-water bodies of 

Turgai depression (upper Ubagan River Basin, Kustanai Region, Kazakhstan) (Starobogatov, Andreeva, 

1981; Andreeva, 1987). 

Caspiohydrobia sogdiana Starobogatov et Izzatullaev, 1974 B 


● Type locality: river sediments near Lyaur village, Dangarinsky region, Tajikistan. 


● Distribution: type locality; Aral Sea and brackishwater bodies of Turgai depression (upper Ubagan River 

Basin, Kustanai Region, Kazakhstan) (Starobogatov, Izzatullaev, 1974; Andreeva, 1987). 

Caspiohydrobia subconvexa (Logvinenko et Starobogatov, 1968) M,B 

Pyrgohydrobia subconvexa Logvinenko, Starobogatov, 1968: 349, fig. 358 (10). 



● Distribution: Caspian Sea, 0-10 m; Aral Sea (Logvinenko, Starobogatov, 1968; Andreeva, 1987). 


Caspiohydrobia tadzhikistanica Starobogatov et Izzatullaev, 1974 B 


● Type locality: river sediments near Laur village, Dangarinskij district, Tajik SSR. 


● Distribution: brackish waters of Tajikistan, Aral Sea (Andreeva, 1987). 

Caspiohydrobia turrita (Logvinenko et Starobogatov, 1968) M 

Pyrgohydrobia turrita Logvinenko, Starobogatov, 1968: 348, fig. 358 (5). 

● Type locality: southern Caspian Sea, 0-40 m. 

● Holotype: not traced in ZIN. 

● Distribution: Caspian Sea; 0-40 m (Logvinenko, Starobogatov, 1968). 

Euxinipyrgula Sitnikova et Starobogatov, 1999 

Type species: Pyrgula milachevitchi Golikov et Starobogatov, 1966 (OD) 

Euxinipyrgula azovica (Golikov et Starobogatov, 1966) F,B 

Pyrgula azovica Golikov, Starobogatov, 1966: 357, fig. 2 (7).

● Type locality: off Taganrog, Azov Sea. 


● Distribution: Taganrog Bay (Azov Sea), lower reaches of Dnieper River, Dniestr liman (Alexenko, Starobogatov, 

1987; Anistratenko, 1998a). 

Euxinipyrgula borysthenica (Alexenko et Starobogatov, 1987) F,B 

Turricaspia derbentina borysthenica Alexenko, Starobogatov, 1987: 34-35, fig. 6. 

● Type locality: lower reaches of Dnieper River, sta. 14. 


● Distribution: lower reaches of Dnieper River and Dnieper-Bug liman (Anistratenko, 1998; Starobogatov et 

al., 2004). 

Euxinipyrgula grigorievi (Alexenko et Starobogatov, 1987) F,B 

Turricaspia grigorievi Alexenko, Starobogatov, 1987: 35, fig. 7. 



● Distribution: Dnieper and Dniester limans, lower reaches of Dnieper River, Yalpug Lake (Anistratenko, 

1998a). 

Euxinipyrgula limanica (Golikov et Starobogatov, 1966) F,B 

Pyrgula limanica Golikov, Starobogatov, 1966: 357, fig. 2 (6). 

● Type locality: Bug River, off Nikolaev. 


● Distribution: Dnieper-Bug and Dniester limans, lower reaches of Dnieper river, Yalpug Lake (Anistratenko, 

1998a). 

Euxinipyrgula lincta (Milaschewitsch, 1908) F 

Micromelania lincta Milaschewitsch, 1908: 991. 

● Type locality: Lake Katlabuh, near Danube delta. 

● Lectotype (Golikov, Starobogatov, 1966): ZIN 4401/1. 

● Distribution: Northwest Black Sea Maritime Territory (Anistratenko, 1998a). 

Euxinipyrgula milachevitchi (Golikov et Starobogatov, 1966) B 

[= Micromelania lincta Milashewitsch, 1916 (partim)] 

Pyrgula milachevitchi Golikov, Starobogatov, 1966: 359, fig. 2 (15). 

● Type locality: off Chufutovka, Dnieper-Bug liman. 


● Distribution: limans of northern and northwestern coast of Black Sea, Taganrog Bay (Golikov, Starobogatov, 


Euxinipyrgula ostroumovi (Golikov et Starobogatov, 1966) B 

[= Micromelania lincta Milachewitch, 1916 (partim)] 

Pyrgula ostroumovi Golikov, Starobogatov, 1966: 358, fig. 2 (13). 

● Type locality: Katlabukh Lake, near delta of Danube River. 


● Distribution: salt lakes and limans of northwestern coast of Black Sea (Golikov, Starobogatov, 1966; Anistratenko, 

1998a). 

Euxinipyrgula ukrainica (Alexenko et Starobogatov, 1987) F,B 

Turricaspia meneghiniana ukrainica Alexenko et Starobogatov, 1987: 35, fig. 9. 


● Holotype: not traced, should be in ZIN.

● Distribution: Dnieper-Bug and Dniester limans, lower reaches of Dnieper River, Yalpug lake (Anistratenko, 

1998a). 

Pyrgula Cristofori et Jan, 1832 

Type species: Pyrgula annulata Jan, 1832 (monotypy) 

Pyrgula abichi Logvinenko et Starobogatov, 1968 M 

[= Caspia cincta sensu Kolesnikov, 1947 (partim), non Abich, 1859] 




● Distribution: southern and western part of middle Caspian Sea; 36-120 m (Logvinenko, Starobogatov, 

1968). 


Pyrgula aenigma Logvinenko et Starobogatov, 1968 M 


● Type locality: northward of Apsheron Peninsula, Caspian Sea, 75 m. 


● Distribution: type locality (Logvinenko, Starobogatov, 1968). 

Pyrgula bakuana (Kolesnikov, 1947) M 

Turricaspia bakuana Kolesnikov, 1947: 108, 112. 

● Type locality: not specified (Caspian Sea – from title). 


● Distribution: southern and middle Caspian Sea; 25-75 m (Logvinenko, Starobogatov, 1968). 


Pyrgula basalis basalis (B. Dybowski et J. Grochmalicki, 1915) M 

Micromelania dimidiata var. basalis B. Dybowski, Grochmalicki, 1915b: 131, pl. 3, fig. 36 a-b. 

● Type locality: Caspian Sea. 

● Lectotype (Logvinenko, Starobogatov, 1968): ZIN 4368/1. 

● Distribution: middle and southern Caspian Sea; 35- 200 m (Logvinenko, Starobogatov, 1968). 

Pyrgula basalis laticarinata Logvinenko et Starobogatov, 1968 M 

[= Micromelania dimidiata sensu W. Dybowski, 1888 (partim), non Rissoa dimidiata Eichwald, 1838] 

Pyrgula laticarinata Logvinenko, Starobogatov, 1968: 359, fig. 359 (3). 

● Type locality: southern Caspian Sea, mouth of Kura River, 39°07.5’N, 49°35’E, 88 m. 

● Lectotype (here designated): ZIN, No. 1 in systematic catalogue. 



in the original description, their statement was false. Nevertheless, the specimen No. 1 in systematic 

catalogue bears the remark “holotype” and therefore it is obvious, that the authors intended to isolate holotype, 

but this was missed in publication. Therefore we designate here the same specimen as lectotype. 

Pyrgula behningi Logvinenko et Starobogatov, 1968 M 


● Type locality: western part of southern Caspian Sea, in the vicinities of Kura River mouth, 39°05’N, 

49°48’E, 120 m. 


● Distribution: western part of southern Caspian Sea; 75-120 m, only dead specimens (Logvinenko, Starobogatov, 

1968). 

● Remark: no identified material was found in ZIN.

Pyrgula cincta (Abich, 1859) M 

[= Caspia orthii Clessin et W. Dybowski in W. Dybowski, 1888] 

Rissoa cincta Abich, 1859: 57, Taf. 2, Fig. 6. 

● Type locality: not specified [Caspian Sea from the title]. 



Pyrgula columna Logvinenko et Starobogatov, 1968 M 




● Distribution: western part of southern Caspian Sea; 25-50, only dead shells (Logvinenko, Starobogatov, 

1968). 


Pyrgula concinna Logvinenko et Starobogatov, 1968 M 

[= Hydrobia spica sensu Grimm, 1876 (partim), sensu W. Dybowski, 1888, non Eichwald, 1855; Turricaspia 

spica var. nova Kolesnikov, 1947 (partim)] 

Logvinenko et Starobogatov, 1968: 365, fig. 362 (3). 

● Type locality: middle Caspian Sea, 25-80 m. 

● Syntypes: not traced, may be in ZIN. 

● Distribution: middle Caspian Sea; 25-80 m (Logvinenko, Starobogatov, 1968). 


in the original description, their statement is false. No identified material was found in ZIN. 

Pyrgula curta (Naliwkin, 1914) M 

Micromelania curta Naliwkin, 1914: 21-22, 31, pl. 6, figs. 1-4, 7, 9-14. 

● Type locality: not stated (hill south of Bibi-Eibat, west of Shikhova village, Apsheron Peninsula, Azerbaijan 

[fossil] – from preface to paper). 

● Lectotype (Logvinenko, Starobogatov, 1968): All-Russian Scientific Research Geological Insitute, St.- Petersburg. 



Pyrgula derzhavini Logvinenko et Starobogatov, 1968 M 






Pyrgula dimidiata (Eichwald, 1838) M 

[= Micromelania dimidiata var. eucalia B. Dybowski et Grochmalicki, 1915; Micromelania dimidiata var. 

gracilis B. Dybowski et Grochmalicki, 1915; Micromelania dimidiata var. ptychophora B. Dybowski et Grochmalicki, 

1915; Micromelania dimidiata var. bicarinata B. Dybowski et Grochmalicki, 1915; Pyrgula dimidiata 

grossui Golikov et Starobogatov, 1966] 

Rissoa dimidiata Eichwald, 1838: 156. 

● Type locality: “in eodem lapide calcareo, fossilis” (in quaternary limestones, fossil, probably Daghestan). 



● Remark: Micromelania dimidiata var. eucalia B. Dybowski et Grochmalicki, 1915 was listed as a valid 

species of Turricaspia by Sitnikova and Starobogatov (1998), though without discussion of its taxonomy.

Pyrgula dubia Logvinenko et Starobogatov, 1968 M 




● Distribution: western part of middle Caspian Sea; 75-100 m, only dead shells (Logvinenko, Starobogatov, 

1968). 


Pyrgula ebersini Logvinenko et Starobogatov, 1968 M 




● Distribution: western part of middle Caspian Sea; 0-50 m, only dead shells (Logvinenko, Starobogatov, 

1968). 


Pyrgula eulimellula (B. Dybowski et J. Grochmalicki, 1915) M 

[= Turricaspia grimmi sensu Kolesnikov, 1947 (partim), non Clessin et W. Dybowski, 1888] 

Micromelania (Turricaspia) eulimellula B. Dybowski, Grochmalicki, 1915b: 123-125, pl. 3, fig. 27, a-b. 



● Distribution: middle Caspian Sea; deeper than 80 m (Logvinenko, Starobogatov, 1968). 

Pyrgula fedorovi Logvinenko et Starobogatov, 1968 M 




● Distribution: western part of middle Caspian Sea; 80 m, only dead shells (Logvinenko, Starobogatov, 

1968). 


Pyrgula grimmi (Clessin et W. Dybowski in W. Dybowski, 1888) M 

Micromelania grimmi Clessin et W. Dybowski — W. Dybowski, 1888: 27-28, pl. 1, fig. 2a-c. 

● Type locality: “Kaspiu-See”. 



Pyrgula isseli Logvinenko et Starobogatov, 1968 M 






Pyrgula kolesnikoviana Logvinenko et Starobogatov in Golikov et Starobogatov, 1966 M 

[= Rissoa conus sensu Eichwald, 1841 (partim), non Eichwald, 1838; Caspia conus sensu Kolesnikov, 1948 

(partim), non Eichwald, 1838] 

Golikov, Starobogatov, 1966: 357-358, fig. 2 (8, 9). 

● Type locality: Caspian Sea, northward of Apsheron peninsula, northwestward from “Kamni Dva Brata” 

Island, 40°47’N, 49°42’E, 30 m. 


● Distribution: southern Caspian Sea; 25-120 m (Logvinenko, Starobogatov, 1968).

Pyrgula kowalewskii (Clessin et W. Dybowski in W. Dybowski, 1888) M 

Caspia Kowalewskii Clessin et W. Dybowski — W. Dybowski, 1888: 40-41, pl. 3, fig. 9a-b. 




Pyrgula lencoranica Logvinenko et Starobogatov, 1968 M 

[= Turricaspia sieversi sensu Kolesnikov, 1947, non Clessin in W. Dybowski, 1888] 






Pyrgula lirata (B. Dybowski et J. Grochmalicki, 1915) M 

[= Micromelania spica var. incisata B. Dybowski et Grochmalicki, 1915] 

Micromelania (Turricaspia) spica var. lirata B. Dybowski, Grochmalicki, 1915b: 117, pl. 2, fig. 18. 



● Distribution: western part of middle and southern Caspian Sea; 25-50 m (Logvinenko, Starobogatov, 1968). 

Pyrgula marginata (Westerlund, 1902) M 

Nematurella marginata Westerlund, 1902: 45. 

● Type locality: “Kaspisches Meer bei Krasnojarsk”( sic!). 




Pyrgula nana Logvinenko et Starobogatov, 1968 M 

Logvinenko, Starobogatov, 1968: 379-380, fig. 367(12). 



● Distribution: western part of southern Caspian Sea; 75-120 m, only empty shells (Logvinenko, Starobogatov, 

1968). 


Pyrgula nossovi (Kolesnikov, 1947) M 

Turricaspia nossovi Kolesnikov, 1947: 108-111. 




Pyrgula pallasii (Clessin et W. Dybowski in W. Dybowski, 1888) M 

Caspia pallasii Clessin, W. Dybowski — W. Dybowski, 1888: 37, pl. 3, fig. 3a-b. 



● Distribution: Southern Caspian Sea; 50-150 m, only empty shells (Logvinenko, Starobogatov, 1968). 


Pyrgula pseudobacuana Logvinenko et Starobogatov, 1968 M 


● Type locality: not specified.


● Distribution: southern Caspian Sea; 50-80 m, only empty shells (Logvinenko, Starobogatov, 1968). 


Pyrgula pseudodimidiata (B. Dybowski et Grochmalicki, 1915) M 

Micromelania (Turricaspia) pseudodimidiata B. Dybowski, Grochmalicki, 1915b: 126-128, pl. 3, fig. 32, a-b. 

● Type locality: not stated [Caspian Sea — from the title]. 



Pyrgula pseudospica Logvinenko et Starobogatov, 1968 M 






Pyrgula pulla (B. Dybowski et Grochmalicki, 1915) M 

Micromelania (Turricaspia) caspia var. pulla B. Dybowski, Grochmalicki, 1915b: 111, pl. I, fig. 6a. 

● Type locality: no stated [Caspian Sea — from the title]. 


● Distribution: western coast of middle and southern Caspian Sea; 30-80 m (Logvinenko, Starobogatov, 

1968). 

Pyrgula rudis Logvinenko et Starobogatov, 1968 M 




● Distribution: middle and southern Caspian Sea; 50- 100 m, only empty shells (Logvinenko, Starobogatov, 

1968). 


Pyrgula schorygini Logvinenko et Starobogatov, 1968 M 


● Type locality: Caspian Sea; off Apsheron Peninsula, 40°07.5’N, 50°57.5’E, 88 m. 



“Pyrgula” sieversi (O. Boettger, 1881) F 

[= Pyrgula syriaca Pallary, 1930] 

Hydrobia sieversi O. Boettger, 1881a: 246-247, pl. 9, fig. 23. 

● Type locality: Araks River. 

● Holotype: SMF 141487. 

● Distribution: Araks River near Nakhichevan (Shadin, 1952). 

“Pyrgula” sieversi (Clessin in W. Dybowski, 1888) M 

Nematurella Sieversi Clessin — W. Dybowski, 1888: 45-46, pl. 2, fig. 1. 

● Type locality: “Kaspisee”. 


● Distribution: Caspian Sea (Shadin, 1952, as Caspiella sieversi). 

● Remark: Pyrgula sieversi (O. Boettger, 1881) and Pyrgula sieversi (Clessin in W. Dybowski, 1888) are 

secondary homonyms. We did not find any reference to these species after Shadin (1952). Since the generic

diagnostics of Pyrgulinae recently is the subject of permanent changes and corrections, it is possible that in 

time these species will be placed in separate genera and thus the secondary homonymy will be eliminated. 

Pyrgula similis Logvinenko et Starobogatov, 1968 M 

[= Micromelania grimmi Clessin et W. Dybowski in W. Dybowski, 1888 (partim)] 




● Distribution: eastern part of middle Caspian Sea; 20-50 m (Logvinenko, Starobogatov, 1968). 


Pyrgula simplex Logvinenko et Starobogatov, 1968 M 

Logvinenko, Starobogatov, 1968: 367-368, fig. 363 (4). 



● Distribution: middle Caspian Sea; 40-120 m, only empty shells (Logvinenko, Starobogatov, 1968). 


Pyrgula sowinskyi Logvinenko et Starobogatov, 1968 M 






Pyrgula turkmenica Logvinenko et Starobogatov, 1968 M 




● Distribution: eastern part of southern Caspian Sea; 80-150 m, only empty shells (Logvinenko, Starobogatov, 

1968). 


Pyrgula ulskii (Clessin et W. Dybowski in W. Dybowski, 1888) M 

Caspia ulskii Clessin, W. Dybowski — W. Dybowski, 1888: 38-39, pl. 3, fig. 8a-b. 

● Type locality: north-east of Naphtabank, middle Caspian. 

● Probable syntypes: ZIN 4608/1. 


Pyrgula uralensis Logvinenko et Starobogatov, 1968 M 




● Distribution: northern Caspian Sea (Logvinenko, Starobogatov, 1968). 


Turricaspia B. Dybowski et Grochmalicki, 1915 

Type species: Micromelania turricula B. Dybowski et Grochmalicki, 1915 (SD Wenz, 1928) 

Turricaspia andrussowi (B. Dybowski et Grochmalicki, 1915) M 

[= Hydrobia spica sensu Grimm, 1876, part., non Eichwald, 1855; Micromelania turricula var. major B. 

Dybowski et Grochmalicki, 1915; Micromelania turricula var. nova B. Dybowski et Grochmalicki, 1915 

(partim); Micromelania spica var. lordosa B. Dybowski et Grochmalicki, 1915; Micromelania spica f. striata

B. Dybowski et Grochmalicki, 1915; Micromelania elegantula B. Dybowski et Grochmalicki, 1915; Micromelania 

elegantula var. mediocincta B. Dybowski et Grochmalicki, 1915; Micromelania brusinae B. Dybowski 

et Grochmalicki, 1915; Turricaspia turricula sensu Kolesnikov, 1947 (partim)] 

Micromelania (Turricaspia) andrussowi B. Dybowski, Grochmalicki, 1915b: 125-126, pl. 3, fig. 31, a-b. 

● Type locality: no data [Caspian Sea — from the title]. 



Turricaspia astrachanica (Pirogov, 1971) F 

Pyrgula astrachanica Pirogov, 1970[1971]: 249-251, fig. 1. 

● Type locality: Volga delta. 


● Distribution: Volga delta (Starobogatov et al., 2004). 

Turricaspia bogatscheviana (Logvinenko et Starobogatov, 1968) M 

Pyrgula bogatscheviana Logvinenko et Starobogatov, 1968: 367, fig. 363 (2). 



● Distribution: western part of middle Caspian Sea; 60 m, only empty shells (Logvinenko, Starobogatov, 

1968). 


Turricaspia bogensis (Dubois in Küster, 1852) B,M 

[= Clessiniola variabilis Ilyina, 1966, part.; Pyrgula pseudotriton Golikov, Starobogatov, 1966] 

Paludina bogensis Dubois — Küster, 1852: 31. 

● Type locality: Bug River. 


● Distribution: brackish waters of northwestern Black Sea Maritime Territory (Dnieper-Bug and Dniester 

limans), Caspian Sea (Anistratenko, 1998a; Starobogatov et al., 2004). 

Turricaspia boltovskoji (Golikov et Starobogatov, 1966) B 

Pyrgula boltovskoji Golikov, Starobogatov, 1966: 357, fig. 2 (4). 

● Type locality: Azov Sea off Taganrog. 


● Distribution: Taganrog Bay (Azov Sea), lower reaches of Dnieper River (Alexenko, Starobogatov, 1987; 

Anistratenko, 1998a). 

Turricaspia borceana (Golikov et Starobogatov, 1966) M 

Pyrgula borceana Golikov, Starobogatov, 1966: 359, fig. 2 (16). 

● Type locality: off Crimea coast near Alushta, 44°47.12’N, 34°31.7’E. 


● Distribution: subfossil on phaseoline silt off Crimea coast (Golikov, Starobogatov, 1966). 

Turricaspia caspia (Eichwald, 1838) M 

[= Buliminus goebeli Westerlund, 1896; Micromelania caspia f. typica B. Dybowski et Grochmalicki, 1915; 

Micromelania caspia var. eichwaldi B. Dybowski et Grochmalicki, 1915; Micromelania caspia var. brunnea 

B. Dybowski et Grochmalicki, 1915; Micromelania caspia var. major B. Dybowski et Grochmalicki, 1915] 

Rissoa caspia Eichwald, 1838: 154-155. 

● Type locality: “in eodem lapide calcario Dagesthanico, fossilis” (in quaternary limestones, Dagestan, fossil) 

(see remark). 

● Lectotype (Alexenko, Starobogatov, 1987): ZIN, No. 1 in systematic catalogue. 

● Distribution: middle and southern Caspian Sea; 30- 150 m (Logvinenko, Starobogatov, 1968).

● Remarks: the species was described as fossil, from Quaternary deposits. In designating a lectotype, Alexenko 

and Starobogatov (1987) mentioned that the label reads “Caspian Sea”. This may indicate that the “lectotype” 

is a Recent shell and does not belong to the type series. Micromelania caspia var. brunnea B. Dybowski 

et Grochmalicki, 1915 was listed as a valid species of Turricaspia by Sitnikova and Starobogatov 

(1998), though without discussion of taxonomy. 

Turricaspia chersonica Alexenko et Starobogatov, 1987 F 

Alexenko, Starobogatov, 1987: 35-36, fig. 10. 

● Type locality: delta of Dnieper River. 


● Distribution: delta of Dnieper River (Alexenko, Starobogatov, 1987; Anistratenko, 1998a). 

Turricaspia conus conus (Eichwald, 1838) M 

Rissoa conus Eichwald, 1838: 155. 

● Type locality: “in eodem lapide calcareo, fossilis” (in Quaternary limestones, not specified where, probably 

in Daghestan). 



Turricaspia conus lindholmiana (Golikov et Starobogatov, 1966) M,B 

[= Micromelania lincta Milashevitsch, 1916 (partim)] 

Pyrgula (Caspiella) lindholmiana Golikov, Starobogatov, 1966: 357, fig 2 (5). 

● Type locality: Aksai River, Don District (Azov Sea). 


● Distribution: lower reaches of Don and Dnieper rivers, limans of Azov Sea and northwestern Black Sea 

(Golikov, Starobogatov, 1966; Alexenko, Starobogatov, 1987; Anistratenko, 1998a). 

Turricaspia crimeana (Golikov et Starobogatov, 1966) M 

Pyrgula crimeana Golikov, Starobogatov, 1966: 358, fig. 2 (10). 

● Type locality: of coast of Crimea, Black Sea (RV “Meotida”, sta. 24/229), 44°47.15’N, 35°50’20’’E, 7 

fathoms. 


● Distribution: subfossil on phaseoline silt from type locality (Golikov, Starobogatov, 1966). 

Turricaspia dagestanica (Logvinenko et Starobogatov, 1968) M 

Pyrgula dagestanica Logvinenko, Starobogatov, 1968: 361, fig. 360 (3). 



● Distribution: western coast of middle Caspian Sea; 20-30 m, only empty shells (Logvinenko, Starobogatov, 

1968). 


Turricaspia derbentina (Logvinenko et Starobogatov, 1968) M 

Pyrgula derbentina Logvinenko, Starobogatov, 1968: 374, fig. 366 (8). 



● Distribution: southern and middle Caspian Sea; 16- 131 m (Logvinenko, Starobogatov, 1968). 


Turricaspia eburnea (Logvinenko et Starobogatov, 1968) M 

Pyrgula eburnea Logvinenko, Starobogatov, 1968: 370, fig. 365 (1). 


● Holotype: not traced, may be in ZIN.

● Distribution: eastern part of southern Caspian Sea; 50-75 m (Logvinenko, Starobogatov, 1968). 


Turricaspia elegantula (Clessin et W. Dybowski in W. Dybowski, 1888) M 

Micromelania elegantula Clessin, W. Dybowski — W. Dybowski, 1888: 33, pl. 1, fig. 7 a-c. 


● Possible syntype: ZIN No. 1 in systematic catalogue. 

● Distribution: eastern part of southern Caspian Sea; 40-60 m (Logvinenko, Starobogatov, 1968). 

Turricaspia iljinae (Golikov et Starobogatov, 1966) M 

Pyrgula iljinae Golikov, Starobogatov, 1966: 358, fig. 2 (14). 

● Type locality: off coast of Crimea, near Alushta, 44°47’12’’N, 34°31’07’’E, 89-46 fathoms. 



Turricaspia ismailensis (Golikov et Starobogatov, 1966) F 

Pyrgula ismailensis Golikov, Starobogatov, 1966: 358, fig. 2 (11). 

● Type locality: delta of Danube River, lakes Yalpug and Kugurlui. 


● Distribution: northwestern Black Sea Maritime Territory (Anistratenko, 1998a). 

Turricaspia lirata marisnigri Starobogatov in Alexenko et Starobogatov, 1987 M 

[= Turricaspia elegantula sensu Golikov, Starobogatov, 1966, 1972, non Clessin et W. Dybowski in W. Dybowski, 

1888] 

Alexenko, Starobogatov, 1987: 33 (nom. nov. pro Turricaspia elegantula sensu Golikov, Starobogatov, 

1966, 1972, non Clessin et W. Dybowski in W. Dybowski, 1888). 

● Type locality: subfossil, on phaseoline silt near coast of Crimea, “Meotida”, sta. 24, tow 229. 


● Distribution: Black Sea, empty shells, may be extinct (Golikov, Starobogatov, 1972; Alexenko, Starobogatov, 

1987). 

Turricaspia martensii (Clessin et W. Dybowski in W. Dybowski, 1888) F,B 

[= Pyrgula triton sensu Logvinenko, Starobogatov, 1968; Golikov, Starobogatov, 1972 (partim), non Eichwald, 

1838; Pyrgula variabilis sensu Logvinenko et Starobogatov, 1968, Golikov et Starobogatov, 1972 (partim), 

non Eichwald, 1838] 

Clessinia Martensii Clessin, W. Dybowski — W. Dybowski, 1888: 43, pl. 2, fig. 5. 

● Type locality: “Kaspi-See bei Baku”. 


● Distribution: brackish waters of northwestern Black Sea Maritime Territory, northwestern Azov Maritime 

Territory, and Caspian Sea (Anistratenko, 1998a; Starobogatov et al., 2004). 

Turricaspia meneghiniana (Issel, 1865) M 

[= Micromelania subulata Westerlund, 1902; Micromelania caspia var. inflata B. Dybowski et Grochmalicki, 

1915] 

Bithynia meneghiniana Issel, 1865: 405, pl. 1, figs. 12, 13. 

● Type locality: Baku (fossil). 


● Distribution: Middle and southern Caspian Sea; 0-35 m (Logvinenko, Starobogatov, 1968). 

Turricaspia nevesskae (Golikov et Starobogatov, 1966) M 

Pyrgula nevesskae Golikov, Starobogatov, 1966: 359, fig. 2 (17). 

● Type locality: off Crimea coast (RV “Meotida”, sta. 4, tow 358) near Alushta, 44°47’12’’N, 34°31’07’’E, 

89-46 fathoms.



Turricaspia ovum Logvinenko et Starobogatov, 1968 M 

Pyrgula ovum Logvinenko, Starobogatov, 1968: 374, fig. 366 (9). 





Turricaspia pullula (B. Dybowski et Grochmalicki, 1915) M 

Micromelania (Turricaspia) caspia var. pullula B. Dybowski, Grochmalicki, 1915b: 111-112, pl. 1, fig. 7. 



● Distribution: western part of middle Caspian Sea; 60 m (Logvinenko, Starobogatov, 1968). 

Turricaspia sajenkovae (Logvinenko et Starobogatov, 1968) M,B 

Pyrgula sajenkovae Logvinenko, Starobogatov, 1968: 361, fig. 360 (4). 



● Distribution: northern Caspian Sea, Volga delta (Logvinenko, Starobogatov, 1968; Starobogatov et al., 

2004). 


Turricaspia spasskii (Logvinenko et Starobogatov, 1968) M 

[= Turricaspia spica sensu Kolesnikov, 1947, non Eichwald, 1855] 

Pyrgula spasskii Logvinenko, Starobogatov, 1968: 361, fig. 360 (8). 



● Distribution: western part of middle Caspian Sea; 40-70 m (Logvinenko, Starobogatov, 1968). 


Turricaspia spica (Eichwald, 1855) M,B 

[= Micromelania spica f. typica B. Dybowski et J. Grochmalicki, 1915] 

Paludina spica Eichwald, 1855: 303-304, pl. 10, figs. 8-10. 

● Type locality: “im Kaspischen Meere, am ufer der Inseln Tschetschnja, vorzüglich nordostwärts der Insel 

im Meeresgrunde”. 


● Distribution: western and northern parts of middle Caspian Sea, Volga delta; 0-30 m (Logvinenko, Starobogatov, 


Turricaspia triton (Eichwald, 1838) M,B 

[= Clessinia ahngeri Westerlund, 1902; Clessiniola variabilis sensu Kolesnikov, 1947, non Eichwald, 1838] 

Paludina triton Eichwald, 1838: 152. 

● Type locality: “in eodem lapide Daghestanico fossilis” (quaternary limestone, Daghestan). 

● Lectotype (Alexenko, Starobogatov, 1987): ZIN No. 1 in taxonomic catalogue. 

● Distribution: Caspian Sea, 20-25 m; Black Sea limans and mouths of rivers; 8-10 m (Golikov, Starobogatov, 

1966; Logvinenko, Starobogatov, 1968; Anistratenko, 1998a; Starobogatov et al., 2004). 

Turricaspia trivialis (Logvinenko et Starobogatov, 1968) M 

[= Micromelania grimmi Clessin et W. Dybowski in W. Dybowski, 1888 (partim); Clessiniola grimmi sensu 

Kolesnikov, 1947 (partim), non Clessin et W. Dybowski in W. Dybowski, 1888] 

Pyrgula trivialis Logvinenko, Starobogatov, 1968: 374-375, fig. 366 (10).





Turricaspia turricula (Clessin et W. Dybowski in W. Dybowski, 1888) M 

[= Micromelania turricula f. typica B. Dybowski et Grochmalicki, 1915] 

Micromelania turricula Clessin, W. Dybowski — W. Dybowski, 1888: 34, pl. 1, fig. 3 a-c. 


● Possible syntypes: ZIN 4574/1, 4575/2. 

● Distribution: middle and southern Caspian Sea; deeper than 60 m (Logvinenko, Starobogatov, 1968). 

Turricaspia variabilis (Eichwald, 1838) B,F 

[= Bithynia eichwaldi Krynicki, 1837, non auct.; Clessiniola martensi sensu Kolesnikov, 1947] 

Paludina variabilis Eichwald, 1838: 151-152. 

● Type locality: “in Volgae ostio prope Astrachanum, et versus mare Caspium; etiam fossilis in calcario 

lapide conglutinato recentissimo Daghestanici littoris” (Volga near Astrakhan, Caspian Sea, fossil in quaternary 

limestones in Dagestan). 


● Distribution: Caspian Sea, 20-25 m; fresh water parts of Azov and Black seas (Logvinenko, Starobogatov, 


Turricaspia vinogradovi (Logvinenko et Starobogatov, 1968) M,B 

Pyrgula vinogradovi Logvinenko, Starobogatov, 1968: 368, fig. 363 (9). 



● Distribution: northern Caspian Sea, Volga delta (Logvinenko, Starobogatov, 1968; Starobogatov et al., 

2004). 


LITHOGLYPHIDAE Troschel, 1857 

LITHOGLYPHINAE Troschel, 1857 


Lithoglyphus Megerle von Mühfeld in Hartmann, 1821 

Type species: Lithoglyphus eburneus Megerle von Mühfeld in Hartmann, 1821 (nom. nud.; = Paludina naticoides 

Férussac in Pfeiffer, 1828) (SD Herrmannsen, 1847) 

Lithoglyphus apertus (Küster, 1852) F 

Paludina aperta Küster, 1852: 45-46, pl. 9, figs. 14-18. 

● Type locality: "in der Save bei Agram in Croatien". 


● Distribution: northern Black Sea maritime area, rivers (Anistratenko, Stadnichenko, 1994 [1995]; Anistratenko, 

1998b). 

Lithoglyphus naticoides naticoides (C. Pfeiffer, 1828) F 

Paludina naticoides C.Pfeiffer, 1828: 45-46, pl. 8. fig. 1, 2, 4. 

● Type locality: “Donau bei Wien, Ungarn bei Pesth” 

● Types: probably lost. Pfeiffer’s collection was dispersed after his death (Dance, 1986). 

● Distribution: rivers of Azov-Black Sea maritime areas, entered the Volga River delta (Anistratenko, Stadnichenko, 

1994[1995]; Anistratenko, 1998b; Starobogatov et al., 2004).

Lithoglyphus naticoides berolinensis Westerlund, 1886 F 

Lithoglyphus naticoides var berolinensis Westerlund, 1886: 85. 

● Type locality: "Germania, Berlin". [lectotype label]. 

● Lectotype (Alexenko et al., 1990): ZIN systematic catalogue No. 1. 

● Distribution: rivers of Baltic Sea drainage (Alexenko, Levina, Starobogatov, 1990; Starobogatov et al., 

2004). 

Lithoglyphus pyramidatus Möllendorff, 1873 F 

Möllendorff, 1873: 59, fig. 20. 

● Type locality: River Urbas, Bosnia. 


● Distribution: Ukraine except Carpathians, delta of Volga river, in rivers and lakes]; Anistratenko, 1998b; 

Starobogatov et al., 2004); outside former USSR the species is found in waterbodies of northern Balkan Peninsula 

and north-western Anatolia (Glöer, 2002). 

● Remark. This species until recently was identified as L. fuscus (Pfeiffer, 1828) (Anistratenko, Stadnichenko, 

1994[1995]). 

Shadinia Akramowski, 1976 

Type species: Pyrgula terpoghassiani Shadin, 1952 (OD) 

Shadinia akramowskii (Shadin, 1952) F 

Hydrobia akramowskii Shadin, 1952: 226, fig. 149. 

● Type locality: spring in Dzhabachalu village, Zangibassar District, Armenia. 


● Distribution: Armenia (Shadin, 1952). 

Shadinia terpoghassiani (Shadin, 1952) F 

[= Pyrgula terpoghassiani Akramowski, 1952, nom. nud.] 

Pyrgula terpoghassiani Shadin, 1952: 227, fig. 151 (by indication). 

● Type locality: Aiger-Lich Lake (Armenia). 


● Distribution: Armenia, Araks River Basin, Azerbajdzhan (Akramowski, 1976). 

BENEDICTIINAE Clessin, 1880 


Benedictia W. Dybowski, 1875 

Type species: Benedictia fragilis W. Dybowski, 1875 (OD) 

Benedictia baicalensis (Gerstfeldt, 1859) F 

[= Benedictia lindholmi Kozhov, 1929] 

Paludina baicalensis Gerstfeldt, 1859: 6, fig. 8. 

● Type locality: Kultuk, 3 fathoms (southern Baikal). 

● Neotype (Sitnikova, 1987): ZIN. 

● Distribution: Baikal Lake, 1.5-100 m (Sitnikova et al., 2004). 

Benedictia distinguenda distinguenda Lindholm, 1924 F 

Benedictia fragilis distinguenda Lindholm, 1924d: 22. 

● Type locality: Maloe More, "Kobylya Golova" (Baikal). 

● Lectotype (Sitnikova, 1987??): ZIN. 

● Distribution: northern Baikal Lake, Maloe More, 20-200 m (Sitnikova et al., 2004).

Benedictia distinguenda lamuana Sitnikova, 1987 F 

Sitnikova, 1987: 1473, fig. 3, 5. 

● Type locality: of the mouth of Bolshaya Kotinka river (western coast of southern Baikal). 


● Distribution: southern Baikal Lake, 20-200 m (Sitnikova et al., 2004). 

Benedictia fragilis W. Dybowski, 1875 F 

Benedictia fragilis W. Dybowski, 1875: 5-21, pl. 1, figs. 1–5; pl. 5, figs. 1–16; pl. 7, figs. 17–19; pl. 8, figs. 

1–4. 

● Type locality: Kultuk [restricted by Sitnikova et al., 2004]. 

● Lectotype (Sitnikova, 1987): ZIN. 

● Distribution: southern and central Baikal Lake, 30-1300 m (Sitnikova et al., 2004). 

Benedictia kotyensis Matiokin, Dzuban et Sitnikova, 1988 F 

Matiokin et al., 1988: 123-134, fig. 6 (1-3). 

● Type locality: at the mouth of Bolshie Kotinki River, 20-30 m (southern Baikal). 



Benedictia limnaeoides limnaeoides (Schrenck, 1867) F 

Paludina limnaeoides Schrenck, 1867: 619, pl. 26, figs. 2- 6. 

● Type locality: Amur River, Bureya mountains [erroneous label – Sitnikova et al., 2004]. 



Benedictia limnaeoides ongurensis Kozhov, 1936 F 

Benedictia limnaeoides var. ongurensis Kozhov, 1936: 44, pl. 2, figs. 30-32/ 

● Type locality: Olhonskiye Vorota, 11.5 fathoms (Baikal). 


● Distribution: northern Baikal Lake, Maloe More, 5-120 m (Sitnikova et al., 2004). 

Benedictia limnaeoides litoralis Kozhov, 1936 F 

Benedictia limnaeoides var. litoralis Kozhov, 1936: 45, pl. 1, figs. 11, 27. 

● Type locality: Boguchanskaya Inlet, 9.5 m (northern Baikal). 


● Distribution: Baikal Lake, 10-60 m (Sitnikova et al., 2004). 

Benedictia maxima maxima (W. Dybowski, 1875) F 

Hydrobia maxima W. Dybowski, 1875: 27, pl. 1, fig. 24. 

● Type locality: Kultuk (southern Baikal) (restricted Sitnikova et al., 2004). 

● Lectotype (Beckman, Starobogatov, 1975): ZIN. 


Benedictia maxima marisminus Sitnikova, 1987 F 

Sitnikova, 1987: 1474, fig. 4, 3. 

● Type locality: Maloe More. 


Distribution:northern Baikal Lake, Maloe More, 20-220 m (Sitnikova et al., 2004). 

Benedictia nana Beckman et Starobogatov, 1975 F 

Benedictia (Baicalocochlea) nana Beckman, Starobogatov, 1975: 101, fig. 3, G. 

● Type locality: near Sosnovka [Baikal Lake], depth 300 m. 

● Holotype: ZIN.


Benedictia pulchella pulchella Sitnikova, 1987 F 

Sitnikova, 1987: 1472, fig. 3, 8. 

● Type locality: Kultuk. 



Benedictia pulchella sarmensis Sitnikova, 1987 F 

Sitnikova, 1987: 1472, fig. 3, 7. 

● Type locality: Sarma in Maloe More [Baikal]. 


● Distribution: northern Baikal Lake, 20-220 m (Sitnikova et al., 2004). 

● Remark: After examination of the morphology and radula of B. fragilis, B. distinguenda and B. pulchella, it 

appears that the species are very similar and two latter ones are probably the local morphs of B. fragilis (Sitnikova 

et al., 2004). 

Benedictia pumyla (Lindholm, 1924) F 

Kobeltocochlea pumyla Lindholm, 1924a: 17. 

● Type locality: northern Baikal: Tukalaranga Bay. 



Benedictia shadini Beckman et Starobogatov, 1975 F 

[= Benedictia maxima (part.) sensu Shadin, 1952] 

Beckman, Starobogatov, 1975: 100-101, fig. 35 [B]. 

● Type locality: Maloe More [Baikal Lake]. 


● Distribution: Baikal Lake: Maloe More, 39-135 м (Beckman, Starobogatov, 1975). 

Kobeltocochlea Lindholm, 1909 

Type species: Hydrobia martensiana W. Dybowski, 1875 (OD) 

Kobeltocochlea falsipumyla Sitnikova, 2000 F 

Sitnikova, 2000: 105, fig.1–5 

● Type locality: Akademiya Ridge (border between central and northern Baikal). 



Kobeltocochlea lindholmiana Sitnikova, 1987 F 

Sitnikova, 1987: 1471; fig. 2, 3. 

● Type locality: Maloe Morye, Olchonskiye Vorota, Baikal. 


● Distribution: Maloe Morye, northern Baikal, 2-20 m (Sitnikova et al., 2004). 

Kobeltocochlea martensiana (W. Dybowski, 1875) F 

Hydrobia martensiana W. Dybowski, 1875: 24, pl. 1, figs. 1823. 

● Type locality: Kultuk (southern Baikal) [restricted by Sitnikova et al., 2004]. 



Kobeltocochlea olchonensis Lindholm, 1909 F 

Kobeltocochlea martensiana var.olchonensis Lindholm, 1909: 37.

● Type locality: Olchonskiye Vorota (Baikal). 


● Distribution: central and northern Baikal, 2-40 m (Sitnikova et al., 2004). 

Pseudobenedictia Sitnikova, 1987 

Type species: Kobeltocochlea michnoi Lindholm, 1929 (OD) 

Pseudobenedictia michnoi (Lindholm, 1929) F 

Kobeltocochlea michnoi Lindholm, 1929c: 316, fig. 2. 

● Type locality: Hovsogöl Lake, Mongolia. 



● Remark: Although the label, accompanying the holotype states that the specimen was collected in 

Hovsogöl Lake, it has been never found there again. 

Yaroslawiella Sitnikova, 2001 

Type species: Yaroslawiella eximia Sitnikova, 2001 (OD) 

Yaroslawiella eximia Sitnikova, 2001 F 

Sitnikova, 2001: 499, figs.1, 2, 3. 

● Type locality: Baikal, Akademia Ridge 102-157 m (border between central and northern Baikal). 

● Holotype: Limnological Institute of Siberian Branch of Russian Academy of Sciences, No. 827. 


POMATIOPSIDAE Stimpson, 1865 

TRICULINAE Annandale, 1924 


Sibirobythinella Johansen et Starobogatov, 1982 

Type species: Sibirobythinella kuznetzkiana Johansen et Starobogatov, 1982 (OD) 

Sibirobythinella almaatina Izzatullaev, Sitnikova et Starobogatov, 1985 F 

Izzatullaev et al., 1985: 56-57, fig. 3, 2. 

● Type locality: Alma-Ata Region [Kazakh SSR], Zailijskij Alatau [Ridge], spring Belbulak and vicinities of 

Talgar. 


● Distribution: Kazakhstan (Izzatullaev et al., 1985). 

Sibirobythinella kuznetzkiana Johansen et Starobogatov, 1982 F 

Johansen, Starobogatov, 1982: 1142-1144, fig. a, v. 

● Type locality: Kuznetzk Alatau Mountains [southern Siberia], "lime-tree island", stream on the right bank 

of Tom' River 2.5 km downstream from Kazyr, station 95, samples 163-165. 


● Distribution: Mountain Shoriya (Tomsk Region) (Starobogatov et al., 2004). 

CLADE HETEROBRANCHIA 

(Kantor Yu. I., Vinarski M.V., Sysoev A.V.)

VALVATOIDEA Gray, 1840 

VALVATIDAE Gray, 1840 

VALVATINAE Gray, 1840 

Cincinna Hubner, 1810 

Type species: Nerita piscinalis (Müller, 1774) (OD) 

Cincinna aliena (Westerlund, 1876) F 

Valvata aliena Westerlund, 1876a: 63, fig. 15. 

● Type locality: “Jenissei, Nischnij Inbatsk (lat. 63°` 50’)”. 

● Lectotype (Starobogatov, Streletzkaya, 1967): ZIN, No. 8 in systematic catalog. Paralectotypes in NMG, No. 

4664. 

● Distribution: rivers of Yenissei River basin, estuaries of large tributaries of Baikal, north of Western Siberia 

(Starobogatov, Streletzkaya, 1967; Starobogatov et al., 2004, Sitnikova et al., 2004; Lazutkina, 2004). 

Cincinna ambigua (Westerlund, 1873) F 

Valvata ambigua Westerlund, 1873: 439. 

● Type locality: “Sverige vid Göteborg”. 


● Distribution: rivers of Europe and south of Western Siberia (Irtysh River basin) (Anistratenko, 1998b; Starobogatov 


Cincinna amurensis Moskvicheva in Starobogatov et Zatrawkin, 1985 F 

Moskvicheva – Starobogatov, Zatrawkin, 1985: 1156-1157, fig. 1. 

● Type locality: Kutumanda River (Amur basin). 


● Distribution: Amur River basin (except its upper part), Ussuri River, Khanka Lake (Bogatov, Zatrawkin, 1990 

[1992]; Starobogatov et al., 2004). 

Cincinna antiqua (Morris in Morris et Sowerby, 1838) F 

Valvata antiqua Morris – Morris, Sowerby, 1838: 547, fig. 26. 

● Type locality: the Thames valley [from the publication title]. 


● Distribution: lakes and rivers of the basin of the Baltic Sea and North Sea, lower and central Dnieper (Anistratenko, 


● Remark: Starobogatov et al. (2004) believe that Cincinna antiqua sensu Anistratenko, 1998b is probably a 

separate species. Glöer (2002) considers this species as subspecies of Valvata piscinalis. 

Cincinna bathybia (W. Dybowski, 1886) F 

Valvata (Cincinna) bathybia W. Dybowski, 1886: 129, pl. 4, fig. 2. 

● Type locality: Kultuk (southern Baikal) (restricted in Sitnikova et al., 2004). 


● Distribution: southern and central Baikal, 50-400 (Sitnikova et al., 2004). 

Cincinna brevicula (Kozhov, 1936) F 

Valvata (Cincinna) aliena var. brevicula Kozhov, 1936: 18, pl. 1, fig. 30. 

● Type locality: Bogushanskaya Inlet (Baikal). 

● Lectotype (Sitnikova et al., 2004): ZIN.

● Distribution: basins of Yenissei, Lena, and lower Ob’ rivers, Azabachje Lake in Kamchatka, Baikal; in slowrunning 

rivers and river floodland lakes (Starobogatov et al., 2004; Sitnikova et al., 2004; Lazutkina, 2004). 

Cincinna bureensis Starobogatov et Zatrawkin, 1985 F 

Starobogatov, Zatrawkin, 1985: 1158, fig. 6. 

● Type locality: lake on the left bank of the River Bureja, near Chekunda (old) settlement, Khabarovsk Territory 

[Far East]. 


● Distribution: basins of Bureya and Manoma rivers and taiga tributaries of Ussuri River(Bogatov, Zatrawkin, 

1990 [1992]; Starobogatov et al., 2004). 

Cincinna chereshnevi Bogatov, Zatrawkin et Starobogatov in Bogatov et Zatrawkin, 1990 [1992] F 

Bogatov, Zatrawkin, 1990 [1992]: 35-36, fig. 8. а-в [a-v]. 

● Type locality: southern Chukchi Peninsula, basin of Khatyrka River, Lake Egergytgyn. 

● Holotype: ZIN No. 1 in systematic catalog. 

● Distribution: glacial lakes of southern Chukchi Peninsula (basin of Khatyrka River) and northern Okhotsk Sea 

maritime area (Starobogatov et al., 2004). 

Cincinna chersonica Chernogorenko et Starobogatov, 1987 F 

Chemogorenko, Starobogatov, 1987b: 150. 

● Type locality: mouth of Dnieper river. 


● Distribution: southern areas of eastern Europe to Altai, rivers (Anistratenko, 1998b; Starobogatov et al., 2004). 

Cincinna chishimana Prozorova et Starobogatov, 1999 F 

[= Cincinna kizakikoensis Fujita et Habe, 1991] 

Prozorova, Starobogatov, 1999: 54-55, figs. 2B, 3. 

● Type locality: Iturup Island, Reidovoe Lake, 1-5 m. 


● Distribution: lakes of Iturup Island (South Kurile Islands) (Starobogatov et al., 2004). 

Cincinna confusa (Westerlund, 1897) F 

[= Valvata stelleri W. Dybowski, 1904] 

Valvata (Cincinna) confusa Westerlund, 1897b: 130. 

● Type locality: "Sibirien. Thal des Olenek". 

● Lectotype (Starobogatov, Streletzkaya, 1967): ZIN No. 32 in systematic catalog. Paralectotypes in NMG, No. 

4656. 

● Distribution: from north of western Siberia to Kamchatka and Chukchi Peninsula, North Kurile Islands (Shumshu 

Island), in lakes and slow-running rivers (Bogatov, Zatrawkin, 1990 [1992]; Starobogatov et al., 2004). 

Cincinna contorta (Müller, 1774) F 

[Valvata piscinalis var. fluviatilis Colbeau, 1859] 

Nerita contorta Müller, 1774: 187-188. 

● Type locality: Fridrichsdal (Denmark). 

● Types: unknown. Possibly, in ZMUC. 

● Distribution: lakes of the Baltic Sea basin, central and lower parts of Dnieper basin (Anistratenko, 1998b as 

Cincinna fluviatilis; Starobogatov et al., 2004). 

● Remark: Glöer (2002) considers this species as a synonym of Valvata piscinalis antiqua. Anistratenko and 

Anistratenko (2001) cited the opinion of Ya.I.Starobogatov, that Valvata piscinalis var. fluviatilis Colbeau, 1859 

is a synonym of Cincinna contorta. Most of european malacologists under the name Valvata fluviatilis 

identified separate species, for which Starobogatov proposed a new name Cincinna (Cincinna) falsifluviatilis 

(see below).

Cincinna depressa (C. Pfeiffer, 1821) F 

[= Valvata borealis Milachevitch, 1881] 

Valvata depressa Pfeiffer, 1821: 100, pl. 4, fig. 33. 

● Type locality: "in einem schlammingen Wassergraben, unweit Hanau, bey dem Dorfe Enkheim". 

● Types: not traced. Pfeiffer’s collection was dispersed after his death (Dance, 1986). 

● Distribution: Europe and south of Western Siberia (Irtysh basin), lakes and large ponds (Anistratenko, 1998b; 

Starobogatov et al., 2004; Lazutkina, 2004). 

Cincinna dilatata (Eichwald, 1830) F 

Paludina dilatata Eichwald, 1830: 219. 

● Type locality: Quaternary deposits near Grodno. 

● Lectotype (Starobogatov et al., 1994): ZIN, № 1 in systematic catalogue. 

● Distribution: Europe, Western Siberia (Irtysh and central Ob’ rivers basins), lakes and rivers (Anistratenko, 

1998b; Starobogatov et al., 2004; Lazutkina, 2004). 

Cincinna discors (Westerlund, 1886) F 

Valvata discors Westerlund, 1886: 133. 

● Type locality: "Schweden in See Ringsjön". 


● Distribution: lakes and rivers of the Baltic Sea basin, central Dnieper basin (Anistratenko, 1998b; Starobogatov 

et al., 2004); Western Siberia (Irtysh River basins) (Lazutkina, 2004). 

● Remark: Glöer (2002) regarded C. discors as a subspecies of Valvata piscinalis (Müll.). 

Cincinna falsifluviatilis Starobogatov in Anistratenko et Anistratenko, 2001 F 

Cincinna (Cincinna) falsifluviatilis Starobogatov – Anistratenko, Anistratenko, 2001: 139-140. fig. 110 

[nom. nov. pro Valvata fluviatilis sensu Westerlund, 1886; non Colbeau, 1859] 

● Type locality: “Belgien, England, Deutschland, Russland”. 


● Distribution: Europe, in Russia – basin of the Balthis Sea, in Ukraine – in middle and lower pargt of the 

Dnieper basin, rivers (Anistratenko, Anistratenko, 2001; Starobogatov et al., 2004). 

● Remark: in Starobogatov et al., 2004 this species is attributed to 1996, which is obviously erroneous. Kantor 

and Sysoev (2005) overlooked the description of the species in Anistratenko and Anistratenko (2001) and 

therefore erroneously concluded that the name is unavailable. 

Cincinna frigida (Westerlund, 1873) F 

Valvata frigida Westerlund, 1873: 436-437. 

● Type locality: “Från Naustejaur I Pite Lappmark”. 

● Syntypes: NMG, No. 4677. 

● Distribution: northern Europe and Siberia to Okhotsk Sea basin, lakes (Starobogatov et al., 2004). 

● Remark: Glöer (2002) considers this species as a synonym of Valvata sibirica. 

Cincinna gafurovi (Izzatullaev, 1977) F 

Valvata (Cincinna) gafurovi Izzatullaev, 1977: 948-949, textfig. 

● Type locality: Tajik SSR, Gorno-Badakhmanskaya Autonomous Republic, lake Sylykty-Sai (area about 300 

m 2 ), 10 km south-westward from Kyzil-Rabat, on depth 2-3 m. 


● Distribution: Tajikistan (Izzatullaev, 1977). 

Cincinna hankensis Prozorova, 1988 F 

Prozorova, 1988b: 1736-1738. 

● Type locality: small lake near the shore of Khanka Lake. 

● Holotype: ZIN No. 1 in systematic catalogue.

● Distribution: basin of Khanka Lake and southern part of Primorje, in lakes and semipermanent waterbodies 

(Bogatov, Zatrawkin, 1990 [1992]; Starobogatov et al., 2004). 

Cincinna helicoidea (Dall, 1905) F 

Valvata lewisi helicoidea Dall, 1905: 123, pl. 2, figs. 1, 2 

● Type locality: Lake Bennett, Yukon Territory; Old Fort Yukon, Alaska; and others. 

● Syntype: USNM No. 180304. 

● Distribution: Chukchi Peninsula, lower reaches of Kolyma River, northwestern Kamchatka, Eastern Siberia 

(from Khatanga to Kolynma) and Western Siberia (lower Ob’ basin) (Bogatov, Zatrawkin, 1990 [1992]; Dolgin, 

2001; Starobogatov et al., 2004; Lazutkina, 2004). 

Cincinna hokkaidoensis (Miyadi, 1935) F 

Valvata cristata hokkaidoensis Miyadi, 1935: 61-62, Pl.3, fig. 4. 

● Type locality: Tôro-ko, Hokkaido. 

● Holotype: Ôtu Hydrobiological Station. 

● Distribution: within Russian – southern Sakhalin, outside – Hokkaido (Prozorova, Bogatov, Saenko, 2004). 

Cincinna iturupensis Prozorova in Prozorova et Starobogatov, 1998 F 

Prozorova, Starobogatov, 1998c: 61, 63, fig. 3D. 

● Type locality: Lake Dobroye, Iturup Island. 


● Distribution: lakes of Iturup (Kurile Islands) (Prozorova, Starobogatov, 1998c; Starobogatov et al., 2004). 

Cincinna japonica (Martens, 1877) F 

Valvata japonica Martens, 1877: 116. 

● Type locality: Lake Hakone, Japan. 

● Types: not traced. Possibly, in ZMB (fide Dance, 1986). 

● Distribution: lakes of southern Sakhalin (Starobogatov et al., 2004). 

Cincinna kamchatica Prozorova et Starobogatov, 1998 F 

Prozorova, Starobogatov, 1998c: 63, 65, fig. 3A. 

● Type locality: eastern Kamchatka, puddle on Bolshoi Kamchatskyi Island in the valley of Kamchatka River. 


● Distribution: Kamchatka, Kolyma basin, northern Okhotsk Sea maritime area, lakes (Starobogatov et al., 

2004). 

Cincinna kliniensis (Milaschewitsch, 1881) F 

Valvata fluviatilis var. kliniensis Milaschewitsch, 1881: 236. 

● Type locality: "Moujevo" (vicinities of Moscow). 

● Syntypes: ZIN, № 1 in systematic catalogue. 

● Distribution: Europe, south of Western Siberia, mostly in lakes and large ponds (Anistratenko, 1998b; Starobogatov 


Cincinna klucharevae Starobogatov in Starobogatov et Zatrawkin, 1985 F 


● Type locality: southern Sakhalin Island, near Bousset lagoon, Bolshoj Vavaj Lake, depth 2.4 m. 


● Distribution: lakes of southern Sakhalin Island (Bogatov, Zatrawkin, 1990[1992]; Starobogatov et al., 2004). 

Cincinna korotnevi (Lindholm, 1909) F 

Valvata (Cincinna) korotnevi Lindholm, 1909: 73, pl. 1, fig. 63. 

● Type locality: Angarskyi sor (northern Baikal). 

● Lectotype (Starobogatov, Streletzkaya, 1967): ZIN, No. 1 in systematic catalogue.

● Distribution: southern part of central Siberia, Baikal Lake (Starobogatov et al., 2004, Sitnikova et al., 2004). 

Cincinna laethmophila (Beckman et Starobogatov, 1975) F 

Valvata (Pseudomegalovalvata) Beckman, Starobogatov, 1975: 95, fig. 1D. 

● Type locality: near Listvennichnoe [Baikal Lake], depth 1380 m. 


● Distribution: type locality (Sitnikova et al., 2004). 

Cincinna lewisi (Currier, 1868) F 

Valvata lewisi Currier, 1868: 9 [new name for Valvata striata sensu Lewis, non Philippi, 1834]. 

● Type locality: Grattam, Kent (Michigan). 


● Distribution: lakes (excluding thermokarst) of southern Chukchi Peninsula (Khatyrka River basin) (Starobogatov 

et al., 2004). 

Cincinna lilljeborgi (Westerlund, 1897) F 

Valvata lilljeborgi Westerlund, 1897a: 137. 

● Type locality: "Suecia in fluvio Fyrisån ad Upsala". 

● Possible syntypes: NHG, No. 4654. 

● Distribution: lakes of the Baltic Sea basin, central part of Dnieper basin, south of Western Siberia (Anistratenko, 


Cincinna macrostoma (Steenbuch in Mörch, 1864) F 

[= Valvata macrostoma Steenbuch in Beck, 1847, nom. nud; Valvata umbilicalis Westerlund, 1878] 

Valvata macrostoma Mörch, 1864: 59. 

● Type locality: Denmark (“Talrig i en Mosegröft paa sognefogdens Lod i Rudi (Stb.); Sorösö (Stp.); Viborgsö 

(Fedd.)”. 

● Types: not traced. According to Dance (1986), Mörch collection is dispersed between ZMUC and BMNH. 

● Distribution: Europe, in temporary waterbodies (Anistratenko, 1998b; Starobogatov et al., 2004). 

Cincinna mergella (Westerlund, 1883) F 

Valvata mergella Westerlund, 1883c: 166. 

● Type locality: Port Clarence, NW America. 


● Distribution: Chukchi Peninsula: Khatyrka River basin in the south to Chaun lowland in the north (Starobogatov 

et al., 2004). 

Cincinna middendorffi Moskvicheva in Starobogatov et Zatrawkin, 1985 F 

Cincinna sibirica middendorffi Moskvicheva – Starobogatov, Zatrawkin, 1985: 1158-1159, fig. 7. 

● Type locality: Amur. 


● Distribution: small semipermanent and temporary waterbodies of Amur basin and Primorje (Starobogatov et 

al., 2004). 

Cincinna obtusa (Draparnaud, 1801) Пм 

Cyclostoma obtusum Draparnaud, 1801: 39-40. 

● Type locality: "France septemtrionale". 

● Types: not traced. Possibly, in NHMV. 

● Distribution: rivers of the Baltic Sea basin, central and lower Dnieper basin, south of Western Siberia (Anistratenko, 


Cincinna olkhonica (Beckman et Starobogatov, 1975) F 

Valvata (Pseudomegalovalvata) olkhonica Beckman, Starobogatov, 1975: 94, fig. 1V.

● Type locality: Kharin-Irgi Bay (Oikhon Gates) (Baikal Lake], depth 32-39 m, 14.9.1966. 



Cincinna pamirensis Starobogatov, 1972 F 

Valvata (Cincinna) pamirensis Starobogatov, 1972a: 170-171, fig. 9. 

● Type locality: Tajik SSR, Gorno-Badakhshan Autonomous Region, Shaimak, 7 km from Kyzyl-Ravat, warm 

spring on right bank of the Aksu River, sample 136. 



Cincinna piscinalis (Müller, 1774) F 

Nerita piscinalis Müller, 1774: 172. 


● Types: unknown, possibly in ZMUC. 

● Distribution: Europe, western Siberia, lakes and rivers (Anistratenko, 1998b; Starobogatov et al., 2004); Armenia 

(Akramowski, 1976). 

Cincinna profundicola (Beckman et Starobogatov, 1975 ) F 

Valvata (Pseudovalvata) profundicola Beckman, Starobogatov, 1975: 95, fig. 1A. 

● Type locality: near Bolsodej Cape [Baikal Lake], depth 300 m. 



Cincinna pulchella (Studer, 1820) F 

Valvata pulchella Studer, 1820: 23. 

● Type locality: “Umgebung des Bielersee’s in Wassergräben”. 

● Lectotype (Forcart, 1957): NMBE. 

● Distribution: Europe, western Siberia, in temporary waterbodies (Starobogatov et al., 2004); Armenia (Akramowski, 

1976). 

● Remark: Boeters and Falkner (1998) revealed that the species name Valvata pulchella Studer, 1820 is invalid 

and these authors have described this species as a new for science one, Valvata studeri Boeters et Falkner, 1998. 

However, the authors believe that V. studeri is distributed in the Alps only, hence this species has not to be included 

to the catalogue of continental molluscs of Russia. The revision of Eastern European valvatid species 

known previously as V. pulchella is needed. 

Cincinna saghalinensis (Miyadi, 1935) F 

Valvata pulchella saghalinensis Miyadi, 1935: 61, pl. 3, fig. 3. 

● Type locality: “along the shore of a lakelet Tyatya-numa on the western coast of South Sakhalin”. 

● Holotype: Ôtu Hydrobiological Station. 

● Distribution: small semipermanent waterbodies of southern Sakhalin (Bogatov, Zatrawkin, 1990[1992]; Starobogatov 

et al., 2004). 

Cincinna sibirica (Middendorff, 1851) F 

Valvata cristata var. sibirica Middendorff, 1851: 299. 

● Type locality: Barnaul (on the basis of the lectotype). 

● Lectotype (Prozorova, Starobogatov, 1998c): ZIN. 

● Distribution: from north of western Siberia and Altai mountains to Okhotsk Sea; Transbaikalia and Chita region, 

Baikal Lake, in lakes (Starobogatov et al., 2004, Sitnikova et al., 2004). 

Remark: the species is mentioned in CLECOM database as inhabiting Norway, Sweden and Finland (the authors 

of the CLECOM database [Falkner et al., 2001] consider C. sibirica and C. frigida to be conspecific; in the Russian 

literature, all findings of this species from Europe are attached to C. frigida).

Cincinna simusyuensis (Miyadi, 1935) F 

Valvata piscinalis simusyuensis Miyadi, 1935: 60-61, pl. 3, fig. 2. 

● Type locality: “Shumshir Island” [Kurile Islands]. 

● Lectotype (Fujita, Habe, 1991): Ôtu Hydrobiological Station. 

● Distribution: Shumshu Island (North Kurile Islands), eastern Kamchatka, lakes (Starobogatov et al., 2004). 

Cincinna sirotskii Starobogatov et Zatrawkin, 1985 F 


● Type locality: near Novyj Mir settlement, Komsomolskij district, Khabarovsk Territory (Far East), depth 0.2 

m. 

● Holotype: ZIN, No. 1 in systematic catalog. 

● Distribution: basin of Amur River (except its upper part), Ussuri and Tugur rivers, northwestern Sakhalin 


Cincinna skorikovi (Lindholm, 1911) F 

Valvata (Cincinna) skorikovi Lindholm, 1911: 299-300. 

● Type locality: 5 stations in "Newabucht bei Kronstadt". 

● Syntype: ZIN, № 1 in systematic catalogue. 

● Distribution: Europe, Black Sea maritime area, Western Siberia (Irtysh and central Ob’ rivers basins), rivers 

and lakes (Anistratenko, 1998b; Starobogatov et al., 2004; Lazutkina, 2004). 

Cincinna ssorensis (W. Dybowski, 1886) F 

Valvata (Cincinna) ssorensis W. Dybowski, 1886: 113, pl. 4, figs. 1, 3, 5. 

● Type locality: Posolskyj sor (eastern coast of Baikal). 


● Distribution: Yenissei River basin, Baikal, Lena and lower Ob’ rivers basins, Kamchatka, North Kurile Islands 

(Shumshu Island), in lakes and slow-running rivers (Bogatov, Zatrawkin, 1990 [1992]; Dolgin, 2001; Starobogatov 

et al., 2004, Sitnikova et al., 2004; Lazutkina, 2004). 

Cincinna sujfunensis Prozorova in Prozorova et Starobogatov, 1998 F 

Prozorova, Starobogatov, 1998c: 65, 67, fig. 4H. 

● Type locality: Razdolnaya River near Razdolnoye settlement [Far East]. 


● Distribution: Razdolnaya (Sujfun) River and rivers entering Khanka Lake from south-east (Starobogatov et al., 

2004). 

Cincinna tenagobia (Beckman et Starobogatov, 1975) F 

Valvata (Pseudomegalovalvata) tenagobia Beckman, Starobogatov, 1975: 93, риC. 1S. 

● Type locality: Kharin-Irgi Bay (Oikhon Gates) [Baikal Lake], depth 32-39 m. 



Cincinna tymiensis Starobogatov in Starobogatov et Zatrawkin, 1985 F 


● Type locality: Sakhalin Island, right bank of the River Tym' near Nogliki settlement. 


● Distribution: Sakhalin Island, basin of the River Tym (Bogatov, Zatrawkin, 1990[1992]; Starobogatov et al., 

2004). 

Megalovalvata Lindholm, 1909 

Type species: Valvata baicalensis Gerstfeldt, 1859 (OD)

Megalovalvata baicalensis (Gerstfeldt, 1859) F 

[= Valvata grubii W. Dybowski, 1875] 

Paludina baicalensis Gerstfeldt, 1859: 510, figs. 8-10. 

● Type locality: "Baikalsee". 

● Lectotype (Sitnikova, 1983): ZIN, № 104 in systematic catalogue. 

● Distribution: Baikal Lake, 3-50 m, Angara River (Starobogatov et al., 2004; Sitnikova et al., 2004). 

Megalovalvata demersa (Lindholm, 1909) F 

Valvata (Liratina) baicalensis var. demersa Lindholm, 1909: 79. 

● Type locality: Berchin Bay, 25 fathoms (western coast of central Baikal). 



Megalovalvata kozhovi Sitnikova, 1983 F 

Sitnikova, 1983: 35. 

● Type locality: Zavorotnaya Bay (western coast of northern Baikal). 



Megalovalvata lauta (Lindholm, 1909) F 

Valvata (Atropidina) lauta Lindholm, 1909: 74, pl. 1, figs. 68-70. 

● Type locality: Davsha Inlet, 9 fathoms (eastern coast of northern Baikal). 


● Distribution: northern Baikal Lake and eastern coast of central Baikal, 2-70 m (Sitnikova et al., 2004). 

Megalovalvata parvula (Kozhov, 1936) F 

Valvata (Megalovalvata) lauta var. parvula Kozhov, 1936: 25. 

● Type locality: off Listvennichnyi Island, 5 m (eastern coast of central Baikal). 



Megalovalvata piligera piligera (Lindholm, 1909) F 

[= Valvata (Liratina) baicalensis sensu Lindholm, 1909 and Thiele, 1929, non Gerstfeldt, 1859] 

Valvata baicalensis var. piligera Lindholm, 1909: 77. 

● Type locality: Maloe More, 26 fathoms [Baikal]. 


● Distribution: Maloe More and southern Baikal Lake, 5-40 m (Sitnikova et al., 2004). 

Megalovalvata piligera minor (Lindholm, 1909) F 

Valvata (Liratina) baicalensis f. minor Lindholm, 1909: 78. 

● Type locality: Boguchanskaya Guba (western coast of northern Baikal). 



Megalovalvata piligera nudicarinata (Lindholm, 1924) F 

Valvata (Liratina) piligera nudicarinata Lindholm, 1924b: 217. 

● Type locality: southern Baikal, ? Murinskaya bank. 



Valvata Müller, 1774 

Type species: Valvata cristata Müller, 1774 (monotypy)

Valvata andreaei Menzel, 1904 F 

Valvata (Cincinna) andreana (lapsus calami) Menzel, 1904: 77, textfig. 

● Type locality: "in der diluvialen (interglacialen) pflanzenreichen Süsswasserablagerung von Wallensen im 

Südlichen Hannover und in dem altalluvialen Kalktufflager von Alfeld a.d. Leine". 


● Distribution: lakes of the basin of the Baltic Sea, Western Bug basin, south of Western Siberia (Anistratenko, 


● Remark: the species was named after German naturalist A. Andreae and therefore the spelling of species epithet 

andreana is obviously lapsus calami. Menzel corrected himself the spelling in the same issue of Nachrichtsblatt 

der Deutschen Malakozoologischen Gesellschaft on page 96. Nevertheless, in Russian literature 

the species epithet is traditionally spelled incorrectly (including Kantor, Sysoev, 2005). 

Valvata cristata Müller, 1774 F 

[= Nerita valvata Gmelin, 1778] 

Müller, 1774: 198-199. 


● Types: possibly, in ZMUC. 

● Distribution: Europe, western Siberia, central Dnieper basin (Anistratenko, 1998b; Starobogatov et al., 2004). 

Valvata elatior Menzel, 1904 F 

Valvata (Cincinna) andreana var. latior Menzel, 1904: 77-78, textfig. 



● Distribution: lakes of the basin of the Baltic Sea (Starobogatov et al., 2004). 

Valvata geyeri Menzel, 1904 F 

Valvata (Cincinna) geyeri Menzel, 1904: 78-79, textfig. 

● Type locality: "in Weissen See bei Fuessen in Bayern". 


● Distribution: lakes of the basin of the Baltic Sea, central Dnieper, south of Western Siberia (Anistratenko, 


● Remark: Glöer (2002) regards V. geyeri as a subspecies of Valvata piscinalis (Müll.). 

Valvata lietuvensis Chernogorenko et Starobogatov, 1987 F 

Chernogorenko, Starobogatov, 1987b: 149. 

● Type locality: Vyshtitis Lake, at the border of Kaliningrad district and Lithuania. 



Valvata nana Westerlund, 1886 F 

[= Valvata (?) minuta “Draparnaud” Westerlund, 1873: 437). 

Valvata nana Westerlund, 1886: 141 

● Type locality: Denmark. 

● Types: Possibly, in NMG. 

● Distribution: lakes of the basin of the Baltic Sea (Starobogatov et al., 2004). 

Valvata planorbulina Paladilhe, 1867 F 

Paladilhe, 1867: 50, pl. 21, figs. 23-26. 

● Type locality: "dans des alluvions du Lez". 


● Distribution: basin of the Baltic Sea, central Dnieper basin (Anistratenko, 1998b; Starobogatov et al., 2004).

Valvata pusilla (Müller, 1774) F 

Nerita pusilla Müller, 1774: 171. 

● Type locality: "In lacu Ruppinensi". 

● Types: unknown. Possibly, in ZMUC (fide Dance, 1986). 

● Distribution: lakes of the basin of the Baltic Sea, Western Bug river basin (Anistratenko, 1998b; Starobogatov 

et al., 2004). 

Valvata spirorbis Draparnaud, 1805 F 

Draparnaud, 1805: 41. 


● Types: not traced. Possibly, in NHMV (fide Dance, 1986). 

● Distribution: Europe, central Dnieper basin (Anistratenko, 1998b; Starobogatov et al., 2004). 

Valvata trochoidea Menke, 1846 F 

Valvata contorta var. trochoidea Menke, 1846: 116. 


● Types: not traced. Menke’s collection was dispersed after his death (Dance, 1986). 

● Distribution: lakes of Europe, except extreme north and south, south of Western Siberia (Anistratenko, 1998b; 

Starobogatov et al., 2004; Lazutkina, 2004). 

Valvata vystitiensis Chernogorenko et Starobogatov, 1987 F 

Chernogorenko, Starobogatov, 1987b: 148. 

● Type locality: Vyshtitis lake, at the border of Kaliningrad district and Lithuania. 



BORYSTHENIINAE Starobogatov, 1983 

Borysthenia Lindholm, 1913 

Ann. Mus. zool. Ac. Sci, 18: 167, nom. nov. pro Jelskia 

Type species: Valvata naticina Menke, 1845 (так в 1927) 

Borysthenia alligans (Lindholm, 1927) F 

Valvata naticina f. alligans Lindholm, 1927a: 22. 

● Type locality: "Weichsel bei Plock" (Poland). 

● Types: ZIN, № 1 in systematic catalogue. 

● Distribution: basins of Dnieper, Dniester, Neman, and Visla rivers; Kaliningrad Region (Anistratenko, 1998b; 


Borysthenia jelskii (Crosse, 1863) F 

Valvata jelskii Crosse, 1863: 382-384, pl. 13, fig. 3. 

● Type locality: Dnieper, near Kiev. 

● Holotype: MNHN. 

● Distribution: known on the single specimen from Dnieper near Kiev (Starobogatov et al., 2004). 

Borysthenia menkeana (Jelski, 1863) F 

Valvata menkeana Jelski, 1863: 136-137, pl. 6, fig. 4. 

● Type locality: Dnieper, off Romanovka, near Kiev. 

● Syntype: MNHN. 

● Lectotype: ZIN, № 2 in systematic catalogue; paralectotype - MNHN. 

● Distribution: basins of Dnieper, South Bug, Dniester, Danube, Neman, Visla rivers; Kaliningrad Region (Anistratenko, 

1998b; Starobogatov et al., 2004).

Borysthenia naticina (Menke, 1846) F 

Valvata naticina Menke, 1846: 129. 

● Type locality: “Hungaria, ad Pestinum”. 

● Types: not traced. Menke’s collection was dispersed after his death (Dance, 1986). 

● Distribution: basins of Dnieper, South Bug, Dniester, Danube, Neman, Visla rivers; Kaliningrad Region (Anistratenko, 


PULMONATA 

Informal Group BASOMMATOPHORA Keferstein, 1865 

ACROLOXOIDEA Thiele, 1931 

ACROLOXIDAE Thiele, 1931 


Acroloxus Beck, 1837 

Type species: Patella lacustris Linnaeus, 1758 (SD Herrmannsen, 1846) 

Acroloxus arachleicus Kruglov et Starobogatov, 1991 F 

Kruglov, Starobogatov, 1991: 74, fig. 2, 9. 

● Type locality: Arakhlej Lake, Zabaikalje, [SE Siberia]. 

● Holotype: ZIN No 1 in systematic catalogue. 


Acroloxus baicalensis Kozhov, 1936 F 

Acroloxus lacustris var. baicalensis Kozhov, 1936: 184, 314, pl. 7, fig. 39. 

● Type locality: Krutaya Inlet (Chivyrkuiski Bay, Baikal), 3 m. 


● Distribution: Angara, Yenissei, Baikal (Starobogatov et al., 2004; Sitnikova et al., 2004). 

Acroloxus caucasicus Kruglov et Starobogatov, 1991 F 

Kruglov, Starobogatov, 1991: 73-74. fig. 2, 7. 

● Type locality: Shaitankazak Lake, Chontaul’sk reservation, Dagestan [Caucasus]. 

● Holotype: ZIN. No 1 in systematic catalogue. 

● Distribution: Dagestan, Stavropol Region, in lakes (Starobogatov et al., 2004). 

Acroloxus hassanicus Kruglov et Starobogatov, 1991 F 

[= Acroloxus hassanicus Kruglov et Starobogatov in Bogatov et Zatrawkin, 1990 [1992]] 

Kruglov, Starobogatov, 1991: 77, fig. 1 4, 3 5. 

● Type locality: Pojma River (former Adimi), near Bamburovo village, Khasanskij district, Primorje Territory 

[Far East]. [The holotype label contains infromation that it was collected in an unnamed ox-bow in the 

floodplain of the Adimi River]. 


● Distribution: Primorje: basins of Ussuri River, Khanka Lake and rivers of southern Primorje (Starobogatov 

et al., 2004). 

Acroloxus klucharevae Kruglov et Starobogatov, 1991 F 

[= Acroloxus klucharevae Kruglov et Starobogatov in Bogatov et Zatrawkin, 1990 (1992)] 

Kruglov, Starobogatov, 1991: 78, fig. 1 (6). 

● Type locality: Lake Bolshoi Vavai, near Aniva Bay, south Sakhalin Island.



Acroloxus kolhymensis Prozorova et Starobogatov, 1998 F 

Prozorova, Starobogatov, 1998a: 39-41, figs. 1A, 2A-B. 

● Type locality: Shuchie Lake, lower Kolyma basin in vicinities of Cherskyi settlement. 


● Distribution: lower Kolyma basin (Starobogatov et al., 2004). 

Acroloxus lacustris (Linnaeus, 1758) F 

Patella lacustris Linnaeus, 1758: 783. 


● Probable syntypes: UUZM Nos. 1269 a-b. 

● Distribution: Europe and western Siberia, in rivers and permanent waterbodies (Starobogatov et al., 2004). 

Acroloxus likharevi Moskvicheva, Kruglov et Starobogatov in Kruglov et Starobogatov, 1991 F 

[= Acroloxus likharevi Moskvicheva, Kruglov et Starobogatov in Bogatov et Zatrawkin, 1990 [1992]] 

Kruglov, Starobogatov, 1991: 75-76, fig. 1 2, 3 5-6. 

● Type locality: Olenevka River, basin of the River RazdoFnaya near Ussurijsk [Far East]. 


● Distribution: basins of Amur and Tugur (southern Okhotsk Sea maritime area) rivers and rivers entering 

Japan Sea and Tatar Strait (Starobogatov et al., 2004). 

Acroloxus lindholmi Kruglov et Starobogatov, 1991 F 

Kruglov, Starobogatov, 1991: 72, fig. 2 (4). 

● Type locality: Luga River near Preobrazhenskoe, Leningrad Region. 



Acroloxus oblongus (Lightfoot, 1786) F 

Patella oblonga Lightfoot, 1786: 168, pl. 3, figs. 1-5. 

● Type locality: near Beaconsfield, Buckinghamshire. 


● Distribution: Europe (probably except north-eastern part), western Siberia, in still waterbodies (Starobogatov 

et al., 2004). 

Acroloxus okaensis Kruglov et Starobogatov, 1991 F 

Kruglov, Starobogatov, 1991: 70-72, fig. 1 3, 3 3. 

● Type locality: Oka River [central Russia], zaton Studenetz. 


● Distribution: Europe (except, probably, north-east), in large and shallow still waterbodies (Starobogatov et 

al., 2004). 

Acroloxus orientalis Kruglov et Starobogatov, 1991 F 

[= Acroloxus orientalis Kruglov et Starobogatov in Bogatov et Zatrawkin, 1990 [1992]] 

Kruglov, Starobogatov, 1991: 75, fig. 2 10. 

● Type locality: spring, connecting lake Kotika with Tym' River, Sakhalin Island. 


● Distribution: northern Sakhalin, upper part of Zeya River basin (Amur Region), in rivers and lakes (Starobogatov 

et al., 2004). 

Acroloxus regelae Prozorova et Starobogatov, 1998 F 

Prozorova, Starobogatov, 1998a: 41-42, figs. 1B, 2C.

● Type locality: Shuchie Lake, lower Kolyma basin in vicinities of Cherskyi settlement. 


● Distribution: lower Kolyma basin (Starobogatov et al., 2004). 

Acroloxus rossicus Kruglov et Starobogatov, 1991 F 


● Type locality: Myshega River, surroundings of Aleksin, Kaluga Region [central Russia]. 


● Distribution: Europe (probably except north-east), in lakes (Starobogatov et al., 2004). 

Acroloxus shadini Kruglov et Starobogatov, 1991 F 

Kruglov, Starobogatov, 1991: 73, fig. 2 6. 

● Type locality: Shaitankazak Lake, Chontaul'sk reservation, Dagestan [Caucasus]. 


● Distribution: entire Europe, south of western Siberia, north of Kazakhstan, Altai, in rivers and lakes (Starobogatov 

et al., 2004). 

Acroloxus ussuriensis Moskvicheva, Kruglov et Starobogatov in Kruglov et Starobogatov, 1991 F 

[= Acroloxus ussuriensis Moskvicheva, Kruglov et Starobogatov in Bogatov et Zatrawkin, 1990 [1992]] 


● Type locality: river Malyi Sungachik, 1 km from the mouth, [Far East]. 


● Distribution: Amur and southern Primorje rivers basins, in rivers and lakes (Starobogatov et al., 2004). 

Acroloxus victori Prozorova, 1996 F 

Prozorova, 1996a: 495-497, figs. 1 (1), 2. 

● Type locality: Chistovodnaya River in vicinities of Vanino City, near railroad bridge. 



Acroloxus zarjaensis Kruglov et Starobogatov, 1991 F 

[= Acroloxus zarjaensis Kruglov et Starobogatov in Bogatov et Zatrawkin, 1990 (1992)] 

Kruglov, Starobogatov, 1991: 77-78, fig. 1 5. 

● Type locality: Lake Zarja, Lazovsk reservation, Primorje [coast of the Japan Sea]. 



Baicalancylus Starobogatov 1967 

Type species: Ancylus dybowskii var. laricensis W. Dybowski, 1913 (OD) 

Baicalancylus boettgerianus (Lindholm, 1909) F 

Ancylus (Pseudancylastrum) boettgerianus Lindholm, 1909: 28, pl. 2, figs. 37-38. 

● Type locality: near cape Kurma in Maloe More (border between central and northern Baikal). 


● Distribution: Maloe More, northern Baikal, 2-10 m (Starobogatov, 1989; Sitnikova et al., 2004). 

Baicalancylus kobelti (W. Dybowski, 1885) F 

Ancylus kobelti W. Dybowski, 1885: 313, fig.1. 

● Type locality: near Listvyanka (western coast of southern Baikal). 


● Distribution: southern Baikal, Angara, 5-10 m (Starobogatov, 1989; Sitnikova et al., 2004).

Baicalancylus laricensis (W. Dybowski, 1913) F 

Ancylus dybowskii var. laricensis W. Dybowski, 1913: 140 (nom. nov. pro Ancylus dybowskii sensu Lindholm, 

1909). 

● Type locality: near Listvyanka (western coast of southern Baikal). 



Baicalancylus njurgonicus Starobogatov, 1989 F 

Starobogatov, 1989: 72, fig. 2, 13 5, z, l. 

● Type locality: Maloe More, Cape Njurgon [Baikal Lake], 3 m. 



Gerstfeldtiancylus Starobogatov, 1989 

Type species: Gerstfeldtiancylus gerstfeldti Starobogatov, 1989 (OD) 

Gerstfeldtiancylus benedictiae Starobogatov, 1989 F 

[= Ancylus (Pseudancylastrum) troscheli sensu Lindholm, 1909, non W. Dybowski, 1875, part.] 


● Type locality: vicinities of Listvennichnyj [SW coast of Baikal Lake], 3-14 m. 


● Distribution: Baikal, 3-20 m, mostly on shells of Benedictia (Sitnikova et al., 2004). 

● Remark: Among paratypes there are 6 paralectotypes of Ancylus sibiricum Gerstfeldt, 1859, coll. 

A.A.Maak, 1854 from "Angara and Baikal Lake". 

Gerstfeldtiancylus caputiformis Starobogatov, 1989 F 


● Type locality: Zavorotnaya Inlet, western coast of northern Baikal Lake, 4 m. 



Gerstfeldtiancylus kotyensis Starobogatov, 1989 F 

[Gerstfeldtiancylus kozhovi Starobogatov, 1989] 


● Type locality: near Bolshiye Koty, NW part of southern Baikal Lake, 25-40 m. 



● Remark: synonymy of G. kotyensis and G. kozhovi was proved by Shirokaya (2005). 

Gerstfeldtiancylus pileolus Starobogatov, 1989 F 

Starobogatov, 1989: 67, fig. 2 8 5, z, l. 

● Type locality: Zavorotnaya Inlet, western coast of northern Baikal Lake, 4 m. 


● Distribution: type locality (Starobogatov, 1989; Sitnikova et al., 2004). 

Gerstfeldtiancylus porfirievae Starobogatov, 1989 F 

[= Ancylus (Pseudancylastrum) troscheli sensu Lindholm, 1909, non W. Dybowski, 1875, part.] 


● Type locality: Cape Kocherikovskij, W coast of northern Baikal Lake, 6-8.5 m. 


● Distribution: northern Baikal, 4-12 m (Sitnikova et al., 2004).

Gerstfeldtiancylus renardii (W. Dybowski, 1884) F 

[Gerstfeldtiancylus gerstfeldti Starobogatov, 1989] 

“Ancylus” renardii W. Dybowski, 1884: 157, pl. 4, figs. 2, 4. 

● Type locality: southern Baikal. 

● Syntypes: Zoological Museum of Lviv University (Ukraine). 

● Distribution: western coast of Baikal, 4-20 m (Sitnikova et al., 2004). 

● Remark: synonymy of G. renardii and G. gerstfeldti was proved by Shirokaya (2005). 

Gerstfeldtiancylus roepstorfi Shirokaya, Röpstorf et Sitnikova, 2003 F 

Shirokaya, Röpstorf, Sitnikova, 2003: 117, 119, fig. 3C-D, 4 A-I. 

● Type locality: littoral of Maly Uskany Island (Baikal), depth 13 m. 

● Holotype: collections of Limnological Institute, No. 898. 

● Distribution: southern and northern Baikal, 13-20 m (Shirokaya, Röpstorf, Sitnikova, 2003). 

Gerstfeldtiancylus ushunensis Shirokaya, 2007. F 

Shirokaya, 2007: 56, fig. 3-7. 

● Type locality: Lake Baikal, Malye Olkhonskie Vorota, Ushun Bay, 53°00’54’’ N 106°56’96’’ E, 20-22 m 

depth, on stones. 

● Holotype: ZIN, No 1-521-2006. 

● Distribution: type locality (Shirokaya, 2007). 

Pseudancylastrum Lindholm, 1909 

Type species: Ancylus sibiricus Gerstfeldt, 1859 (OD) 

Pseudancylastrum aculiferum Starobogatov, 1989 F 

[= Ancylus sibiricus Gerstfeldt, 1859, part.] 

Pseudancylastrum aculiferum Starobogatov, 1989: 55, fig. 1, 10, 5, z, l. 

● Type locality: Kultuk, SW corner of Baikal Lake. 


● Distribution: southern Baikal, 4-36 m (Starobogatov, 1989; Sitnikova et al., 2004). 

● Remark: Holotype is at the same time a paralectotype of Ancylus sibiricum Gerstfeldt, 1859. 

Pseudancylastrum beckmanae Starobogatov, 1989 F 

Pseudancylastrum beckmanae Cтаробогатов, 1989: 49, fig. 1, 4 5, z, l. 

● Type locality: near Cape Solontzovyj, western coast of the northern Baikal Lake, 40-18 m. 


● Distribution: western coast of northern Baikal, 4-20 m (Sitnikova et al., 2004). 

Pseudancylastrum cornu Starobogatov, 1989 F 


● Type locality: southern Baikal Lake. 


● Distribution: southern Baikal (Sitnikova et al., 2004). 

Pseudancylastrum dorogostajskii Starobogatov, 1989 F 

Pseudancylastrum dorogostajskii Strobogatov, 1989: 57, fig. 1, 12 5, z, l. 

● Type locality: vicinities of Listvennichnyj (Baranchik) [SW coast of the Baikal Lake], 6-15 m. 


● Distribution: western coast of southern Baikal and eastern coast of northern Baikal, 3-35 m (Starobogatov, 

1989; Sitnikova et al., 2004). 

Pseudancylastrum dybowskii (Clessin, 1882) F 

Ancylus dybowskii Clessin, 1882b: 38, pl. 7, fig. 1.



● Distribution: Baikal, Angara River downstream to Irkutsk (Starobogatov et al., 2004; Sitnikova et al., 

2004). 

Pseudancylastrum frolikhae Sitnikova et Starobogatov, in Sitnikova, Fialkov, Starobogatov, 1993 F 

Sitnikova et al., 1993: 134, fig. 1. 

● Type locality: Frolikha Bay (south-western coast of northern Baikal), 340-360 m. 

● Holotype: Limnological Institute, No. 263-264. 


Pseudancylastrum irindaense Starobogatov, 1989 F 

Pseudancylastrum irindaense Starobogatov, 1989: 52, fig. 1, 7 5, z, l. 

● Type locality: Irinda Bay, east coast of northern Baikal Lake, 5 m. 


● Distribution: type locality, Maloe More [Baikal], 2-10 m (Starobogatov, 1989). 

Pseudancylastrum korotnevi Starobogatov, 1989 F 

Pseudancylastrum korotnevi Cтаробогатов, 1989: 51, fig. 1, 6 5, z, l. 

● Type locality: near Cape Kocherikovskij, western coast of northern Baikal Lake, 6-8.5 m. 


● Distribution: western coast of Baikal, 4-10 m (Sitnikova et al., 2004). 

Pseudancylastrum olgae Starobogatov, 1989 F 

Pseudancylastrum olgae Starobogatov, 1989: 48s, fig. 1 (3), 5 z, l. 

● Type locality: near Kluevka, SE coast of southern Baikal Lake, 8 m. 


● Distribution: south-eastern coast of Baikal, 2.5-10 m (Sitnikova et al., 2004). 

Pseudancylastrum poberezhnyi Starobogatov, 1989 F 

Pseudancylastrum poberezhnyi Starobogatov, 1989: 56, fig. 1, 11 5, z, l. 

● Type locality: near Bolshie Koty [SW Baikal Lake]. 


● Distribution: Baikal, 4-15 m (Starobogatov, 1989; Sitnikova et al., 2004). 

Pseudancylastrum sibiricum (Gerstfeldt, 1859) F 

Ancylus sibiricum Gerstfeldt, 1859: 23, fig. 30. 

● Type locality: Angara River within borders of Irkutsk. 


● Distribution: southern Baikal, Angara River downstream to Irkutsk (Starobogatov et al., 2004; Sitnikova et 

al., 2004). 

Pseudancylastrum troschelii (W. Dybowski, 1875) F 

Ancylus troschelii W. Dybowski, 1875: 64, pl. 4, figs. 35-37. 

● Type locality: Kultuk (southern Baikal) [restricted Sitnikova et al., 2004]. 


● Distribution: southern Baikal, 5-25 m (Starobogatov, 1989; Sitnikova et al., 2004). 

Pseudancylastrum werestschagini Starobogatov, 1989 F 

Pseudancylastrum werestschagini Starobogatov, 1989: 52, fig. 1, 8 5. z. l. 

● Type locality: near Bolshie Koty [southern Baikal Lake], 3-5 m. 


● Distribution: southern Baikal, 3-20 m; Angara River downstream to Irkutsk (Starobogatov, 1989; Star-

obogatov et al., 2004; Sitnikova et al., 2004). 

LYMNAEOIDEA Rafinesque, 1815 

LYMNAEIDAE Rafinesque, 1815 


Aenigmomphiscola Kruglov et Starobogatov, 1981 

Type species: Aenigmomphiscola europaea Kruglov et Starobogatov, 1981 (OD) 

Aenigmomphiscola europaea Kruglov et Starobogatov, 1981 F 

Kruglov, Starobogatov, 1981: 969, fig. 1, 1, 7, 12;3A;4A. 

● Type locality: vicinities of Ufa city, bog near Urgun village (in reality, Urgun village is situated 230 km 

eastern from Ufa city; M.E. Grebennikov, personal communication). 


● Distribution: eastern part of European Russia (Kruglov, Starobogatov, 1993b; Starobogatov et al., 2004). 

Aenigmomphiscola kazakhstanica Kruglov et Starobogatov, 1981 F 

Kruglov, Starobogatov, 1981: 973, fig. 1, 3, 9, 14; 3 B; 4B. 

● Type locality: Kokchetav Region, Shchucghinskij district [northern Kazakhstan], pasture Kyzyl-Agach. 


● Distribution: south of western Siberia and northern Kazakhstan, eastward to Altai (Starobogatov et al., 

2004). 

Aenigmomphiscolauvalievae Kruglov et Starobogatov, 1981 F 

Kruglov, Starobogatov, 1981: 971-973, fig. 1, 2, 8, 13; 2; 3B; 4B; 5. 

● Type locality: Kokchetav Region, Shchucghinskij district [northern Kazakhstan], sovkhoz imeni Frunze, 

pasture. 


● Distribution: eastern part of European Russia, south of western Siberia and northern Kazakhstan, eastward 

to Altai (Starobogatov et al., 2004). 

Lymnaea Lamarck, 1799 

Type species: Helix stagnalis Linnaeus, 1758 (monotypy). 

Lymnaea aberrans (Westerlund, 1897) F 

Limnaea ovata var. aberrans Westerlund, 1897b: 125. 

● Type locality: Kamchatka River basin. 

● Lectotype (Kruglov, Starobogatov, 1984a): ZIN, No. 1 in systematic catalogue. 

● Distribution: eastern Kamchatka, northern and western shores of the Okhotsk Sea, Kolyma River, North 

Kurile Islands (Kruglov, Starobogatov, 1993b; Starobogatov et al., 2004). 

Lymnaea almaatina Izzatullaev, Kruglov et Starobogatov, 1983 

Izzatullaev et al., 1983a: 323, fig. 1 (3-4). 

● Type locality: spring in the valley of Bolshaja Almaatinka River [southern Kazakh SSR] 


● Distribution: mountain part of Central Asia, probably in inner Altai (Kruglov, Starobogatov, 1993a; Starobogatov 

et al. 2004) 

Lymnaea alticola Izzatullaev, Kruglov et Starobogatov, 1983 F 

Izzatullaev et al., 1983b: 53, fig. 1-2.

● Type locality: Pamir Mountains, hot spring near Jashikul Lake [Tajik SSR]. 


● Distribution: mountains of Central Asia (thermal springs) (Kruglov, Starobogatov, 1993a). 

Lymnaea ampla (Hartmann, 1821) F 

[= Lymnaea patula sensu Starobogatov, 1977 non Da Costa, 1778] 

● Type locality”Rhine near Reineck” (locality of the lectotype collection) 

● Lectotype designated by Vinarski and Glöer (2007) is housed in Naturmuseum Sain-Gallne, Switzerland. 

● Distribution: Europe, southern Siberia eastward to Baikal Lake (Vinarski, in press). 

Lymnaea ampullacea (Rossmässler, 1835) F 

Limnaeus ampullaceus Rossmässler, 1835b: 19, pl. 7, fig. 124. 

● Type locality: “im Lac de Joux im Jura (5. Charp.)”. 

● Types: lost (R. Janssen, pers. comm.). 

● Distribution: Europe, southern Siberia eastward to Baikal Lake, Baikal Lake (Kruglov, Starobogatov, 

1993b; Sitnikova et al., 2004). 

Lymnaea amurensis Kruglov, Moskvicheva et Starobogatov in Kruglov et Starobogatov, 1984 F 

Kruglov, Starobogatov, 1984a: 28, fig. 1, 12; 2, 14. 

● Type locality: Khabarovsk Territory [Far East], valley of the Amur River, shallow water body near the 

Petropavlovskoe Lake, near Kukhari village. 


● Distribution: Amur basin, Primorje (Kruglov, Starobogatov, 1993b). 

Lymnaea arachleica Kruglov et Starobogatov, 1989 F 

Kruglov, Starobogatov, 1989b: 23, fig. 1, 6, 2, 2. 

● Type locality: Lake Arakhlei (Transbaikalia). 


● Distribution: Lake Arakhlei (Chita Region) (Kruglov, Starobogatov, 1993a). 

Lymnaea araratensis Kruglov et Starobogatov, 1985 F 

[= Limnaea elophila sensu Kobelt, 1877, non Bourguignat, 1862; Lymnaea omsiana sensu Hubendick, 1951, 

non Locard, 1883]. 

Kruglov, Starobogatov, 1985c: 26. fig. 1 d, 3d. 

● Type locality: pond near the railway station Masis, Armenian SSR. 


● Distribution: Armenia, western Middle East, Kirghizstan (lake Son-Kul), Pamir (Kruglov, Starobogatov, 

1993a). 

Lymnaea archangelica Kruglov et Starobogatov, 1986 F 

Kruglov, Starobogatov, 1986: 60, fig. 1; 2, 2. 

● Type locality: pool in the Hood lands of the Uemljanka River, Arkhangelsk Region, 10-12 km from entering 

the Severnaja Dvina River. 


● Distribution: northern part of East Europe, West Siberia (Kruglov, Starobogatov, 1993a). 

Lymnaea atkaensis Dall, 1884 F 

[= Lymnaea randolphi Baker, 1904] 

Lymnaea ovata var. atkaensis Dall, 1884: 343. 

● Type locality: Aleutians. 

● Syntypes: USNM 29031. 

● Distribution: Chukchi Peninsula (Amguema river drainage and eastwards), Koryak upland (Kruglov, Starobogatov, 

1993a; Starobogatov et al., 2004).

Lymnaea atra atra (Schrank, 1803) F 

Buccinum atrum Schrank, 1803: 288. 

● Type locality: “in der Donau”. 

● Types: lost (G. Falkner, personal communication). 

● Distribution: Europe (excluding extreme northeastern part) (Kruglov, Starobogatov, 1993a). 

Lymnaea atra starobogatovi Lazareva, 1967 F 

Lymnaea starobogatovi Lazareva, 1967a: 1343-1344, figs. 1 (2-2a), 2 (2-2a). 

● Type locality: former river-bed of Tobol River, Kustanai Region, Kazakhstan. 


● Distribution: steppes of Kazakhstan and the Irtysh basin (Kruglov, Starobogatov, 1993a). 

Lymnaea atra zebrella (B. Dybowski, 1913) F 

Costolimnaea zebrella B. Dybowski, 1913: 186, pl.4, fig. 10 a, b. 

● Type locality: vicinities of Kultuk (southern Baikal). 

● Lectotype (Starobogatov, Streletzkaya, 1967): ZIN, No. 1 in systematic catalogue. 

● Distribution: northeastern Europe, West Siberia, Yenissei basin, Baikal (Kruglov, Starobogatov, 1993a; 

Sitnikova et al., 2004). 

Lymnaea auricularia (Linnaeus, 1758) F 

[= Radix auriculatus Montfort, 1810] 

Helix auricularia Linnaeus, 1758: 774-775. 


● Types: possibly in UUZM. 

● Distribution: Europe, Siberia, Uzbekistan, Baikal, rivers of Pacific coast of Far East from Kamchatka to 

Primorje (Kruglov, Starobogatov, 1993a; Starobogatov et al., 2004; Sitnikova et al., 2004). 

Lymnaea azabatschensis Kruglov et Starobogatov, 1989 F 

Kruglov, Starobogatov, 1989a: 18, fig. 1, 5, 2, 4. 

● Type locality: Kamchatka, channel from Azabachje Lake to the Kamchatka River. 


● Distribution: southeastern Kamchatka (Kruglov, Starobogatov, 1993a). 

Lymnaea bactriana (Hutton, 1849) F 

Limnaea bactriana Hutton, 1849: 656. 

● Type locality: Quetta (fide Annandale, Prashad, 1919) [Afghanistan]. 

● Holotype: Asiatic Society of Bengal (fide Annandale, Prashad, 1919). 

● Distribution: western Middle East, eastern Transcaspia, Central Asia (Kruglov, Starobogatov, 1993a). 

Lymnaea badia (Küster, 1862) F 

Limnaeus badius Küster, 1862: 23, pl. 4, figs. 18-19. 

● Type locality: River Narenta, Dalmatia. 


● Distribution: within Russia – Volga delta (Starobogatov et al., 2004). 

Lymnaea bakowskyana (Clessin, 1879) F 

Lymnaea peregra var. bakowskyana Clessin, 1879: 12, pl. 1, fig. 8. 

● Type locality: “Plickow. Galicia”. 


● Distribution: southern Europe, Caucasus (Kruglov, Starobogatov, 1993b). 

Lymnaea balthica (Linnaeus, 1758) F

[Turbo patulus Da Costa, 1778; Limnaea inflata Kobelt, 1870] 

Helix balthica Linnaeus, 1758: 775. 

● Type locality: “M. Balthici littora”. 

● Neotype (Kruglov, Starobogatov, 1983b): ZIN. 

● Distribution: Europe, southern Siberia, Baikal (Kruglov, Starobogatov, 1993b; Sitnikova et al., 2004). 

● Remark: synonymy of Lymnaea patula and L. balthica was suggested by Vinarski and Glöer (2007), who 

also designated lectotype of the former species. 

Lymnaea berlani (Bourguignat, 1870) F 

Limnaea berlani Bourguignat, 1870: 44-45. 

● Type locality: not stated in the original description (lower Danube – from title). Hubendick (1951: 182) 

states that the type locality is situated somewhere in Bulgaria. 


● Distribution: South Europe, northern Caucasus, Transcaucasia (Kruglov, Starobogatov, 1993a; Starobogatov 

et al., 2004). 

Lymnaea blauneri (Shuttleworth in Küster, 1862) F 

Limnaeus blauneri Shuttleworth – Küster, 1862: 56, pl. 12, fig. 7-8. 

● Type locality: “In der Scweitz im Canton Wallis, auf dem Matterhorn”. 


● Distribution: Ciscaucasia (Kruglov, Starobogatov, 1993b; Starobogatov et al., 2004). 

Lymnaea bodamica (K. Miller, 1873) F 

Limnaea stagnalis var. bodamica K. Miller, 1873: 4, pl. 1, fig. 2. 

● Type locality: Bodensee. 


● Distribution: basins of Baltic and White seas (Starobogatov et al., 2004). 

Lymnaea bowelli (Preston, 1909) F 

Limnaea bowelli Preston, 1909: 115. 

● Type locality: “Te-ring Gompa, in a small hill stream arising from a spring, 14, 000 feet; also from Mangtsa, 

14, 500 feet; High Hill, Gompa, Gyantse valley in a small hill stream, among moss and stones, 14, 500 

feet; and Gyantse, 13, 120 feet” [Tibet]. 


● Distribution: mountain part of Central Asia, probably in Tuva and Altai (Kruglov, Starobogatov, 1993a; 


Lymnaea callomphala (Servain, 1881) 

[= Limnaea palustris var. transsylvanica Kimakowicz, 1884] 

Limnaea callomphala Servain, 1881: 78. 

● Type locality: "lac [Balaton], entre Füredet et Tihany". 


● Distribution: western and southern parts of European Russia, Ukraine (Kruglov, Starobogatov, 1993a; Starobogatov 

et al., 2004). 

● Remark: empty shells that seems conspecific with L. callomphala were found in Western Siberia, although 

the taxonomic position should be confirmed by anatomical data (M.V. Vinarski, unpublished). 

Lymnaea carelica Kruglov et Starobogatov, 1983 F 

Kruglov, Starobogatov, 1983b: 1467, fig. 2, 6; 3J. 

● Type locality: shallow water body near Svjatozero (Karelian ASSR) 


● Distribution: Karelia (Baltic drainage area) (Kruglov, Starobogatov, 1993b).

Lymnaea chereshnevi Kruglov et Starobogatov, 1989 F 

Kruglov, Starobogatov, 1989b: 27, fig. 1, 24; 2, 22. 

● Type locality: Maloe Lake, basin of the river Bolshaya, western Kamchatka. 


● Distribution: western and southern Kamchatka, North Kurile Islands (Kruglov, Starobogatov, 1993a; Starobogatov 

et al., 2004). 

Lymnaea clavata (Westerlund, 1885) F 

[= Galba glabra sensu Jackiewicz, 1959, non Müller, 1774]. 

Limnaea glabra var. clavata Westerlund, 1885a: 49. 

● Type locality: “Norddeutschland, Schweeden”. 

● Syntypes: most probably in NMG. 

● Distribution: Baltic Sea basin (Luga River) (Kruglov, Starobogatov, 1993a; Starobogatov et al., 2004). 

Lymnaea coreana (Martens, 1886) F 

Limnaea auricularia var. coreana Martens, 1886: 80. 

● Type locality: “Changjin, Prov. Hangyöngdo, Korea”. 


● Distribution: Amur basin, southern part of Primorje (Kruglov, Starobogatov, 1993a; Starobogatov et al., 

2004). 

Lymnaea corvus (Gmelin in Linnaeus, 1791) F 

Helix corvus Gmelin, 1791: 3665 

● Type locality: "in Thuringiae". 


● Distribution: Baltic Sea basin, north-western part of the Dnieper drainage area (right tributaries of the Pripyat 

river) (Kruglov, Starobogatov, 1993a; Starobogatov et al., 2004). 

Lymnaea curtacorvus Kruglov et Starobogatov, 1984 F 

[=Limnaea palustris var. curta Clessin, 1873] 

Kruglov, Starobogatov, 1984b: 65, fig. 1, J; 2 5, 6. 

● Type locality: Karelia, Pildhzskoe Lake. 


● Distribution: Baltic Sea basin (Kruglov, Starobogatov, 1993a; Starobogatov et al., 2004). 

Lymnaea danubialis (Schrank, 1803) F 

Buccinum danubialis Schrank, 1803: 286. 



● Distribution: northern Caucasus and Rostov Region (Starobogatov et al., 2004), Southern Urals (Vinarski, 

unpublished). 

Lymnaea dipkunensis Gundrizer et Starobogatov, 1979 F 

Gundrizer, Starobogatov, 1979: 1134, fig. 1, 4. 

● Type locality: Gornoe Lake, flood land of the Kurejka River upper mouth of river Dipkun (NW Siberia]. 


● Distribution: northern part of Central Siberia (Kruglov, Starobogatov, 1993b). 

Lymnaea dolgini Gundrizer et Starobogatov, 1979 F 

Gundrizer, Starobogatov, 1979:1132, fig. 1, 2; 2, 2. 

● Type locality: lake in flood land of the Kurejka River 20 km upper its mouth [NW Siberia]. 

● Holotype: ZIN No 1 in systematic catalogue.

● Distribution: lower part of the Ob basin, Irtysh and Yenissei basins (Kruglov, Starobogatov, 1993b; Dolgin, 

2001; Vinarski, 2008). 

Lymnaea doriana (Bourguignat, 1862) F 

Limnaea doriana Bourguignat, 1862a: 60. 

● Type locality: "Sicile". 

● Types: MHNG. 

● Distribution: south of East Europe, Caucasus, northern part of Kazakhstan, Tuva (Kruglov, Starobogatov, 

1993a; Prozorova, Sharyi-Ool, 1999; Starobogatov et al., 2004). 

Lymnaea dupuyi (Locard, 1893) F 

Amphipeplea dupuyi Locard, 1893: 47. 

● Type locality: "marais de Peychaud (Gironde)" (France). 


● Distribution: Baltic Sea basin (Kruglov, Starobogatov, 1993b; Starobogatov et al., 2004). 

Lymnaea dvoriadkini Kruglov et Starobogatov, 1984 F 


● Type locality: Primorje Territory [Far East], northern coast of the Vostok Bay, stream near Avangard settlement. 


● Distribution: Amur basin, Primorje (Kruglov, Starobogatov, 1993b). 

Lymnaea euphratica (Mousson, 1874) F 

Limnaea euphratica Mousson, 1874: 40-41. 

● Type locality: “Samava” [lower Mesopothamia]. 


● Distribution: southern Daghestan, Azerbaijan, Tajikistan, Uzbekistan, Turkmenistan (Kruglov, Starobogatov, 

1993a). 

Lymnaea falsipalustris Kruglov et Starobogatov, 1989 F 


● Type locality: Kamchatka, channel from Azabachje Lake to the Kamchatka River. 


● Distribution: southeastern Kamchatka (Kruglov, Starobogatov, 1993a). 

Lymnaea fontinalis (Studer, 1820) F 

[= Lymnaea acutalis Morelet, 1845] 

Limneus fontinalis Studer, 1820: 93. 

● Type locality: “Umgebund von Bern, in Bächen”. 


● Distribution: Europe, Siberia, Baikal (Kruglov, Starobogatov, 1993b, Sitnikova et al., 2004). 

Lymnaea fragilis (Linnaeus, 1758) F 

[= Limnaea stagnalis var. producta Colbeau, 1859; Limnaea raphidia Bourguignat, 1860; Limnaea stagnalis 

var. subulata Westerlund, 1873] 

Helix fragilis Linnaeus, 1758: 774. 


● Types: most probably in the UUZM collection. 

● Distribution: North Europe, entire Siberia, eastward to Kolyma basin (Kruglov, Starobogatov, 1993a; Prozorova, 

1998b). 

Lymnaea fulva (Zigler in Küster, 1862) F

Limnaeus fulva Zigler – Küster, 1862: 16, pl. 4, fig. 19-21. 

● Type locality: “in Krain in Gebirgsquellen”. 


● Distribution: Ciscaucasia (Kruglov, Starobogatov, 1993b; Starobogatov et al., 2004). 

Lymnaea fusca fusca (C. Pfeiffer, 1821) F 

Lymnaeus fuscus C. Pfeiffer, 1821: 92, pl. 4, fig. 25. 

● Type locality: “entre Füred et Tihany” [Balathon Lake]. 


● Distribution: Baltic Sea basin, small waterbodies (Kruglov, Starobogatov, 1993a; Starobogatov et al., 

2004). 

● Remark: in recent western European malacological literature (eg. Falkner et al., 2001; Glöer, 2002) this 

name is used for different species from that of Kruglov and Starobogatov (1993a). 

Lymnaea fusca maritima (Clessin, 1878) B 

Limnaea palustris var. maritima Clessin, 1878: 76, pl. 3, fig. 17. 

● Type locality: “Küste des Bottnischen busens der Lanschaft Medelpad”. 


● Distribution: brackish waters of Baltic Sea (Kruglov, Starobogatov, 1993a; Starobogatov et al., 2004). 

Lymnaea gebleri (Middendorff, 1851) F 

Middendorff, 1851: 292, pl. 30, fig. 1-3. 

● Type locality: “Barnaul” (Middendorff, 1851). Lazareva (1967) believed that this designation is erroneous 

and the true type locality of the species is Zaisan Lake (eastern Kazakhstan). 

● Lectotype (Lazareva, 1967b): ZIN, No. 1 in systematic catalogue. 

● Distribution: eastern Kazakhstan, Tuva, Altai? (Lazareva, 1967b; Prozorova, Sharyi-Ool, 1999; Kruglov, 

Starobogatov, 1993a; Starobogatov et al., 2004;). 

Lymnaea gedrosiana (Annandale et Prashad, 1919) F 

Limnaea gedrosiana Annandale, Prashad, 1919: 48-49, pl.7, fig. 2-4. 

● Type locality: “Pishin district (Chaman), Kandahar, Quetta and in the Hamun-i-Helmand in Seistan”. 

● Syntypes: Zoological Survey of India, M 11533/2. 

● Distribution: southern Tajikistan (Kruglov, Starobogatov, 1993a). 

Lymnaea glabra (Müller, 1774) F 

Buccinum glabrum Müller, 1774: 135-136. 


● Types: ZMC (see Dance, 1986). 

● Distribution: probably in Baltic Sea basin (Starobogatov et al., 2004). 

Lymnaea glutinosa (Müller, 1774) F 

Buccinum glutinosum Müller, 1774: 129. 



● Distribution: Europe, south part of West Siberia, northern part of Central Siberia (eastward to Ob mouth), 

northern Kazakhstan (Kruglov, Starobogatov, 1993b; Starobogatov et al., 2004). 

Lymnaea goupili (Moquin-Tandon, 1856) F 

Limnaea truncatula var. goupili Moquin-Tandon, 1856: 474. 

● Type locality: “dans la Sarthe (Goupil)” [France]. 


● Distribution: Europe (except for extreme north-east) (Kruglov, Starobogatov, 1993a).

Lymnaea gueretiniana (Servain, 1881) F 

Limnaea gueretiniana Servain, 1881: 76. 

● Type locality: Save, near Belgrad. 


● Distribution: Baltic, Black and Azov seas drainages (Kruglov, Starobogatov, 1993a). 

Lymnaea gundrizeri Kruglov et Starobogatov, 1983 F 

Kruglov, Starobogatov, 1983a: 141. 

● Type locality: water body near the shore of Surulu-Kol’ Lake, Ulaganskij district. Altai Territory [SW Siberia]. 


● Distribution: Altai (Kruglov, Starobogatov, 1993b). 

Lymnaea hadutkae Kruglov et Starobogatov, 1989 F 


● Type locality: hot spring Khadutka. Kamchatka. 


● Distribution: eastern Kamchatka, thermal springs (Kruglov, Starobogatov, 1993a). 

Lymnaea hakusyensis Kruglov et Starobogatov, 1989 F 


● Type locality: hot spring entering Baikal Lake in the vicinities of Khakusa Bay. 


● Distribution: eastern shore of Baikal, thermal springs (Kruglov, Starobogatov, 1993a). 

Lymnaea hartmanni (Studer, 1820) F 

Limneus hartmanni Studer, 1820: 93. 

● Type locality: Bodensee. 


● Distribution: Europe (Kruglov, Starobogatov, 1993b). 

Lymnaea hookeri Reeve, 1850 F 

[= Lymnaea heptapotamica Lazareva, 1967] 

Reeve, 1850: 49, textfig. 

● Type locality: "Thibetan or north side of Sikkim Himalaya, at 18,000 feet elevation". 

● Types: BMNH. 

● Distribution: eastern central Asia, eastern Kazakhstan, Altai, Transbaikalia, Amur drainage including 

Khanka Lake and southern Primorje (Lazareva, 1967; Kruglov, Starobogatov, 1993b; Starobogatov et al., 

2004). 

Lymnaea igarkae Gundrizer et Starobogatov, 1979 F 

Gundrizer, Starobogatov, 1979:1133, fig. 1, 3; 2, 3. 

● Type locality: water body in the city Igarka [NW Siberia]. 



Lymnaea iliensis Lazareva, 1967 F 

Lazareva, 1967a: 1347-1348, figs. 1 (6-6a), 2 (6-6a). 

● Type locality: river floodland lake near Kaskelenka River entrance to Ili River (Kazakhstan). 


● Distribution: Balkhash Lake drainage area (Kruglov, Starobogatov, 1993a), southern part of the Western 

Siberian plain (Vinarski, 2008).

Lymnaea impura (Troschel, 1837) F 

Limnaeus impurus Troschel, 1837: 172. 

● Type locality: river Ganges, India (see Hubendick, 1951). 


● Distribution: Tajikistan, southern Uzbekistan (Kruglov, Starobogatov, 1993a), Turkmenia (Krugov, 2005). 

Lymnaea intercisa (Lindholm, 1909) F 

[= Gulnaria intercisa var. sorica B. Dybowski, 1912] 

Limnaea auricularia var. intercisa Lindholm, 1909: 5, pl. 1, fig.72. 

● Type locality: Maloe More [central Baikal]. 

● Lectotype (Sitnikova et al., 2004): ZIN 

● Distribution: southern part of Central Siberia, Baikal, northeastern Kazakhstan (Kruglov, Starobogatov, 

1993a; Sitnikova et al., 2004). 

● Remark. Sitnikova & Takhteev (2006) considered L. intercisa to be a form of the subspecific rank of L. 

auricularia. 

Lymnaea intermedia Lamarck, 1822 F 

Lymnaea intermedia Lamarck, 1822: 162. 

● Type locality: “en France, dans le Quercy”. 

● Syntype: MHNG. 

● Distribution: Europe, Siberia (Kruglov, Starobogatov, 1993b). 

Lymnaea iturupica Kruglov et Starobogatov, 1989 F 


● Type locality: Lebyazhje Lake, Iturup Island [Kurile Islands], vicinities of Kuril'sk city. 


● Distribution: South Kurile Islands (Kruglov, Starobogatov, 1993a; Starobogatov et al., 2004). 

Lymnaea jacutica Starobogatov et Streletzkaja, 1967 fig. 3B F 

Starobogatov, Streletzkaja, 1967: 233, fig. 27. 

● Type locality: mouth of the Kolyma river, near small settlement Chajchaja. 


● Distribution: northern Siberia from Ural mountains to Kolyma drainage, Far East: northern Okhotsk Sea 

maritime area and northern Sakhalin (Kruglov, Starobogatov, 1993b; Starobogatov et al., 2004). 

Lymnaea japonica (Jay, 1856) F 

Lymnea japonica Jay, 1856: 294-295, pl. 5, figs. 10-12. 

● Type locality: Simoda [Japan]. 


● Distribution: South Kurile Islands (Kruglov, Starobogatov, 1993a). 

Lymnaea juribeica Kruglov et Starobogatov, 1984 F 

Kruglov, Starobogatov, 1984a: 32, fig. 1, 17. 

● Type locality: Tyumen’ Region [Siberia], Gydan Peninsula, Pisi-To Lake in the Juribej River basin. 


● Distribution: Yamal Peninsula, lower Ob’ basin (Kruglov, Starobogatov, 1993b; Dolgin, 2001). 

Lymnaea kafanovi Kruglov et Starobogatov, 1984 F 


● Type locality: Sakhalin Region [Sakhalin Island], vicinities of Nogliki settlement. 

● Holotype: ZIN No 1 in systematic catalogue 

● Distribution: northern Sakhalin (basins of Tym and Poronai rivers) (Kruglov, Starobogatov, 1993b; Starobogatov 

et al., 2004).

Lymnaea kamtschatica (Middendorff, 1851) F 

Limnaeus kamtschaticus Middendorff, 1851: 295, pl. 30, fig. 11-12. 

● Type locality: Kamchatka River (probably plane). 

● Lectotype (Kruglov, Starobogatov, 1984a): ZIN, No. 1 in systematic catalogue. 

● Distribution: eastern Kamchatka, Chukchi Peninsula, Lena, Yana, Indigirka, and Kolyma rivers basins 

(Kruglov, Starobogatov, 1993b; Dolgin, 2001). 

Lymnaea kazakensis Mozley, 1934 F 

Lymnaea palustris kazakensis Mozley, 1934: 3-4, pl. 1, fig.7. 

● Type locality: small dry lake bottom near the village of Novo Troetskaya, northern Kazakstan. 

● Holotype: USNM 470457. 

● Distribution: steppes of West Siberia and Kazakhstan, southern Ural, basin of Ural River and Orenburg 

district (Kruglov, Starobogatov, 1993a; Starobogatov et al., 2004; M.V. Vinarski, unpublished). 

● Remark: although the holotype of Lymnaea palustris kazakensis was designated in the original publication 

(as “type”), in the type collection of USNM under the catalogue number 470457 there is a lot containing 7 

specimens, labeled as syntypes on a later printed label. It was impossible to isolate the “holotype” and non of 

the specimen matches the original illustration. 

Lymnaea kunashirica Kruglov et Starobogatov, 1989 F 


● Type locality: Lagunnoe Lake, Kunashir Island [Kurile Islands]. 


● Distribution: Kunashir Island (Kruglov, Starobogatov, 1993a). 

Lymnaea kurejkae Gundrizer et Starobogatov, 1979 F 

Gundrizer, Starobogatov, 1979: 1131, fig. 1, 1; 2, 1. 

● Type locality: lake in flood land of the Kurejka River 20 km upper its mouth [NW Siberia]. 


● Distribution: northern part of Central Siberia, Irtysh River basin (Kruglov, Starobogatov, 1993b; Vinarski, 

2003). 

Lymnaea kurenkovi Kruglov et Starobogatov, 1989 F 


● Type locality: eastern Kamchatka, Tumrok Ridge, hot spring (up to 38 C°). 


● Distribution: southeastern Kamchatka (thermal springs of Tumrok area), North Kurile Islands (Shumshu 

and Paramushir) (Kruglov, Starobogatov, 1993a; Starobogatov et al., 2004). 

Lymnaea kurilensis Kruglov et Starobogatov, 1989 F 


● Type locality: Lebyazhje Lake, Iturup Island [Kurile Islands], vicinities of Kuril'sk city. 


● Distribution: South Kurile Islands (Kruglov, Starobogatov, 1993a; Starobogatov et al., 2004). 

Lymnaea lagotis (Schrank, 1803) F 

[Gulnaria lagotis lapidaria B. Dybowski, 1912; G. l. forma ssorensis B. Dybowski, 1912; Lymnaea 

(Radix)auricularia var. lagotis morpha ssorensis Kozhov, 1936] 

Buccinum lagotis Schrank, 1803: 290. 



● Distribution: Europe, West Siberia, Baikal (Kruglov, Starobogatov, 1993b; Sitnikova et al., 2004).

Lymnaea lenaensis Kruglov et Starobogatov, 1985 F 


● Type locality: River Anga valley (upper Lena River, Irkutsk Region) [Siberia]. 


● Distribution: Central Siberia, Amur drainage, Primorje of the Far East (Kruglov, Starobogatov, 1993b). 

Lymnaea likharevi Lazareva, 1967 F 

Lazareva, 1967a: 1346- 1347, figs. 1 (5-5a), 2 (5-5a). 

● Type locality: Borovoe Lake, Kazakhstan. 


● Distribution: southeastern part of East European plain, southern part of West Siberia, Kazakhstan (Kruglov, 

Starobogatov, 1993a). 

Lymnaea liogyra (Westerlund, 1897) F 

Limnaea palustris var. liogyra Westerlund, 1897b: 125. 

● Type locality: “Sibirien. Süd-Ussuri-Gebiet”. 


● Distribution: Amur drainage and Primorje (Kruglov, Starobogatov, 1993a). 

Lymnaea luteola Lamarck, 1822 F 

Lymnaea luteola Lamarck, 1822: 160. 

● Type locality: Bengalia. 

● Types: most probably MHNG (see Dance, 1986). 

● Distribution: Uzbekistan, Tajikistan, Turkmenia (Izzatullaev, 1972). 

Lymnaea mabillei (Locard, 1893) F 

Amphipelpea mabillei Locard, 1893: 47. 

● Type locality: "environs de Troyes (Aube)" (France). 



Lymnaea magadanensis Kruglov et Starobogatov, 1985 F 


● Type locality: Magadan city [Far East], water reservoir of hydroelectric power station. 


● Distribution: northern coast of Okhotsk Sea and upper part of Kolyma basin (Kruglov, Starobogatov, 

1993b). 

Lymnaea manomaensis Kruglov, Starobogatov et Zatrawkin in Kruglov et Starobogatov, 1984 F 


● Type locality: Khabarovsk Territory [Far East], flood land of the Amur River, near Nizhnyaya Manoma 

settlement. 


● Distribution: basin of Amur and rivers entering Tatar Strait, northern Primorje (Kruglov, Starobogatov, 


Lymnaea media (Hartmann, 1840) F 

Limnaea stagnalis var. media Hartmann, 1840: 20, Hartmann 1841: 44. 

● Type locality: "oberbayrischen Seen". 


● Distribution: basins of Baltic and White seas (Starobogatov et al., 2004). 

● Remark: M.V. Vinarski considers Lymnaea media sensu Kruglov and Starobogatov (1985) as in fact different 

species from Limnaea stagnalis var. media sensu Hartmann. This opinion is based on apparent differences

etween shells of L. media sensu Kruglov & Starobogatov and the picture of L. media shell given by Kobelt 

(1877). In the original description, the picture of shell is not given. 

Lymnaea middendorffi (W. Dybowski, 1904) F 

Limnaea peregra var. middendorffi W. Dybowski, 1904: 52, fig. 7. 

● Type locality: "See Sněźinskoje" (Snezhinskoye Lake, Kamchatka). 


● Distribution: eastern Kamchatka, northern and western shores of the Okhotsk Sea, Kolyma River, Lena, 

Yana, and Indigirka rivers basins (Kruglov, Starobogatov, 1993b; Dolgin, 2001; Starobogatov et al., 2004). 

Lymnaea mongolica (Yen, 1939) F 

Pseudosuccinea mongolica Yen, 1939: 68, pl. 15, fig. 54. 

● Type locality: “Dsha-ra-tai, Da-ba-sun, Alashan, Innere Mongolia”. 

● Holotype: SMF 40332. 

● Distribution: along southern and western shores of Khanka Lake (Kruglov, Starobogatov, 1993a; Starobogatov 

et al., 2004), Hövsgöl Lake, Mongolia (Sitnikova et al., 2006). 

Lymnaea monnardi (Hartmann, 1841) F 

Gulnaria monnardi Hartmann, 1841 [1840-1844]: 71, pl. 18 (pl. 6). 

● Type locality: Switzerland. 

● Types: Naturmuseum Saint Gallen, Switzerland. 

● Distribution: Europe (Kruglov, Starobogatov, 1993b). 

Lymnaea mucronata (Held, 1836) F 

Limnaea mucronata Held, 1836: 278. 

● Type locality: not designated by the author, however, his article is devoted to desciption of mollusks of Bavaria, 

so, this country have to be considered as the type locality. 

● Types: Held’s types were destroyed during the World War II (G. Falkner, personal comunication). 


● Remark: The author himself (Held, 1836: 278) believed that this species is identical with Buccinum lagotis 

Schrank. As the type materials of both species are lost, it is impossible to determine the nomenclatorial relatioships 

between these binomens. 

Lymnaea napasica Kruglov et Starobogatov, 1983 F 


● Type locality: near Napas settlement, Tym River, Tomsk Region [W Siberia]. 


● Distribution: middle part of the Ob basin (Kruglov, Starobogatov, 1993b), Mountain Altai (unpublished 

findings from the ZIN collection). 

Lymnaea narzykulovi Izzatullaev, Kruglov et Starobogatov, 1983 F 

Izzatullaev et al., 1983b: 55, fig. 3-4. 

● Type locality: Pamir Mountains, Turumtaikul Lake [Tajik SSR]. 


● Distribution: Pamir (Kruglov, Starobogatov, 1993a). 

Lymnaea nogoonica Kruglov et Starobogatov, 1983 F 


● Type locality: vicinitiesi of Kosh-Agach settlement, Altai Territory [SW Siberia]. 


● Distribution: eastern Altai (Kruglov, Starobogatov, 1993b). 

Lymnaea novikovi Kruglov et Starobogatov, 1983 F


● Type locality: Tarkhatinskoe Lake, Koshagachskij district, Altai Territory [SW Siberia]. 


● Distribution: southeastern part of West Siberia, Altai, southern and central Ural, lower Ob’ basin, Irkutsk 

Region (possibly entire Western Siberia) (Kruglov, Starobogatov, 1993b; Starobogatov et al., 2004; 

M.V.Vinarski, unpublished). 

Lymnaea nuttaliana Lea, 1841 F 

Lea, 1841: 33. 

● Type locality: Oregon. 


● Distribution: Eastern Kamchatka, Koryak upland (Kruglov, Starobogatov, 1993a). 

Lymnaea obensis Kruglov et Starobogatov, 1984 F 


● Type locality: lake on the left bank of the Ob' River, against the Polovinnyj Island [Siberia]. 


● Distribution: lower part of the Ob basin (Kruglov, Starobogatov, 1993b). 

Lymnaea obliquata (Martens, 1864) F 

Limnaeus obliquatus Martens, 1864a: 116, pl. 3, fig. 9-10. 

● Type locality: Issyk-kul. 


● Distribution: Lake Issyk-Kul', Tuva (Prozorova, Sharyi-Ool, 1999; Kruglov, Starobogatov, 1993a). 

Lymnaea oblonga (Puton, 1847) F 

Lymnea oblonga Puton, 1847: 60. 

● Type locality: “Doller (Haut-Rhein)” [upper Rein]. 


● Distribution: Baltic Sea basin and northern Caucasus (Starobogatov et al., 2004). 

Lymnaea ollula (Gould, 1859) F 

Limnaea ollula Gould, 1859: 40. 

● Type locality: streams and marshes of Hong Kong Island. 

● Lectotype (Johnson, 1964): USNM 831. 

● Distribution: temporary waterbodies of southern Primorje (Kruglov, Starobogatov, 1993b; Starobogatov et 

al., 2004). 

Lymnaea ovata (Draparnaud, 1805) F 

[= Gulnaria ovata petricola Lindholm, 1909; Lymnaea (Radix) ovata var. petricola morpha angarensis, morphatschiwirkuensis 

Kozhov, 1936] 

Limnaeus ovatus Draparnaud, 1805: 50, pl. 2, fig. 30-31. 

● Type locality: not specified, in general — France. 

● Types: NHMV. 

● Distribution: Europe, Siberia, Baikal (Kruglov, Starobogatov, 1993b; Sitnikova et al., 2004). 

Lymnaea pachyta (Westerlund, 1885) F 

Limnaea taurica var. pachyta Westerlund, 1885a: 48. 

● Type locality: not specified, probably the same as for taurica (“Krim”). 

● Types: most probably in NMG. 

● Distribution: known from Crimea, Odessa Region, and Volga delta (Kruglov, Starobogatov, 1993a; Son, 

2004).

Lymnaea pacifampla Kruglov et Starobogatov, 1989 F 


● Type locality: mouth of the Gladkaya River, Far East, Posjet Bay [Japan Sea] 


● Distribution: Khanka Lake basin, Primorje (except for northernmost areas) (Kruglov, Starobogatov, 1993a; 


Lymnaea palustris palustris (Müller, 1774) F 

Buccinum palustre Müller, 1774: 131-132. 



● Distribution: Europe, West Siberia (Kruglov, Starobogatov, 1993a). 

Lymnaea palustris syriacus (Mousson, 1861) F 

Linnaeus syriacus Mousson, 1861: 53-54. 

● Type locality: “Damas, Jérusalem”. 


● Distribution: Armenia (Kruglov, Starobogatov, 1993a). 

Lymnaea peregra (Müller, 1774) F 

[= Radix labiata (Rossmässler, 1835) sensu Glöer, 2002]. 

Buccinum peregrum Müller, 1774: 130. 



● Distribution: Europe (Kruglov, Starobogatov, 1993b), Western Siberia (Vinarski et al., 2007). 

Lymnaea perpolita Dall, 1905 F 

Dall, 1905: 78, pl. 2, fig. 6, 8. 

● Type locality: Nushagak, Bristol Bay, Alaska. 


● Distribution: probably eastern Chukchi Peninsula (Kruglov, Starobogatov, 1993b; Starobogatov et al., 

2004). 

Lymnaea persica (Bourgignat in Issel, 1865) F 

Limnaea auricularia var. persica Bourgignat – Issel, 1865: 47. 

● Type locality: “Kerman (Persia meridionale)”. 


● Distribution: southern Azerbaijan (Kruglov, Starobogatov, 1993a). 

Lymnaea petersi Dall, 1905 F 

[= Lymnaea alascensis Baker, 1911] 

Dall, 1905: 66, pl. 2, fig. 3. 

● Type locality: Koyukuk River, N of Yukon, Alaska. 


● Distribution: Chukchi Peninsula (only extreme eastern part), Koryak upland (Kruglov, Starobogatov, 

1993a). 

Lymnaea psilia psilia (Bourguignat, 1862) F 

Limnaea psilia Bourguignat, 1862a: 61. 

● Type locality: “rivière de l’Aube, entre Unienville et Dienville (departement de l’Aube)” [France]. 

● Syntypes: 2 syntypes in MHNG. 

● Distribution: Europe, Siberia, Baikal, Far East: river basins of Kamchatka and Okhotsk Sea to Primorje 

(Kruglov, Starobogatov, 1993a; Starobogatov et al., 2004; Sitnikova et al., 2004).

Lymnaea rectilabrum (Annandale et Prashad, 1919) F 

Limnaea gedrosiana var. rectilabrum Annandale, Prashad, 1919: 49-50, pl. 6, fig. 1-6. 

● Type locality: “Kushdil Khan reservoir, altitude 5000 ft in the hill-country of Baluchistan noerth of Quetta, 

and a small pool in the desert some miles south of Nasratabad in Seistan”. 

● Syntypes: Zoological Survey of India, M 11534/2. 

● Distribution: western Middle East, Tajikistan, southern Uzbekistan, Turkmenistan (waterbodies of Kopet- 

Dagh) (Kruglov, Starobogatov, 1993a). 

Lymnaea saridalensis Mozley, 1934 F 

[Linnaea (Galba) palustris draverti Mozley, 1934] 

Lymnaea palustris saridalensis Mozley, 1934: 2-3, pl. 1, fig. 1. 

● Type locality: small, somewhat saline lake on the Steppe Sari Dala, 15 km southwest of Pavlodar, northern 

Kazakstan. 


● Distribution: entire Western Siberia, Ural, Kazakhstan (Kruglov, Starobogatov, 1993a; M.V.Vinarski, unpublished). 

● Remark: although the holotype of Lymnaea palustris saridalensis was designated in the original publication 

(as “type”), in the type collection of USNM under the catalogue number 469734 there is a lot containing 

15 specimens, labeled as syntypes on a later printed label. It was impossible to isolate the “holotype” and to 

find which specimen matches the original illustration. 

Lymnaea schelechovi Kruglov et Starobogatov, 1989 F 

Kruglov, Starobogatov, 1989b: 28, fig. 1, 2.!; 2, 16. 

● Type locality: Yagelnoye Lake, coast of Yamskaya Inlet, Shelekhov Bay coast, Okhotsk Sea. 


● Distribution: river drainages of Shelekhov Bay and Taui I nlet (Okhotsk Sea) (Kruglov, Starobogatov, 

1993a; Starobogatov et al., 2004). 

Lymnaea schirazensis (Küster, 1863) F 

Lymnaeus schirazensis Küster, 1863: 53, pl. 11, fig. 28-31. 

● Type locality: “Schiraz, Persien”. 


● Distribution: western Central Asia (Kruglov, Starobogatov, 1993a). 

Lymnaea schubinae Kruglov, Starobogatov et Zatrawkin in Kruglov et Starobogatov, 1989 F 


● Type locality: river Bochinka, tributary of the Amur River, area of Amursk city 


● Distribution: Amur basin and Primorje (Kruglov, Starobogatov, 1993a). 

Lymnaea sibirica (Westerlund, 1885) F 

Limnaea truncatula var. sibirica Westerlund, 1885a: 52 

● Type locality: “Sibirien bei Lusino”. 


● Distribution: eastern part of West Siberia, East Siberia, Far East, Amur basin, Primorje, extreme northeastern 

Asia (Kruglov, Starobogatov, 1993a; Starobogatov et al., 2004). 

Lymnaea sihotealinica Kruglov et Starobogatov, 1984 F 


● Type locality: Primorje Territory [Far East], Khorolskij district, stream near the road between Vladimiro- 

Petrovka and Staraja Devitza.


● Distribution: Amur basin, Primorje (Kruglov, Starobogatov, 1993b; Starobogatov et al., 2004). 

Lymnaea sirikulensis (Nevill, 1878) F 

Limnaea defilippi var. sirikulensis Nevill, 1878: 7. 

● Type locality: “Leh in Ladak” [Indian]. 

● Syntypes: Zoological Survey of India, Calcutta. 

● Distribution: Pamir (in particular, the lake Zorkul "Sirikul") (Kruglov, Starobogatov, 1993a). 

Lymnaea solidissima (Kobelt, 1872) F 

Limnaea lagotis var. solidlssima Kobelt, 1872: 77. 

● Type locality: “Himalaya” [probably Issykkul Lake]. 

● Types: ZMB. 

● Distribution: Pamir (Kruglov, Starobogatov, 1993a). 

Lymnaea stagnalis (Linnaeus, 1758) F 

[= Limnaea vulgaris Locard, 1840; Limnaea turgida Locard, 1882; Lymnaea stagnalis var. ssorensiana W. 

Dybowski, 1912; Lymnaea stagnalis var. subulata angarensis B. Dybowski, 1912] 

Helix stagnalis Linnaeus, 1758: 774. 



● Distribution: Europe and North Asia (ponds), Baikal (Kruglov, Starobogatov, 1993a; Sitnikova et al., 

2004). 

Lymnaea streletzkajae Kruglov et Starobogatov, 1985 F 


● Type locality: bog in the Kolyma River (middle part) flood lands, near Agrobaza [NE Siberia]. 


● Distribution: northern coast of Okhotsk Sea and upper part of Kolyma basin (Kruglov, Starobogatov, 

1993b). 

Lymnaea subangulata (Roffiaen, 1868) F 

[= Limnaea truncatula var. ventricosa Moquin-Tandon, 1856, non Hartmann, 1841]. 

Limnaea truncatula var. subangulata Roffiaen, 1868: 78, pl. 1, fig. 9. 

● Type locality: “près d’Alfort” [Switzerland]. 


● Distribution: southern European Russia, Caucasus, western Central Asia (Kruglov, Starobogatov, 1993a; 


Lymnaea subdisjuncta (Nevill, 1878) F 

Limnaea lagotis var. subdisjuncta Nevill, 1878: 9. 

● Type locality: “Leh in Ladak” [Indian]. 

● Syntypes: Zoological Survey of India, Calcutta. 

● Distribution: desert and semidesert regions of Central Asia, to Inner Mongolia in the east (Kruglov, Starobogatov, 

1993a). 

Lymnaea teletzkiana Kruglov et Starobogatov, 1984 F 


● Type locality: Teletzkoe Lake [SW Siberia]. 


● Distribution: Altai (Kruglov, Starobogatov, 1993b). 

Lymnaea tenera (Küster, 1863) F

Limnaeus tener Küster, 1863: 54-55, pl. 12, fig. 1-2. 

● Type locality: “Persien”. 


● Distribution: Tajikistan, southern Uzbekistan (Kruglov, Starobogatov, 1993a). 

Lymnaea tengriana Izzatullaev, Kruglov et Starobogatov, 1983 F 

Izzatullaev et al., 1983a: 323, fig. 1 (5). 

● Type locality: Tajik SSR, Kuljab Region, Muminabad, pool in vicinity of Khan-Tengri Ridge. 


● Distribution: mountain part of Central Asia, probably in inner Altai (Kruglov, Starobogatov, 1993a; Starobogatov 

et al., 2004). 

Lymnaea terebra terebra (Westerlund, 1885) F 

[= Limnaea attenuata Westerlund, 1877, non Say, 1829; Lymnaea (Fossaria) lindholmi W. Dybowski, 1913; 

Galba occulta Jackiewicz, 1959; Lymnaea vulnerata sensu Kruglov & Staroboatov, 1986 non Küster, 1862] 

Limnaea terebra Westerlund, 1885b: 155. 

● Type locality: “Siberia, Tunguska” (collection place of the neotype). 

● Neotype (designated by Vinarski & Glöer, 2008). NMG. 

● Distribution: Central Europe and all Siberia (Vinarski, Glöer, 2008). 

● Remark: Lymnaea (Stagnicola) terebra lindholmi W. Dybowski, 1913 was considered as valid subspecies 

by Kruglov and Starobogatov (1993) and Sitnikova et al. (2004)), but recently Vinarski and Andreev (2003) 

proved that it is not distinguishable from nominotypical subspecies. Starobogatov and Streletzkaya (1967) 

stated, that they designated a lectotype (ZIN, No. 2 in systematic catalogue). Nevertheless, in reality, there is 

no lecotype in the ZIN collection. The sample No. 2 in systematic collection contains 6 empty shells of this 

species collected by Maack in the Lunkha River, Eastern Siberia 

Lymnaea terebra bolotensis Mozley, 1934 F 

Lymnaea (Galba) palustris bolotensis Mozley, 1934: 5-6, pl. 1, fig. 3. 

● Type locality: flooded area between Rivers Chaganak and Chederti, Djala-Uli drainage basin, northern Kazakstan. 


● Distribution: steppes of northern Kazakhstan and south of western Siberia (Kruglov, Starobogatov, 1993a; 


● Remark: the holotype was designated in the original publication as “type” and a lot in USNM type collection 

contains the single specimen labeled as holotype. Nevetheless neither the measurements (shell length 

21.5 vs 22.7 in the original description), not the original illustration do not match the specimen. 

Lymnaea thermobaicalica Kruglov et Starobogatov, 1989 F 

Kruglov, Starobogatov, 1989b: 20, fig. 1, 10-11; 2, 5. 

● Type locality: ZIN (warm spring entering Baikal Lake in the vicinities of Hakusa Bay. 


● Distribution: eastern shore of Baikal, thermal springs (Kruglov, Starobogatov, 1993a). 

Lymnaea thermokamtschatica Kruglov et Starobogatov, 1989 F 


● Type locality: hot spring Khadutka, Kamchatka. 


● Distribution: eastern Kamchatka, thermal springs (Kruglov, Starobogatov, 1993a). 

Lymnaea thiesseae (Clessin, 1879) F 

[= Lymnaea shadini Izzatullaev, Kruglov et Starobogatov, 1983] 

Limnaea truncatula var. thiesseae Clessin, 1879: 4, pl. 1, fig. 2. 

● Type locality: “Ins. Euboaea”.


● Distribution: southeastern Europe, Caucasus (Daghestan), western Central Asia (Kruglov, Starobogatov, 


Lymnaea torquilla (Westerlund, 1876) F 

Limnaea peregra var. torquilla Westerlund, 1876: 55, pl. fig. 11. 

● Type locality: “Jenissei, Surgutskoj (lat. 62° 50’)”. 



Lymnaea truncatula (Müller, 1774) F 

Buccinum truncatulum Müller, 1774: 130-131. 

● Type locality: Thangelstedt, Saxonia (“In agro Thangelstedtiensi Saxoniae”, Müller, 1774: 131). 


● Distribution: Europe, West Siberia eastward to Baikal lake, Tajikistan (Izzatullaev, 1972; Kruglov, Starobogatov, 


Lymnaea tumida (Held, 1836) F 

Limnaea tumida Held, 1836: 278. 

● Type locality: not designated by the author, however, his article is devoted to desciption of mollusks of Bavaria, 

so, this country have to be considered as the type locality. 

● Types: Held’s types were destroyed during the World War II (G. Falkner, personal comunication) 

● Distribution: Europe, South Siberia eastward to Baikal Lake and north Siberia near Norilsk (Kruglov, Starobogatov, 

1993b; Sitnikova et al., 2004:). 

Lymnaea tumrokensis Kruglov et Starobogatov, 1985 F 

Kruglov, Starobogatov, 1985b: 30, fig. 1, 6. 

● Type locality: hot spring, eastern Kamchatka Peninsula, Tumrok Ridge. 


● Distribution: south eastern Kamchatka (thermal sources of Tumrok Range) (Kruglov, Starobogatov, 

1993b). 

Lymnaea turricula (Held, 1836) F 

Limnaea turricula Held, 1836: 278. 

● Type locality: Donautal. 


● Distribution: Europe (Kruglov, Starobogatov, 1993a; Starobogatov et al., 2004). 

● Remark. L. turricula sensu Kruglov & Starobogatov (1986) is not identical with L. turricula sensu 

Jackiewicz (1998). Due to absence of the type series, it is still impossible to deremmine what the true identity 

of the species is. 

Lymnaea ulaganica Kruglov et Starobogatov, 1983 F 


● Type locality: water body near the shore of Surulu-Kol' Lake, Ulaganskij district, Altai Territory [SW Siberia]. 


● Distribution: Altai (Kruglov, Starobogatov, 1993b), southern part of the Western Siberian plain (Vinarski, 

2008). 

Lymnaea ussuriensis Kruglov et Starobogatov, 1989 F 


● Type locality: lake on the border of Khabarovsk [Far East]. 

● Holotype: ZIN No 1 in systematic catalogue.

● Distribution: Amur basin and Primorje (except for northernmost areas) (Kruglov, Starobogatov, 1993a; 


Lymnaea ventricosella B. Dybowski, 1913 F 

Lymnaea (Fossaria) ventricosella B. Dybowski, 1913: 189, pl. 4, fig. 18. 

● Type locality: Bolshoi sor (Baikal). 


● Distribution: Yenissei drainage area, Baikal (Kruglov, Starobogatov, 1993a; Sitnikova et al., 2004). 

Lymnaea viridis Quoy et Gaimard, 1832 F 

Quoy, Gaimard, 1833: 204, pl. 58, figs. 16-18. 

● Type locality: Guam Island (see Hubendick, 1951). 


● Distribution: Primorje Territory, southern Sakhalin, South Kurile Islands (Starobogatov et al., 2004) 

Lymnaea vulnerata (Küster, 1862) F 

Limnaeus vulneratus Küster, 1862: 22, pl. 4, figs. 13-15. 

● Type locality: Cetina River near Omiš, Central Dalmatia, former Yugoslavia (see Jackiewicz, 1988). 


● Distribution: Ukraine, possibly northern Caucasus and Rostov Region (Starobogatov et al., 2004). 

● Remark: From the Jackiewicz’s (1988) opinion, the name L. vulnerata should be attached to a quite different 

species that has another structure of the prostate than L. vulnerata sensu Kruglov & Starobogatov, 1986. 

As Jackiewicz (1988) studied the topotypes, her opinion seems to be preferable (see Vinarski, 2003). Thus, 

the species L. vulnerata cannot be a synonym of Galba occulta Jackiewicz, as Kruglov & Starobogatov 

(1986) believe. However, G. occulta is an apparent junior synonym of an other species, L. terebra, desribed 

from Siberia by Westerlund (1885). The synonymy between Limnaea palustris var. terebra and G. occulta 

was established by Vinarski, Glöer (2008). Thus, L. vulnerata sensu Kruglov & Starobogatov, 1986 is a junior 

synonym of L. terebra (Westerlund) and has to be excluded from the check-list of continental mollusks of 

the former USSR. 

Lymnaea zarenkovi Kruglov et Starobogatov, 1989 F 

Kruglov, Starobogatov, 1989b: 26, fig. I, 23; 2, 18. 

● Type locality: Lebedinskoe Lake, Iturup Island [Kurile Islands]. [According to the paper catalogue of the 

ZIN collection, the holotype of the species is collected in an unnamed lake situated in 1-1.5 km from a river 

that flows out from the Blagodatnoe Lake. The Lebedinskoe Lake is the habitat of on of the paratypes of L. 

zarenkovi] 


● Distribution: Iturup Island (Kruglov, Starobogatov, 1993a). 

Lymnaea zazurnensis Mozley, 1934 F 

Lymnaea (Radix) zazurnensis Mozley, 1934: 6, pl. 1, fig. 2 

● Type locality: Lake Zazurnia, in the mountain range known as Khamar Daban, eastern shore of Lake Baikal 

[Kruglov & Starobogatov (1984a) believe that the true type locality of the species is Snezhnoye Lake near 

Vydrina station, Buryatia]. 


● Distribution: throughout northern Siberia from Ural mountains to Chukchi Peninsula, Altai, mountain parts 

of Transbaikalia, northern part of Amur basin and eastern slopes of Sikhote-Alin, Baikal (Kruglov, Starobogatov, 

1993b; Starobogatov et al., 2004; Sitnikova et al., 2004), China (Anonym, 1985), Hövsgöl Lake, 

Mongolia (Sitnikova et al., 2006). 

● Remark: although the holotype was designated in the original publication (as “type”), the lot in the type 

collection of USNM under the catalogue number 470709 contains single specimen labeled as syntype on a 

later printed label. Since the specimen match the original illustration it is probably a holotype.

PLANORBOIDEA Rafinesque, 1815 

PLANORBIDAE Rafinesque, 1815 


BULININAE P. Fischer et Crosse, 1880 

Culmenella Clench, 1927 

Type species: Bulinus hirasei Clench, 1927 (monotypy) 

Culmenella buldowskii Starobogatov et Prozorova, 1990 F 

Starobogatov, Prozorova, 1990: 33-34, fig. 1 8, 2 5 

● Type locality: Khanka Lake, near village Astrakhanka. 


● Distribution: Primorje except for northeastern part (possibly also in central basin of Amur), in shallow 

permanent waterbodies (Starobogatov et al., 2004). 

Culmenella lindholmi Starobogatov et Prozorova, 1990 F 

Starobogatov, Prozorova, 1990: 31-33, fig. 1 2, 2 4. 

● Type locality: Khanka Lake, near village Astrakhanka. 


● Distribution: Primorje (except for northeastern part) and Amur basin, in shallow permanent and semipermanent 

waterbodies (Starobogatov et al., 2004). 

Culmenella rezvoji (Lindholm, 1929) F 

Glyptophysa rezvoji Lindholm, 1929b: 305-306. 

● Type locality: “Ussuri-Gebiet, Dorf Jakovlevka am Fl. Daubiche”. 


● Distribution: Primorje (except for northeastern part), Amur basin, in shallow shallow permanent waterbodies 


Pettancylus Iredale, 1943 

Type species: Ancylus tasmanicus Tennison-Woods, 1876 (OD) 

Pettancylus wautieri (Mirolli, 1960) F 

Watsonia wautieri Mirolli, 1960: 154, textfigs 1-25. 

● Type locality: Mergozzo Lake (Italy). 


● Distribution: probably Australian species, introduced in Europe (known from Krasnodar Region and cooling 

reservoirs of electric plants, particularly in Moscow Region), Transcaucasia, and central Asia, in permanent 


● Remark: European malacologists usually classify this species in the North American genus Ferrissia 

Walker, 1903 (Falkner et al., 2001; Glöer, 2002). Some taxonomists consider F. watieri to be a junior synonym 

of F. clessiniana (Jickeli, 1882) described from Egypt (Hubendick, 1970; Falkner et al., 2001). Recently, 

a group of American researchers (Walther et al., 2006a, b) have revealed that at least part of European 

populations of F. wautieri are genetically identical with the North American species F. fragilis (Tryon, 1863). 

If so, Australia would have not been the source of this species invasion to Europe. 

Planorbarius Dumeril, 1806 

Type species: Helix carnea Linnaeus, 1758 (monotypy)

Remark: recent molecular studies (Albrecht et al. 2007) revealed that the genus Planorbarius is allied to the 

genera Planorbis, Anisus and Gyraulus belonging to the Planorbinae subfamily and, thus, possibly has not to 

be classified within Bulininae. 

Planorbarius adelosius (Bourguignat, 1859) F 

Planorbis adelosius Bourguignat, 1859: 518, pl. 19, figs. 13-15. 

● Type locality: "les marécages de la Toscane, notamment dans les environs de Pise". 


● Distribution: entire Europe, south Western Siberia, in small permanent waterbodies (Starobogatov et al., 

2004; M.V. Vinarski, unpublished). 

Planorbarius banaticus (Lang in Bourguignat, 1859) F 

Planorbis banaticus Lang – Bourguignat, 1859: 517-518. 

● Type locality: "les eaux stagnantes de la Hongrie, du Banat, de la Transylvanie". 

● Types: possibly in MHNG. 

● Distribution: Europe, possibly south of Western Siberia, in small permanent waterbodies (Starobogatov et 

al., 2004). 

● Remark: Mezzherin et al. (2005) on the basis of the analysis of conchological variability and electrophoretric 

studies synonymised Planorbarius banaticus, P. grandis and P. purpura with Planorbarius 

corneus. 

Planorbarius corneus (Linnaeus, 1758) F 

Helix cornea Linnaeus, 1758: 770. 


● Types: possibly, in UUZM. 

● Distribution: entire Europe, western Siberia to Ob River, in large still and running waterbodies (Pirogov et 

al., 1994; Starobogatov et al., 2004). 

Planorbarius grandis (Dunker in Küster et Dunker, 1850) F 

Planorbis grandis Dunker – Küster, Dunker, 1850: 35, pl. 7, figs. 1-3. 

● Type locality: Germany? (according to the Dunker’s infromation, “Das Vaterland dieser ausgezeichneten 

Schnecke; die Herrn H. Cumings’s Sammlung angehört, ist leider unbekannt”; see Dunker, 1850: 35). 


● Distribution: northwestern Black Sea maritime area, possibly in basin of Baltic Sea, in lakes (Starobogatov 

et al., 2004). 



corneus. 

Planorbarius penchinati (Bourguignat, 1870) F 

Planorbis penchinati Bourguignat, 1870: 39-42, pl. 3, figs. 4-6. 

● Type locality: "le Danube". 

● Types: possibly in NMG. 

● Distribution: northwestern Black Sea maritime area, Baltic Sta basin, in lakes (Starobogatov et al., 2004). 

Planorbarius pinguis (Westerlund, 1885) F 

Planorbis corneus var. pinguis Westerlund, 1885b: 66. 

● Type locality: "Königsberg". 

● Types: not traced, possibly in NMG 

● Distribution: basins of Baltic, Black, Azov and Caspian seas, in lakes (Starobogatov et al., 2004).

Planorbarius purpura (Müller, 1774) F 

Planorbis purpura Müller, 1774: 154-157. 


● Types: unknown, possibly in ZMC (see Dance, 1986). 

● Distribution: entire Europe, western Siberia, Kazakhstan, in still waterbodies (Starobogatov et al., 2004). 



corneus. 

Planorbarius stenostoma (Bourguignat in Servain, 1881) F 

Planorbis stenostoma Bourguignat – Servain, 1881: 82. 

● Type locality: "des bouches du Danube". 


● Distribution: steppe areas of south Europe and Western Siberia to Altai, in temporary waterbodies (Starobogatov 

et al., 2004). 

● Remark: in spite of the data provided by Starobogatov et al. (2004), there is no reliable findings of this species 

in waterbodies of Western Siberia (Karimov, 2005). 

Planorbella Gabb, 1873 

Type species: Planorbella imitans Gabb, 1873 (OD) 

Planorbella hornii (Tryon, 1865) F 

Planorbis hornii Tryon, 1865: 231, pl. 22, fig. 16. 

● Type locality: "Fort Simpson, British America". 


● Distribution: vicinities of Anadyr (Chukotka) (Starobogatov et al., 2004). 

Planorbella subcrenata (Carpenter, 1857) F 

Planorbis subcrenatus Carpenter, 1857: 220-221. 

● Type locality: Oregon. 


● Distribution: so far recorded only in Pleistocene deposits in lower Kolyma basin (Starobogatov et al., 

2004). 

Seminolina Pilsbry, 1934 

Type species: Seminolina eudiscus (fixation of the type species not traced). 

Seminolina eudiscus (Pilsbry, 1934) F 

Helisoma duryi eudiscus Pilsbry, 1934a: 42, pl. 9, figs. 4-9. 

● Type locality: Miami River above Miami, Florida. 

● Holotype: ANSP 77078. 

● Distribution: sometimes settles in waterbodies of warm regions and cooling reservoirs of electric plants; 

particularly, the species was living in irrigation ditches in Dushanbe for several years (Starobogatov et al., 

2004). 

● Remark: European malacologists mentioned this species under the name Planorbella duryi (Wetherby, 

1879); it has been introduced to Europe from Northern America (Glöer, 2002). The genus Seminolina is usually 

regarded as a subgenus of the genus Planorbella. 

PLANORBINAE Rafinesque, 1815 

Ancylus Müler, 1774 

Type species: Ancylus fluviatilis Müller, 1774 (SD ICZN Opinion 363, 1955)

Ancylus armenia Boettger, 1881 F 

Ancylus fluviatilis var. armenia Boettger, 1881c: 128. 

● Type locality: "Eriwan et lacum Chosapinskensem Armeniae". 

● Types: "Lectotype" SMF No. 14976a, "paralectotypes" Nos. 14976b/3, 149674/4. 

● Distribution: mountain springs of Armenia (Soldatenko, Starobogatov, 2004). 

Ancylus benoitianus Bourguignat, 1862 F 

Bourguignat, 1862b: 180-181. 

● Type locality: "Sicile". 

● Types: not traced, possibly in MHNG. 

● Distribution: western Transcaucasia, northern Osetia, southern Daghestan, above water level in mountain 

springs (Starobogatov et al., 2004; Soldatenko, Starobogatov, 2004). 

Ancylus capuloides Jan in Porro, 1838 F 

[= Ancylus fluviatilis var. capuliformis Moquin-Tandon, 1855] 

Porro, 1838: 87-88, pl. 1, fig. 7. 

● Type locality: "lago di Como al prato Pasquee ed allo sbocco de torrente Breccia; ... torrente Bevera in Brianza". 


● Distribution: Ciscaucasia and Transcaucasia, in rivers and springs (Starobogatov et al., 2004; Soldatenko, 

Starobogatov, 2004). 

Ancylus expansilabris Clessin, 1882 F 

Clessin, 1882b: 51, pl. 6, fig. 12. 

● Type locality: "Deutschland. Waldes, Falkenstein, im Regenfluss. Belgien.". 

● Types: the Clessin’s collection has been destroyed after World War II (P. Glöer, pers. communication). 

● Distribution: possibly in Russia, where it is confused with A. fluviatilis (Starobogatov et al., 2004; 

Soldatenko, Starobogatov, 2004). 

Ancylus fluviatilis Müller, 1774 F 

Müller, 1774: 201-202. 

● Type locality: “Saxonia”. 


● Distribution: entire Europe and Ural, Ural River, in rivers and springs (Pirogov et al., 1994; Starobogatov et 

al., 2004). 

Ancylus gibbosus Bourguignat, 1853 F 


● Type locality: “départment de l’Oise (Baudan), des environs de Verdun (Liénard), enfin de plusieurs petits 

russeaux de Vendeuvre-sur-Barge (Aule)”. 

● Syntypes: not traced, possibly in MHNG. 

● Distribution: mountain rivers of Tanscaucasia and Ciscaucasia (Soldatenko, Starobogatov, 2004). 

Ancylus jani Bourguignat, 1853 F 

[= Ancylus fluviatilis sensu Akramowski, 1976, non Linnaeus, 1758] 


● Type locality: “Jan, dans la Lombardie”. 

● Syntypes: not traced, possibly in MHNG. 

● Distribution: mountain rivers of Transcaucasia and Ciscaucasia (Akramowski, 1976; Soldatenko, Starobogatov, 

2004). 

Ancylus major Issel, 1865 F 

Ancylus jani var. major Issel, 1865: 428, pl. 3, figs. 58-61.

● Type locality: "cittadella di Erivan in Armenia". 


● Distribution: mountain springs of Armenia (Soldatenko, Starobogatov, 2004). 

Ancylus orbicularis Held, 1837 F 

Held, 1837: 305. 

● Type locality: Bavaria (judging from the title of the Held’s article). 


● Distribution: western Ukraine, possibly in Euripean Russia where it is confused with A. fluviatilis (Starobogatov 

et al., 2004). 

Ancylus subcircularis Clessin, 1882 F 

Clessin, 1882b: 33. 

● Type locality: "Bis jetzt nur bei Reichenberg in Böhmen". 


● Distribution: running waterbodies of Europe, in Russia in St.-Petersburg (Soldatenko, Starobogatov, 2004). 

Anisus Studer, 1820 

Type species: Helix spirorbis Linnaeus, 1758 (SD Gray, 1847) 

Anisus acronicus (Férussac, 1807) F 

[=Planorbis deformis Férussac, 1807; Planorbis gredleri Bielz in Gredler, 1859, partim; Planorbis devians 

var. euphratica Mousson, 1874; Planorbis rossmässleri Hartwich, 1914] 

Planorbis acronicus Férussac, 1807: 105. 

● Type locality: Germany, Überlinger See (= Acronicus lacus), the northern part of the Bodensee (P. Glöer, 

pers. communication). 


● Distribution: northern Eurasia, eastward to basin of Nizhnyaya Tunguska (Pirogov et al., 1994; Prozorova, 

2003; Starobogatov et al., 2004); Armenia (Akramowski, 1976). 

Anisus agardhi Starobogatov in Prozorova, 2003 F 

Anisus (Bathyomphalus) agardhi “Starobogatov, 1996” – Prozorova, 2003: 100 (nom. nov. pro Planorbis 

contortus var. labiatus Westerlund, 1874 non Planorbis labiatus Benson, 1850). 



● Distribution: Balthic Sea basin, in permanent and semipermanent waterbodies (Prozorova, 2003; Starobogatov 

et al., 2004). 

● Remark: Although this species name was at least twice cited in literature as of “Starobogatov, 1996", it has 

not been published in that year, and, as far as we know, not before 2003. 

Anisus albus (Müller, 1774) F 

[=Planorbis hispidus Draparnaud, 1805; Planorbis albus var. cinctutus Westerlund, 1871] 

Planorbis albus Müller, 1774: 164. 



● Distribution: Europe, Ural River, western Siberia plain, eastward to Altai (Pirogov et al., 1994; Prozorova, 

2003; Starobogatov et al., 2004; Karimov, 2005); Transcaucasia (Akramowski, 1976). 

Anisus amuricus Prozorova et Starobogatov, 1998 F 

Prozorova, Starobogatov, 1998b: 57, fig. 1D. 

● Type locality: Khabarovsk District, lake in the plain of Burea River, 3 km dowstream from mouth of Chekunda 

River. 


● Distribution: central and lower Amur basin, rivers of Primorje (Prozorova, Starobogatov, 1998b; Prozorova, 


Anisus baicalicus (B. Dybowski, 1913) F 

Planorbis (Gyraulus) baicalicus B. Dybowski, 1913: 200, pl. 5, fig 26. 


● Lectotype (Starobogatov, Streletskaya, 1967): ZIN, No. 1 in systematic catalogue. 

● Distribution: Yenissei River basin, northern Baikal Lake area, Baikal, 1-5 m (Starobogatov, Streletskaya, 

1967; Prozorova, 2003; Starobogatov et al., 2004; Sitnikova et al., 2004), south of western Siberia plain 

(Karimov, 2005). 

Anisus bavaricus (Westerlund, 1885) F 

Planorbis charteus var. bavaricus Westerlund, 1885a: 72. 

● Type locality: "Bayern, Schweden". 


● Distribution: entire Europe except extreme north, south of Western Siberia, northern Kazakhstan, in permanent 

waterbodies (Prozorova, 2003; Starobogatov et al., 2004). 

Anisus borealis (Lovén in Westerlund, 1875) F 

[= Anisus (Gyraulus) acronicus, part sensu Starobogatov, 1970, non Férussac, 1807; Gyraulus gredleri var. 

borealis Lovén, 1845] 

Planorbis borealis Lovén – Westerlund, 1875: 112-113, pl. 2, figs. 23-25. 

● Type locality: "Norwegia, Lapponia & Fennia". 


● Distribution: northern Eurasia, eastward to Kamchatka, Baikal Lake, in lakes and rivers (Prozorova, 2003; 

Starobogatov et al., 2004; Sitnikova et al., 2004). 

Anisus buriaticus Prozorova et Starobogatov, 1997 F 

Prozorova, Starobogatov, 1997a: 44, fig. 2A. 

● Type locality: Transbaikalia, vicinity of town Gusinohozyorsk, Buriat Republic. 


● Distribution: Amur Basin, Primorje, Transbaikalia, Selenga River drainage area, Sakhalin (Prozorova, 

Starobogatov, 1997a; Prozorova, 2003; Starobogatov et al., 2004). 

Anisus carinea (Westerlund, 1897) F 

Planorbis (Diplodiscus) vortex var. carinea Westerlund, 1897b: 126. 

● Type locality: "Europäisches Russland. Gouv. Perm, Bilimbajewskaja". 


● Distribution: entire Europe except extreme north, in permanent waterbodies (Prozorova, 2003; Starobogatov 

et al., 2004). 

Anisus centrifugops Рrozorova et Starobogatov, 1997 F 

Рrozorova, Starobogatov, 1997a: 49-50 [nom. nov. pro Anisus Centrifugus sensu Bogatov et Zatrawkin, 

1991[1990], non Planorbis centrifugus Westerlund, 1897]. 

● Type locality: Amur River. 


● Distribution: Amur Basin and Primorje, in permanent and semipermanent waterbodies (Prozorova, 2003; 


Anisus centrifugus (Westerlund, 1897) F 

Planorbis centrifugus Westerlund, 1897b: 141-142. 

● Type locality: "Werchne-Kolymsk". 

● Lectotype (selected by Рrozorova, Starobogatov, 1997a): ZIN, No. 1 in systematic catalogue.

● Distribution: Kolyma, Indigirka, Yana and Lena rivers basins, except upper part, and Taui Inlet river basin 

(Okhotsk Sea), in permanent waterbodies (Prozorova, 2003; Starobogatov et al., 2004; M.V. Vinarski, unpublished). 

Anisus charteus (Held, 1837) F 

Planorbis charteus Held, 1837: 305. 

● Type locality: Bavaria (judging from the title of the Held’s article). 



et al., 2004). 

Anisus chereshnevi Prozorova et Starobogatov, 1997 F 

Prozorova, Starobogatov, 1997a: 48, fig. 3B. 

● Type locality: hot springs in the valey of the Gilmimliveyem River at the oulets of thermal waters, Chukchi 

Peninsula. 


● Distribution: Chukchi Peninsula, in thermal springs (Prozorova, Starobogatov, 1997a; Prozorova, 2003; 


Anisus concinnus (Westerlund, 1881) F 

Planorbis (Gyraulus) concinnus Westerlund, 1881: 63. 

● Type locality: "Suecia ad Ronneby; Norvegia in reg. Gudbrandsdal". 


● Distribution: Baltic Sea basin, in permanent waterbodies (Prozorova, 2003). 

Anisus contortus (Linnaeus, 1758) F 

Helix contorta Linnaeus, 1758: 770. 


● Types: unknown, possibly in UUZM. 

● Distribution: entire Europe, eastward to Ural River, probably in Irtysh basin, in permanent warterbodies 

(Pirogov et al., 1994; Prozorova, 2003; Starobogatov et al., 2004); northern Asia, Ciscaucasia (Akramowski, 

1976); Western Siberia (M.V. Vinarski, unpublished). 

Anisus correctus (Westerlund, 1897) F 

Planorbis correctus Westerlund, 1897a: 123-124. 

● Type locality: "Fennia ad Harjula in Kuopio". 


● Distribution: Baltic Sea basin, in springs (Prozorova, 2003; Starobogatov et al., 2004). 

Anisus convexiusculus (Hutton, 1849) F 

Planorbis convexiusculus Hutton, 1849: 657. 

● Type locality: Candahar, Afghanistan (fide Clessin, 1884). 


● Distribution: Vaksh River, Tajikistan (Izzatullaev, 1972). 

Anisus crassus (Da Costa, 1778) F 

[= Planorbis (Bathyomphalus) contortus sensu Lindholm, 1909; B. Dybowski, 1912; Kozhov, 1936, part., 

non Linnaeus, 1758] 

Helix crassa Da Costa, 1778: 66, pl. 4, fig. 11. 

● Type locality: the Thames near London; rivulets about Peterborough-bouse, Westminster (England). 


● Distribution: Europe, Siberia to Kolyma River, Baikal, in permanent waterbodies (Sitnikova et al., 2004; 

Prozorova, 2003; Starobogatov et al., 2004).

Anisus dazuri (Mörch, 1868) F 

Planorbis dazuri Mörch, 1868: 27 (nom. nov. pro Planorbis spirorbis sensu Rossmässler, 1835, non Müller, 

1774). 

● Type locality: "Bei Kassel, in Schweden und Frankreich" (Rossmässler, 1835a: 106). 


● Distribution: entire Europe, except northern areas, southern Ural and south of Western Siberia, in temporary 

warterbodies (Prozorova, 2003; Starobogatov et al., 2004; Karimov, 2005). 

Anisus discoides (Reinhardt, 1870) F 

Planorbis vortex var. discoides Reinhardt, 1870: 24. 


● Syntypes: SMF No. 147619/10. 

● Distribution: entire Europe except extreme north, in lakes (Prozorova, 2003; Starobogatov et al., 2004). 

Anisus dispar (Westerlund, 1871) F 

Planorbis dispar Westerlund, 1871: 131. 

● Type locality: "In Suecia haec species formis allatis pertotum regnum, a maxime meridionali parte prov. 

Skеne saltem usque in Luleе Lappmark, ad. 67-68 lat. bor., distributa, locis plerisque (prov. Blekinge, ubi 

rara, excepta) frequens. In Norvegia passim as Christiania et Bergen.". 


● Distribution: Europe, in permanent waterbodies (Prozorova, 2003; Starobogatov et al., 2004). 

● Remark: although species was repeatedly reported from Western Siberia, no specimens from this area are 

present in Russian malacological collections (Karimov, 2005). 

Anisus (Gyraulus) draparnaudi (Sheppard, 1823) F 

[= Planorbis limophilus Clessin, 1884] 

Helix draparnaudi Sheppard, 1823: 158-159. 

● Type locality: Holbrook mill-pond. 


● Distribution: Europe, western Siberia, northern Kazakhstan, Tuva, in permanent waterbodies (Prozorova, 


Anisus eichwaldi (Grimm in W. Dybowski, 1888) М 

[= Planorbis micromphalus Grimm, 1876, non Fuchs, 1870] 

Planorbis eichwaldi Grimm – W. Dybowski, 1888: 49-50, pl. 2, fig. 11a-c, pl. 3, fig. 10a-c. 


● Syntypes: ZIN? 


Anisus hypocyrtus Servain, 1888 F 

[= Planorbis compressus Michaud, 1831 non de Serres, 1818] 

Planorbis hypocyrtus Servain, 1888: 299-300, pl. 7, figs. 6-9. 

● Type locality: not stated ("environs de Hambourg" – from title). 


● Distribution: entire Europe except extreme north, Western Siberia, in lakes (Prozorova, 2003; Starobogatov 

et al., 2004; Karimov, 2005). 

Anisus ignotellus (B. Dybowski, 1913) F 

Gyraulus ignotellus B. Dybowski, 1913: 204, pl. 5, fig.. 


● Lectotype (Sitnikova et al., 2004): ZIN, No.1 in systematic catalogue. 

● Distribution: Kultuk, Maloe More (Baikal), 3-110 m (Sitnikova et al., 2004).

Anisus illibatus (Westerlund, 1883) F 

Planorbis (Gyraulus) illibatus Westerlund, 1883b: 53. 

● Type locality: Onuyo, Saitama Pref., Honshu, Japan. 

● Types: SMNH. 

● Distribution: South Kurile Islands (Iturup and Zeiyoniy) (Prozorova, 1996[1997]a; Prozorova, 2003). 

Anisus infirmus Mori, 1938 F 

Anisus (Gyraulus) laevis infirmus Mori, 1938a: 280, 284, 298, pl. 18, fig. 16. 

● Type locality: a river flowing into Itubisinai-ko, Tisima. 


● Distribution: South Kurile Islands (Kunashir and Zeiyony), southern part of Sakhalin Island, in cold springs 

and lakes (Prozorova, 1996[1997]a; Prozorova, 2003; Starobogatov et al., 2004). 

Anisus infraliratus (Westerlund, 1876) F 

Planorbis infraliratus Westerlund, 1876a: 12-13. 

[= Planorbis polaris Westerlund, 1885] 

● Type locality: "Jenissei, Nischnij Inbatsk (63º50')". 


● Distribution: Siberia from Yenissei to Chukotka, Tuva, in permanent waterbodies (Prozorova, 2003; Starobogatov 

et al., 2004). 

Anisus issykulensis (Clessin, 1907) F 

Gyraulus issykulensis Clessin, 1907: 8. 

● Type locality: not stated (Issyk-Kul Lake – from title). 


● Distribution: Issyk-Kul Lake (Izzatullaev, 1972). 

Anisus iturupensis Prozorova et Starobogatov, 1997 F 

Prozorova, Starobogatov, 1997a: 48-49, fig. 3C. 

● Type locality: Solnechnoye Lake, Iturup Island (southern Kurile Islands). 


● Distribution: South Kurile Islands, in permanent waterbodies (Prozorova, Starobogatov, 1997a; Prozorova, 


Anisus japonica (Martens, 1867) F 

[= Anisus (Gyraulus) albus Mori, 1938] 

Planorbis compressus var. japonicus Martens, 1867: 214. 

● Type locality: Japan, near Yokohama. 

● Holotype: whereabouts unknown. 

● Distribution: South Kurile Islands (Prozorova, 1996[1997]a). 

Anisus johanseni (Mozley, 1934) F 

Planorbis (Spiralina) johanseni Mozley, 1934: 6-7, pl. 1, fig. 8. 

● Type locality: Kotur Kulb, near Borovoye, Kazakstan. 


● Distribution: northern Kazakhstan, in lakes (Prozorova, 2003; Starobogatov et al., 2004). 

● Remark: although species was reported from southern part of Western and Central Siberia (eg. by Starobogatov 

et al., 2004), it is still not found there (Karimov, 2005). It is possible that the species is just a junior 

synonym of A. vortex (M.V. Vinarski, unpublished). The holotype was designated in the original publication 

as “type”. Nevertheless in the type collection of USNM the lot with the catalogue number 

470515 contains 21 specimens labeled as “syntypes” on a later printed label. None of them matched the 

original illustration.

Anisus kamtschaticus (Westerlund, 1876) F 

[= Planorbis borealis var. dershavini Rozen, 1927] 

Planorbis (Gyraulus) kamtschaticus Westerlund, 1876b: 127. 

● Type locality: "Sibirien. Halbinsel Kamtschatka, im Flusse Kamtschatka". 


● Distribution: north-east Asia from Lena River basin in the west and Amur River basin in the south, in permanent 


Anisus khabarovskiensis Prozorova et Starobogatov, 1997 F 

Prozorova, Starobogatov, 1997a: 45, fig. 2D. 

● Type locality: Innokentievskoye Lake (Khabarovsk part of Amur drainage area). 


● Distribution: middle and lower parts of Amur drainage area, in permanent or semipermanent waterbodies 

(Prozorova, Starobogatov, 1997a; Prozorova, 2003; Starobogatov et al., 2004). 

Anisus kolesnikovi Logvinenko et Starobogatov, 1966 М 

[= Planorbis eichwaldi var. dybowski Kolesnikov, 1947; Anisus kolesnikovi sublitoralis Logvinenko et Starobogatov, 

1966] 

Logvinenko, Starobogatov, 1966b: 1473-1474, figs. 5,6. 

● Type locality: Caspian Sea, 40°37’ N, 50°52’ E, 115 m. 


● Distribution: middle and southern Caspian Sea; 30-200 m (Logvinenko, Starobogatov, 1966b, 1968). 

Anisus kruglowiae (Johansen, 1937) F 

Planorbis kruglowiae Johansen, 1937: 191, figs. 1-3. 

● Type locality: Mountain Altai, Lake Tenginskoye. 


● Distribution: Lake Tenginskoye, lakes of Burlinsk water system of the plain Altai (Prozorova, 2003; Starobogatov 

et al., 2004; Karimov, 2005). 

Anisus kussakini Prozorova et Starobogatov, 1997 F 

Prozorova, Starobogatov, 1997a: 46, fig. 2F. 

● Type locality: branch of the Anadyr River with slightly brackish water, entering the Ugolnaya Inlet. 


● Distribution: Chukchi Peninsula, Kamchatka, possibly Kolyma River basin, in semipermanent waterbodies 

(Prozorova, Starobogatov, 1997a; Prozorova, 2003; Starobogatov et al., 2004). 

Anisus ladacensis (Nevill, 1878) F 

[= Gyraulus ehrenbergi (Beck) sensu Zhadin, 1950, non Beck, 1837; Gyraulus laevis sensu Zhadin, 1950, 

non Alder, 1833] 

Planorbis (Gyraulus) laevis var. ladacensis Nevill, 1878: 10-11. 

● Type locality: Leh, a region of Ladak (Tibet). 

● Types: Indian Museum (Natural History), Calcutta. 

● Distribution: Central Asia (Izzatullaev, 1972; Prozorova, 2003). 

Anisus laevis (Alder, 1838) F 

Planorbis laevis Alder, 1838: 337. 

● Type locality: “in a pond at Whitley Quarries”. 


● Distribution: Europe, except south and south-east (Prozorova, 2003; Starobogatov et al., 2004); Transcaucasia 


● Remark: Planorbis laevis Faure et Sionest, 1808 – probable homonymy.

Anisus leucostoma (Millet, 1813) F 

[= Planorbis rotundatus W. Dybowski, 1912; Planorbis (Anisus) leucostoma var. nikolensis Kozhov, 1936] 

Planorbis leucostoma Millet, 1813: 16-17. 

● Type locality: St Georges de Boix ca 150 km apart from Angers, departement Charente-Maritime, France 

(the sampling site of the neotype, see Glöer & Meier-Brook, 2008) 

● Neotype: Zoological Museum Hamburg (collection number ZMH 51204) (designated by Glöer and Meier- 

Brook, 2008). 

● Distribution: Europe and western Siberia, eastward to left bank of Lena and Nizniya Tunguska rivers, in 

temporary waterbodies (Pirogov et al., 1994; Prozorova, 2003; Starobogatov et al., 2004; Sitnikova et al., 

2004; M.V. Vinarski, unpublished); Armenia (Akramowski, 1976). 

Anisus minusculus Moskvicheva in Dvoriadkin, 1980 F 

Anisus (Microanisus) minusculus Moskvicheva – Dvoriadkin, 1980: 25, fig. 3. 

● Type locality: Semitka River (Amgun’ River basin) in the vicinities of the settlement Imeni Poliny 

Osipenko. 


● Distribution: Amur basin, Primorje, in temporary and semitemporary waterbodies (Prozorova, 2003; Starobogatov 

et al., 2004). 

Anisus nikolensis (B. Dybowski, 1913) F 

Gyraulus dazuri var. nikolensis B. Dybowski, 1913: 194, pl. 5, fig. 23. 

● Type locality: Bolshoi Posolskyi Sor [Baikal]. 


● Distribution: central and eastern Siberia, Baikal Lake and vicinities, coast of the Okhotsk Sea (lower 

reaches of Uda River), 0-0.5 m (Sitnikova et al., 2004; Prozorova, 2003; Starobogatov et al., 2004). 

Anisus noziriensis Mori, 1938 F 

Anisus (Gyraulus) albus noziriensis Mori, 1938a: 280, 284, 296, pl. 17, fig. 14. 

● Type locality: Noziri-ko, Nagano-ken. 


● Distribution: southern Sakhalin, South Kurile Islands (Kunashir and Zelenyi), in permanent and semipermanent 

waterbodies (Prozorova, 1996[1997]a; Prozorova, 2003; Starobogatov et al., 2004). 

Anisus nummulus (Held, 1837) F 

Planorbis nummulus Held, 1837: 304. 

● Type locality: Bavaria (judging from the title of the Held’s paper). 



et al., 2004). 

Anisus perezii (Graëlls in Dupuy, 1850) F 

Planorbis perezii Graëlls – Dupuy, 1850: 441, pl. 25, fig. 6. 

● Type locality: "les environs d'Arles, de Troyes, la forкt de Raismes [nord]". 


● Distribution: basins of Black and Baltic seas, in shallow temporary waterbodies (Prozorova, 2003; Starobogatov 

et al., 2004). 

Anisus septemgyratus (Ziegler in Rossmässler, 1835) F 

Planorbis septemgyratus Ziegler – Rossmässler, 1835a: 106, pl. 2, fig. 64. 

● Type locality: Bolzano, North Italy (fide Glöer & Meier-Brook, 2008). 


● Distribution: south of Europe, Baltic sea basin, Black Sea maritime area to Altai, in shallow temporary wa-

terbodies (Prozorova, 2003; Starobogatov et al., 2004). 

Anisus sibiricus (Dunker, 1848) F 

Planorbis sibiricus Dunker, 1848: 42. 

● Type locality: "Sibiria". 


● Distribution: eastern Siberia, Yenissei basin, north of Far East, in small semipermanent waterbodies (Prozorova, 


Anisus spirillus (Gould, 1859) F 

Planorbis spirillus Gould, 1859: 40. 

● Type locality: Ousima [Amami-O-Shima, Ryukyu Islands]. 

● Lectotype (Johnson, 1964): USNM 1557. 

● Distribution: South Kurile Islands, southern Sakhalin Island (Prozorova, 1996[1997]a; Prozorova, 2003). 

Anisus spirorbis (Linnaeus, 1758) F 

Helix spirorbis Linnaeus, 1758: 770. 

● Type locality: “in Europae Lacubus, magnitudine Sem. Anethi”. 


● Distribution: entire Europe, western Siberia, northern Kazakhstan, Ural River (Pirogov et al., 1994; Prozorova, 

2003; Starobogatov et al., 2004; Karimov, 2005); Armenia (Akramowski, 1976). 

Anisus spondyloides (Weinland, 1883) F 

Planorbis contortus var. spondyloides Weinland, 1883: 122. 

● Type locality: "Schönthal, Württemberg". 


● Distribution: Baltic Sea basin, in permanent waterbodies waterbodies (Prozorova, 2003; Starobogatov et 

al., 2004). 

Anisus sretenskiensis Prozorova et Starobogatov, 1997 F 

Prozorova, Starobogatov, 1997a: 44-45, fig. 2B. 

● Type locality: Transbaikalia, vicinity of town Sretensk, Shilka River drainage area, Chita Region. 


● Distribution: Transbaikalia, Shilka River drainage area (Prozorova, Starobogatov, 1997a; Prozorova, 2003; 


Anisus stelmachoetius (Bourguignat. 1860) F 

[= Planorbis stelmachoetius var. notatus Westerlund, 1885] 

Planorbis stelmachoetius Bourguignat, 1860b: 139, pl. 2, fig. 10-13. 

● Type locality: small rivers near Dinan. 


● Distribution: Europe and south of Western Siberia, Tuva, in permanent waterbodies (Pirogov et al., 1994; 

Prozorova, 2003; Starobogatov et al., 2004). 

Anisus strauchianus (Clessin, 1884) F 

Planorbis strauchianus Clessin, 1884: 204-205, pl. 31, fig. 5. 

● Type locality: "Rissland Ukiow". 


● Distribution: south of Europe, in temporary waterbodies (Prozorova, 2003; Starobogatov et al., 2004). 

Anisus stroemi (Westerlund, 1881) F 

[= Planorbis (Gyraulus) angasolensis B. Dybowski, 1912] 

Planorbis strömi Westerlund, 1881: 63.

● Type locality: "Norvegia, Fennia (ad Oulu et Nuottasaari Ostrobottniae), Siberia" (from original description). 

● Syntypes: two syntypes NMG (studied by Glöer and Vinarski, 2009). 

● Distribution: northern Europe, western and eastern Siberia westward to the Okhotsk Sea coast, southward 

to Amur basin and Primorje, Baikal, 0-5 m (Prozorova, 2003; Sitnikova et al., 2004; Starobogatov et al., 

2004). 

● Remark: syntypes were collected in Norway. Although in the original description the range rather than the 

locus typicus was given, later Westerlund (1897a: 120) mentioned for Norway the only sampling site “ad 

Eker (in Rökkebjergstjern)” (Glöer, Vinarski, 2009). 

Anisus subfiliaris Moskvicheva in Dworiadkin, 1980 F 

Anisus (Gyraulus) subfiliaris Moskvicheva – Dworiadkin, 1980: 25, fig. 2. 


● Lectotype (Prozorova, Starobogatov, 1998b): ZIN. 

● Distribution: southern Far East from upper reaches of the Amur to Sakhalin, in semipermanent waterbodies 

(Prozorova, 2003; Starobogatov et al., 2004). 

Anisus substroemi Starobogatov et Budnikova, 1976 F 

Starobogatov, Budnikova, 1976: 85-86, fig. 5 V. 

● Type locality: Anadyrskij district, pool near Vakarevo [Chukchi Peninsula] 


● Distribution: Chukchi Peninsula, northern Okhotsk Sea maritime area, in permanent and small semipermanent 


Anisus sulcatus Logvinenko et Starobogatov, 1966 М 

Logvinenko, Starobogatov, 1966b: 1474, fig. 7. 

● Type locality: 42°45’ N, 48°29’ E, 79 m. 


● Distribution: middle Caspian Sea; 75-150 m (Logvinenko, Starobogatov, 1966b, 1968). 

Anisus terekholicus Prozorova et Starobogatov, 1997 F 

Prozorova, Starobogatov, 1997a: 45, fig. 2C. 

● Type locality: Lake Terekhol, Tuva Republic. 


● Distribution: southern Tuva (Lake Terekhol) (Prozorova, Starobogatov, 1997a; Prozorova, 2003; Starobogatov 

et al., 2004), Hövsgöl Lake, Mongolia (Sitnikova et al., 2006). 

Anisus thermochukchensis Prozorova et Starobogatov, 1997 F 

Prozorova, Starobogatov, 1997a: 46, 47, fig. 3A. 

● Type locality: hot springs in the valey of the Gilmimliveyem River at the oulets of thermal waters, Chukchi 

Peninsula. 


● Distribution: eastern Chukchi Peninsula, in warm springs with the temperature up to 42 о (Prozorova, Starobogatov, 

1997a; Prozorova, 2003; Starobogatov et al., 2004). 

Anisus tugurensis Prozorova et Starobogatov, 1997 F 

Prozorova, Starobogatov, 1997a: 45-46, fig. 2E. 

● Type locality: Tugur River valley. 


● Distribution: Tugur River basin (southern Okhotsk Sea maritime area), in permanent and semipermanent 

(Prozorova, Starobogatov, 1997a; Prozorova, 2003; Starobogatov et al., 2004).

Anisus umbiliciferus (Kozhov, 1936) F 

Planorbis (Gyraulus) umbiliciferus Kozhov, 1936: 150, pl. 6, figs. 44-47. 



● Distribution: Baikal: northern part and Maloe More (Sitnikova et al., 2004). 

Anisus vortex (Linnaeus, 1758) F 

Helix vortex Linnaeus, 1758: 770. 



● Distribution: entire Europe except extreme north, Ural River, Western and Central Siberia, in permanent 

waterbodies (Pirogov et al., 1994; Dolgin, 2001; Prozorova, 2003; Starobogatov et al., 2004; Karimov, 2005). 

Anisus vorticulus (Troschel, 1834) F 

Planorbis vorticulus Troschel, 1834: 51. 

● Type locality: not stated, possibly in vicinities of Berlin (“Exemplaria, quibus usus sum, debeo viro harum 

rerum curiosissim Graffio, mechanico Berolinensi”), see Glöer (2002). 


● Distribution: Baltic Sea basin, in lakes (Prozorova, 2003; Starobogatov et al., 2004). 

Anisus zelenensis Prozorova et Starobogatov, 1997 F 

Prozorova, Starobogatov, 1997a: 49, fig. 3D. 

● Type locality: Utinoye Lake, Zeleny Island (Lesser Kurile Islands – Habomai). 


● Distribution: South Kurile Islands, in lakes (Prozorova, Starobogatov, 1997a; Prozorova, 2003; Starobogatov 

et al., 2004). 

Anisus zhirmunskii Prozorova, 2003 F 

Prozorova, 2003: 103-104, fig. 

● Type locality: Lake Bolshoi Pelis, Island Bolshoi Pelis, Peter the Great Bay [Japan Sea]. 


● Distribution: southeastern Primorje (basins of rivers Tumannaya and Gladkaya, southern islands of the Peter 

the Great Bay) (Prozorova, 2003; Starobogatov et al., 2004). 

Armiger Hartmann, 1840 

Type species: Nautilus crista (Linnaeus, 1758) (OD) 

Armiger annandalei (Germain, 1918) F 

Planorbis (Armiger) annandalei Germain, 1918: 282-283, pl. 5, figs. 2-4. 

● Type locality: "Yarkand". 


● Distribution: mountain areas of central Asia, in Russia in lake Tere-Hol (Tuva) (Starobogatov et al., 2004). 

Armiger bielzi (Kimakowicz, 1884) F 

[= Armiger vitoldianus B. Dybowski et Grochmalicki, 1922; A. paravitoldianus B. Dybowski et Grochmalicki, 

1922; Planorbis crista var. inermis Lindholm, 1926, part.] 

Planorbis crista var. bielzi Kimakowicz, 1884: 79. 

● Type locality: "Hammersdorf". 


● Distribution: entire Europe, southern part of Western Siberia, Lena, Yana, Indigirka, and Kolyma rivers 

basins, in still and running waterbodies (Dolgin, 2001; Starobogatov et al., 2004; Karimov, 2005).

Armiger crista (Linnaeus, 1758) F 

[= Turbo nautileus Linnaeus, 1767; Planorbis imbricatus Müller, 1774: 165; Planorbis cristatus Draparnaud, 

1805; Planorbis crista var. spinulosus Clessin, 1873; Armiger jagellonensis B. Dybowski et Grochmalicki, 

1922; A. sobieskianus B. Dybowski et Grochmalicki, 1922; A. gedyminensis B. Dybowski et Grochmalicki, 

1922; Planorbis crista var. inermis Lindholm, 1926, part.] 

Nautilus crista Linnaeus, 1758: 709. 

● Type locality: “in Germaniae”. 

● Types: unknown? 

● Distribution: Europe, western and eastern Siberia, Baikal, Central Asia, Ural River (Akramowski, 1976; 

Pirogov et al., 1994; Starobogatov et al., 2004; Sitnikova et al., 2004). 

Armiger eurasiaticus Prozorova et Starobogatov, 1996 F 

Prozorova, Starobogatov, 1996: 172, fig. 3D. 

● Type locality: Elva, Estonia. 


● Distribution: Europe, northern Asia, in permanent waterbodies (Starobogatov et al., 2004). 

Armiger jankovskajae Izzatullaev, 1984 F 

Izzatullaev, 1984a: ??? 



● Distribution: eastern Pamir (Starobogatov et al., 2004). 

Armiger khoresmicus Prozorova et Starobogatov, 1996 F 

Prozorova, Starobogatov, 1996: 174, fig. 4D. 

Type locallity: Lake Korp-Kul’, Khiva district of Khoresm Region of Uzbekistan. 


● Distribution: lower reaches of Amu-Darya (Starobogatov et al., 2004). 

Armiger shamaricus Izzatullaev, 1972 F 

Izzatullaev, 1972: 47-48, fig. 1 (10 a-б). 

● Type locality: Shamar Lake, Tajik SSR, Tajikabad district, 2600 m above sea level. 


Distibution: type locality (Izzatullaev, 1972). 

Choanomphalus Gerstfeldt, 1859 

Type species: Choanomphalus maacki Gerstfeldt, 1859 (monotypy) 

Choanomphalus amauronius amauronius Bourguignat, 1860 F 

Choanomphalus amauronius Bourguignat, 1860c: 529, pl. 23, figs. 6-10. 

● Type locality: "dans la rivière d'Angara ainsi que dans le lac Baпkal, en Sibérie".. 


● Distribution: southern Baikal, Angara, 1-20 m (Sitnikova et al., 2004). 

Choanomphalus amauronius angulatus (B. Dybowski et Grochmalicki, 1925) F 

Valvatomphalus angulatus B. Dybowski, Grochmalicki, 1925: 878, figs. 29-31. 


● Types: unknown, possibly in the museum of Lvov University (Sitnikova et al., 2004). 

● Distribution: Baikal, Angara in vicinities of Irkutsk, 1.5-20 m (Sitnikova et al., 2004). 

Choanomphalus amauronius westerlundianus Lindholm, 1909 F 

Choanomphalus (Achoanomphalis) amauronius westerlundianus Lindholm, 1909: 20, pl. 2, fig. 1, 3. 

● Type locality: Kultuk (SW coast of southern Baikal).


● Distribution: SE coast of Baikal, 2-40 m (Kozhov, 1936; Sitnikova et al., 2004). 

Choanomphalus annuliformis Kozhov, 1936 F 

Choanomphalus (Sulcifer) annuliformis Kozhov, 1936: 182, pl. 6, figs. 42, 43, 59, 66-68, 68a. 

● Type locality: Ongureny (W coast of Baikal, northward to Bolshuye Olhonskiye Vorota). 


● Distribution: northern Baikal, Maloe More, 1.5-10 m (Sitnikova et al., 2004). 

Choanomphalus anomphalus W. Dybowski, 1901 F 

[= Choanomphalus (Achoanomphalus) dybowskianus Lindholm, 1909] 

W. Dybowski, 1901a: 123. 

● Type locality: Angara. 

● Types: possibly in the museum of Lvov University (Sitnikova et al., 2004). 


Choanomphalus aorus Bourguignat, 1860 F 

Bourquignat, 1860c: 530, pl. 23, figs. 11-15. 

● Type locality: "en Sibérie, dans le lac Baпkal". 


● Distribution: Baikal, Angara River downstream to Bratsk, 1.5-20 m (Sitnikova et al., 2004; Starobogatov et 

al., 2004). 

Choanomphalus bathybius bathybius Beckman et Starobogatov, 1975 F 

Choanomphalus (Kozhovisulcifer) bathybius bathybius Beckman, Starobogatov, 1975: 104, fig. 4A. 

● Type locality: near Sosnovka Bay (northern Baikal), depth 400 m. 



Choanomphalus bathybius meridianus Beckman et Starobogatov, 1975 F 

Choanomphalus (Kozhovisulcifer) bathybius meridianus Beckman, Starobogatov, 1975: 105, рèC. 4B. 

● Type locality: near Sljudjanka city (southern Baikal), depth 170 m. 



Choanomphalus cryptomphalus W. Dybowski, 1901 F 

W. Dybowsky, 1901a: 123. 

● Type locality: Angara. 

● Syntypes: “Museum Lemberg.” 

● Distribution: Balkal, Angara River downstream to Irkutsk (Starobogatov et al., 2004). 

Choanomphalus eurystomus Lindholm, 1909 F 

[= Choanomphalus (Choanomphalus) amauronius f. euristomus Kozhov, 1936] 

Choanomphalus (Achoanomphalus) eurystomus Lindholm, 1909: 16, pl. 11, figs. 22-24. 

● Type locality: Solzan (E coast of southern Baikal). 



Choanomphalus gerstfeldtianus Lindholm, 1909 F 

[= Choanomphalus (Choanomphalus) gerstfeldtianus var. striatus Kozhov, 1936] 

Choanomphalus (Achoanomphalus) gerstfeldtianus Lindholm, 1909: 20, pl. 2, figs. 15-17. 

● Type locality: Baranchiki and Solzan (southern Baikal). 

● Syntypes: ZIN.


Choanomphalus grachevi Sitnikova et Röpstorf, 1999 F 

Choanomphalus (Baicaloplanorbis) grachevi Sitnikova, Röpstorf, 1999: 123, 124, 126, figs. 2-5. 

● Type locality: Talanka Bay (W coast of central Baikal). 


● Distribution: W coast of central Baikal, 10-12 m (Sitnikova et al., 2004). 

Choanomphalus huzhirensis Beckman et Starobogatov, 1975 F 

Choanomphalus (Kozhovisulcifer) huzhirensis Beckman et Starobogatov, 1975: 104-105, fig. 4G. 

● Type locality: Maloe More, against Khuzhiry settlement, 65 m (Baikal Lake). 


● Distribution: Baikal: type locality and Malye Olkhonskiye Vorota, 65-100 m (Sitnikova et al., 2004). 

Choanomphalus hyaliniiformis Moskvicheva in Dworiadkin, 1980 F 

Choanomphalus hyaliniiformis Moskvicheva – Dworiadkin, 1980: 31, fig. 7. 

● Type locality: Khanka Lake, 1 km southward from village Astrakhanka. 


● Distribution: central and lower Amur basin, Primorje (Razdolnaya River basin, small waterbodies near 

lakes Khanka, Khasan and Doritzine), in small permanent and semipermanent waterbodies (Zatrawkin, 


Choanomphalus incertus incertus Lindholm, 1909 F 

Lindholm, 1909: 121, pl. 2, figs. 31-33. 

● Type locality: Guba Ayaya, Besymyannaya Buchta, Insel Bogutschan, Bogutschanskaya Buchta. 


● Distribution: Baikal (Starobogatov et al., 2004). 

Choanomphalus incertus mesospiralis B. Dybowski et Grochmalicki, 1925 F 

Choanomphalus mesospiralis B. Dybowski et Grochmalicki, 1925: 878, figs. 4, 5, 11, 14. 



● Distribution: southern Baikal, Angara River downstream to Irkutsk (Starobogatov et al., 2004). 

Choanomphalus korotnevi Lindholm, 1909 F 

[= Choanomphalus planospiralis B. Dybowski et Grochmalicki, 1925] 




● Distribution: Baikal: Maloe More and SW coast of northern part, 1-15 m (Sitnikova et al., 2004). 

Choanomphalus kozhovi Beckman et Starobogatov, 1975 F 

Choanomphalus (Baicaloplanorbis) kozhovi Beckman, Starobogatov, 1975: 106, fig. 4E. 

● Type locality: near Bolshaya Kosa Cape (NW Baikal). 


● Distribution: type locality and W coast of northern Baikal, 12-70 m (Sitnikova et al., 2004). 

Choanomphalus lindholmi Beckman et Starobogatov, 1975 F 

Choanomphalus (Kozhovisulcifer) lindholmi Beckman, Starobogatov, 1975: 105, fig. 4V. 

● Type locality: near Zavorotnyj Cape (northern Baikal). 


● Distribution: type locality, W coast of northern Baikal, 12-70 m (Sitnikova et al., 2004).

Choanomphalus maacki maacki Gerstfeldt, 1859 F 

[= Choanomphalus bicarinatus W. Dybowski, 1901; Choanomphalus mesospiralis B. Dybowski et Grochmalicki, 

1925] 

Choanomphalus maacki Gerstfeldt, 1859: 24, fig. 31. 



● Distribution: southerrn Baikal, Angara River, 1.5-20 m (Sitnikova et al., 2004). 

Choanomphalus maacki andrussowianus (Lindholm, 1909) F 

[= Choanomphalus elatospiralis B. Dybowski et Grochmalicki, 1925, Choanomphalus andrussovianus B. 

Dybowski et Grochmalicki, 1925] 

Choanomphalus andrussowianus Lindholm, 1909: 13, pl. 2, figs. 28-30. 

● Type locality: Tolstyi Cape, southward to headwaters of Angara (W coast of southern Baikal). 


● Distribution: W coast of southern Baikal, 0-30 m (Sitnikova et al., 2004). 

Choanomphalus maacki elatior Lindholm, 1909 F 

Choanomphalus korotnevi elatior Lindholm, 1909: 13, pl. 2, figs. 25-27. 


● Possible syntype: ZIN. 

● Distribution: Baikal: Maloe More and northern part (Kozhov, 1936; Sitnikova et al., 2004). 

Choanomphalus microtrochus Lindholm, 1909 F 

Choanomphalus (Achoanomphalus) microtrochus Lindholm, 1909: 22, pl. 2, figs. 10-11. 

● Type locality: Ayaya and Dagarskaya bays (E coast of northern Baikal). 


● Distribution: E coast of northern Baikal, 10–12 m (Sitnikova et al., 2004). 

Choanomphalus okhoticus Prozorova et Starobogatov, 1997 F 

Prozorova, Starobogatov, 1997a: 43-44, fig. 1G. 

● Type locality: Tugur River valley. 


● Distribution: type locality (Prozorova, Starobogatov, 1997a; Starobogatov et al., 2004). 

Choanomphalus omphalotus W. Dybowski, 1901 F 

W. Dybowski, 1901a: 122 

● Type locality: Angara River. 

● Syntypes: probably in Lvov University. 

● Distribution: Angara, ? northern Baikal, 2-15 m (Sitnikova et al., 2004). 

Choanomphalus parvus Kozhov, 1936 F 

Choanomphalus (Choanomphalus) dybowskianus var. parvus Kozhov, 1936: 178, pl. 4, figs. 35-36. 

● Type locality: northern Baikal. 


● Distribution: central and northern Baikal, Maloe, 5–25 m (Sitnikova et al., 2004). 

Choanomphalus patulaeformis Lindholm, 1909 F 


● Type locality: near ulus Kurma in Maloe More [Baikal]. 


● Distribution: Baikal, 2-30 m (Sitnikova et al., 2004).

Choanomphalus planorbiformis Beckman et Starobogatov, 1975 F 

Choanomphalus (Antichoanomphalus) planorbiformis Beckman et Starobogatov, 1975: 107; fig. 4 Zh. 

● Type locality: Sosnovka Bay (northern Baikal) depth 400 m. 


● Distribution: type locality, central Baikal, 120-580 m (Sitnikova et al., 2004). 

Choanomphalus pygmaeus Lindholm, 1909 F 

Choanomphalus (Achoanomphalus) pygmaeus Lindholm, 1909: 22, pl. 2, figs. 4-6. 



● Distribution: type locality and Maloe More, 5-25 m (Sitnikova et al., 2004). 

Choanomphalus riparius (Westerlund, 1865) F 

Planorbis riparius Westerlund, 1865: 106. 

● Type locality: “Förekommer fästad pa trästycken, stenar o. s. v. pa östra stranden af Härrstorps sjö vif Ronneby”. 


● Distribution: Europe, except extreme north, southern Ural, plain Altai and lower Ob’ basin, in lakes (Starobogatov 

et al., 2004; Dolgin, personal communication). 

● Remark: FROM the C. Meier-Brook’s opinion (1983), the species Planorbis riparius should be classified 

within the genus Gyraulus. 

Choanomphalus rossmaessleri (Auerswald in A. Schmidt, 1851) F 

[=Planorbis (Gyraulus) planoconcavus Westerlund, 1897] 

Planorbis rossmaessleri Auerswald – Schmidt, 1851: 179. 

● Type locality: "prope Lipsias" [Leipzig]. 


● Distribution: Europe, except north, Altai, Siberia, eastward to left bank of Lena river, in temporary waterbodies 

(Starobogatov et al., 2004; Vinarski et al., 2006). 

● Remark: examination of the type series of Planorbis (Gyraulus) planoconcavus did not reveal significant 

differences from C. rossmaessleri and therefore it is synonimised with the latter (Vinarski et al., 2006). From 

the C. Meier-Brook’s (1983) opinion, the species Planorbis rossmaessleri should be classified within the genus 

Gyraulus 

Choanomphalus schrenkii W. Dybowski, 1875 F 

W. Dybowski, 1875: 60, pl. 2, figs. 27-33. 

● Type locality: Kultuk (SW coast of southern Baikal) [restr. Sitnikova et al., 2004]. 



Choanomphalus valvatoides W. Dybowski, 1875 F 

[= Chоanomphalus. intermedius W. Dybowski, 1901] 

W. Dybowski, 1875: 58, pl. 2, fig. 19, 26; pl. 6, fig. 6. 




Helicorbis Benson, 1855 

Type species: Planorbis umbilicatus Benson, 1855 (OD) 

Helicorbis bactriana Izzatullaev, 1980 F 

Izzatullaev, 1980: 408-409, fig. 2, А, а, б, в. 

● Type locality: Turkmen SSR, Kerkichi, right bank of Amu-Darja.


● Distribution: mountain areas of Central Asia (Starobogatov et al., 2004). 

Helicorbis kozhovi Starobogatov et Streletzkaja, 1967 F 

Starobogatov, Streletzkaja, 1967: 239-240, fig. 36. 

● Type locality: Kotokel Lake, Mostovaja inlet (Baikal Lake area). 

● Holotype: ZIN, No. 1 in systematic catalogue 

● Distribution: Transbaikalia: Kotokel Lake, valley of Manzurka River (upper Lena basin), Yana, Indigirka 

and Kolyma rivers basins, waterbodies of the left bank of Lena, in permanent and semipermanent waterbodies 

(Starobogatov, Streletzkaja, 1967; Dolgin, 2001; Starobogatov et al., 2004; M.V. Vinarski, unpublished). 

Helicorbis kushanica Izzatullaev, 1980 F 

Izzatullaev, 1980: 410, fig. 2, Б, а, б, в. 

● Type locality: Turkmen SSR, Kerkichi, right bank of Amu-Darja. 



Helicorbis levanidovae Zatrawkin et Moskvicheva in Zatrawkin, 1985 F 

Zatrawkin, 1985b: 41, fig. 2. 

● Type locality: Amur within borders of Khabarovsk. 


● Distribution: type locality, possibly upper Sungari basin (Starobogatov et al., 2004). 

Helicorbis rara Moskvicheva in Zatrawkin, 1985 F 

Zatrawkin, 1985b: 40-41, fig. 3. 

● Type locality: Vertopryakha River (Amur basin). 


● Distribution: Amur River shore at Sungari junction (Vertopryakha River mouth), possibly upper Sungari 

basin (Zatrawkin, 1985; Starobogatov et al., 2004). 

Helicorbis shilkaensis Starobogatov, 1996 F 

Starobogatov, 1996a: 1427-1428, textfig. (1). 

● Type locality: vicinities of Sretensk, Shilka River valley, Chita Region. 

● Types: ZIN, No. 1 in systematic catalogue. 

● Distribution: Shilka River and Khanka lake basin (southern Primorje) (Starobogatov et al., 2004). 

Helicorbis sujfunensis Starobogatov, 1957 F 

Starobogatov, 1957: 1004-1005, figs. zh-p. 

● Type locality: vicinities of Voroshilov city (now Ussurijsk City, Far East). 


● Distribution: Amur River basin, southern part of Primorje, in temporary waterbodies (Starobogatov et al., 

2004). 

Hippeutis Agassiz in Charpentier, 1837 

Type species: Helix complanatus Linnaeus, 1758 (SD Clessin, 1878) 

Hippeutis complanatus (Linnaeus, 1758) F 

[= Planorbis (Hippeutis) colchicus Lindholm, 1913] 

Helix complanatus Linnaeus, 1758: 769. 

● Type locality: "in Europae". 

● Types: whereabouts unknown?

● Distribution: Europe and Siberia, Altai, Central Asia (Shadin, 1952; Akramowski, 1976). 

● Remark: according to Starobogatov (1977) and subsequent Russian authors (Stadnichenko, 1990; Starobogatov 

et al., 2004), the species Helix complanatus should be splitted into the three distinct species: Hippeutis 

diaphanella, H. euphaea and Hi. fontana. The reasons of rejection of the species name Hippeutis complanatus 

that is accepted by Western malacologists (Falkner et al., 2001; Glöer, 2002), are not wholly clear. 

Hippeutis diaphanella (Bourguignat, 1864) F 

Planorbis diaphanellus Bourguignat, 1864a, 2: 167, pl. 9, figs. 39-42. 

● Type locality: small rivers of Mostaghanem (Brondel) [Algeria]. 


● Distribution: Europe, south of western Siberia, Ural River, in permanent waterbodies (Pirogov et al., 1994; 


Hippeutis euphaea (Bourguignat, 1864) F 

Planorbis euphaeus Bourguignat, 1864a: 165-167, pl. 9, figs. 35-38. 

● Type locality: "environs d'Alger, notamment а Hussein-Dey; – sur le bords de l'Harrach". 


● Distribution: southern Europe, south of Western Siberia, Kazakhstan, in permanent waterbodies (Starobogatov 

et al., 2004). 

Hippeutis fontana (Lightfoot, 1786) F 

Helix fontana Lightfoot, 1786: 165, pl. 11, fig. 2. 

● Type locality: spring near Bulllstrode, Buckinghamshire. 


● Distribution: Europe, in permanent waterbodies (Starobogatov et al., 2004). 

Kolhymorbis Starobogatov et Streletzkaja, 1967 

Type species: Kolhymorbis shadini Starobogatov et Streletzkaja, 1967 (OD) 

Kolhymorbis angarensis (B. Dybowski et Grochmalicki, 1925) F 

[= Kolhymorbis maacki Starobogatov et Streletzkaya, 1967] 



● Distribution: Eastern Siberia, Amur basin, Central and Easrern Siberia, from Khatanga to Kolyma River, in 

temporary and semipermanent waterbodies (Starobogatov et al., 2004; Dolgin, 2001). 

● This species in Russian literature is attributed to B. Dybowski et Grochmalicki (1925). Nevertheless it was 

not described in the cited work and the formal description was not traced by us. 

Kolhymorbis bogatovi Zatrawkin et Moskvicheva in Zatrawkin, 1985 F 

Zatrawkin, 1985b: 40, fig. 1. 

● Type locality: puddle in the upstream of river Manoma, tributary of Anui (lower Amur basin). 


● Distribution: central and lower Amur basin (Zatrawkin, 1985b; Starobogatov et al., 2004). 

Kolhymorbis dildorae Izzatullaev, 1980 F 

Kolhymorbis dildora Izzatullaev, 1980: 408, fig. 1, В, а, б, в (lapsus calami). 

● Type locality: Uzbek SSR, Tashkent Region, Akkurganskoye water reservoir, Balykchi. 



● Remark: although the name of the species in the original description was spelled as dildora, this is obvious 

lapsus calami, since the species was named after author’s daughter. Besides in the other parts of 

the text the name was spelled as dildorae.

Kolhymorbis shadini Starobogatov et Streletzkaja, 1967 F 


● Type locality: water bodies of the Kolyma delta near Pokhodsk. 

● Holotype: ZIN, No. 1 in systematicv catalogue. 

● Distribution: lower reaches of Siberian rivers from Khatanga in the west to basin of Kolyma in the east, 

Chukotka, in shallow permanent waterbodies (Dolgin. 2001; Starobogatov et al., 2004). 

Planorbis Geoffroy, 1767 

Type species: Helix planorbis Linnaeus, 1758 (tautonomy) 

Planorbis carinatus Müller, 1774 F 

Müller, 1774: 157-158. 



● Distribution: northwest of Russia, south of European part of Russia, south of Ukraine, Ural River, Altai, 

southern Ural, Armenia (Sevan Lake) (Zatrawkin, 1980; Pirogov et al., 1994; Soldatenko, Starobogatov, 


Planorbis dubius Hartmann, 1821 F 

Planorbis marginatus β dubius Hartmann, 1821: 254-255. 

● Type locality: canals of Rhein and in Katzensee in Zurich Kanton. 

● Types: Naturmuseum Sent-Gallen (Switzerland), No. M299. 

● Distribution: Ciscaucasia, Baltic Sea coast, in permanent waterbodies (Starobogatov et al., 2004). 

Planorbis intermixtus Mousson, 1874 F 

[= Planorbis persicus Ancey, 1900; Planorbis sieversi sensu Likharev, Starobogatov, 1967 (part.) et Starobogatov, 

1970, non Mousson, 1874] 

Planorbis (Gyraulus) intermixtus Mousson, 1874: 45. 

● Type locality: "La Basse-Mésopotamie". 


● Distribution: lowland of Central Asia, eastern Transcaucasia, Daghestan, western part of Volga Delta 

(Soldatenko, Starobogatov, 2000). 

Planorbis kubanicus Soldatenko et Starobogatov, 1998 F 

[= Planorbis carinatus var. dilatatus Clessin 1873 non Planorbis dilatatus Gould, 1841] 

Soldatenko, Starobogatov, 1998: 59-60, 62, figs. 1A, 2. 

● Type locality: Russia, Krasnodar Region. Kuban River, Akhtansk group of limans. 


● Distribution: southern Europe, including Ciscaucasia (Soldatenko, Starobogatov, 1996; Starobogatov et al., 

2004). 

Planorbis philippianus Monterosato in Caziot, 1902 F 

[= Planorbis umbilicatus var. subangulatus Philippi, 1844, non Planorbis subangulatus Lamarck, 1807; 

Planorbis philippi Germain, 1908] 

Monterosato – Caziot, 1902: 262 (nom. nov. pro Planorbis subangulatus Philippi, 1844, non Linnaeus, 

1758). 

● Type locality: "In rivulis prope Panormum frequens (ad Nicaem Liguriae legit cl. Kunze)" (Philippi, 1844: 

119). 


● Distribution: Ukraine, Ciscaucasia and Transcaucasia (Soldatenko, Starobogatov, 1998; Starobogatov et al., 

2004).

Planorbis planorbis (Linnaeus, 1758) F 

[= Planorbis marginatus Draparnaud, 1805; 

Helix planorbis Linnaeus, 1758: 769. 



● Distribution: Europe, southern West Siberia to Altai, Baikal Lake, Tajikistan (lake Shamar) (Izzatullaev, 

1972; Starobogatov et al., 2004; Sitnikova et al., 2004). 

Planorbis sieversi Mousson, 1873 F 

Mousson, 1873: 29-30, pl. 7, fig. 9. 

● Type locality: "Daralagher". 

● Syntype: ZMZ No. 521151. 

● Distribution: Central Asia, eastern Transcaucasia, Daghestan, Volga delta, in permanent and semipermanent 

waterbodies (Akramowski, 1976; Starobogatov et al., 2004). 

Planorbis tangitarensis Germain, 1918 F 

[= Planorbis sieversi sensu Likharev, Starobogatov, 1967 (part.), non Mousson, 1873] 

Planorbis planorbis var. tangitarensis Germain, 1918: 276. 

● Type locality: "Tangitar, dans la Kachgarie (Turkestan Oriental)". 


● Distribution: mountain regions of Central Asia, possibly in Altai and Tuva (Izzatullaev, 1972; Starobogatov 

et al., 2004). 

Planorbis umbilicatus Müller, 1774 F 

[= Planorbis marginatus Draparnaud, 1805] 

Müller, 1774: 160-161. 

● Type locality: Fridrichsdal (Denmak). 


● Distribution: entire Europe, Cisbaikalia, in permanent waterbodies (Soldatenko, Starobogatov, 2000; Starobogatov 

et al., 2004). 

Polypylis Pilsbry, 1906 

Type species: Bulimus largillierti Philippi, 1845 (monotypy) 

Polypylis almaatina Starobogatov et Mamilova, 1970 F 

Starobogatov, Mamilova, 1970: 62-63, fig. 1,2. 

● Type locality: eastward from Alma-Ata (Kazakh SSR), spring at the territory of collective farm "Zarja 

Vostoka". 


● Distribution: Kazakhstan, Tajikistan (Izzatullaev, 1972). 

Polypylis likharevi Starobogatov et Streletzkaja, 1967 F 

Starobogatov, Streletzkaja, 1967: 238-239, fig. 35. 

● Type locality: lake on the former river bed in flood-lands of the River Malyj AnjuJ [NE Siberia]. 


● Distribution: Kolyma River basin, Kamchatka, in small permanent waterbodies (Starobogatov, Streletzkaja, 


Polypylis nitidella (Martens, 1877) F 

[= Polypylis (Segmentina) mica Westerlund, 1883] 

Planorbis nitidellus Martens, 1877: 112. 

● Type locality: Yokohama. 

● Types: not traced, possibly in Berlin Museum (fide Dance, 1986).

● Distribution: South Kurile Islands (Prozorova, 1996 [1997]a; Starobogatov et al., 2004). 

Polypylis semiglobosa Moskvicheva in Dworiadkin, 1980 F 

Polypylis semiglobosa Moskvicheva – Dworiadkin, 1980: 28-30, fig. 5. 

● Type locality: Khankaiski District, Astrakhanka village, a channel (information from the holotype label). 


● Distribution: Amur basin, Primorje, in small permanent waterbodies (Starobogatov et al., 2004). 

Polypylis sibirica Starobogatov et Streletzkaja, 1967 F 


● Type locality: lake on the Oblom Cape (eastern coast of the Baikal Lake). 


● Distribution: Cisbaikalia, probably in south of Central Siberia, Kolyma River basin, in lakes (Starobogatov, 

Streletzkaja, 1967; Dolgin, 2001; Starobogatov et al., 2004). 

Polypylis starobogatovi Izzatullaev, 1973 F 

Izzatullaev, 1973a: 65-66, fig. А (1, 2, 3). 

● Type locality: Tajik SSR, natural reserve “Tigrovaya balka”, central cordon, deposits of the Blue Lake. 


● Distribution: type locality (Izzatullaev, 1973a). 

Promenetus Baker, 1935 

Type species: Planorbis exacuous Say, 1821 (OD) 

Promenetus exacuous (Say, 1821) F 

Planorbis exacuous Say, 1821: 165. 

● Type locality: Lake Champlain. 

● Syntypes: ANSP. 

● Distribution: found in stomach of fish caught in central part of Anadyr River (Chukotka) (Starobogatov et 

al., 2004). 

Segmentina Fleming, 1818 

Type species: Nautilus lacustris Lightfoot, 1786 = Planorbis nitidus O.F. Müller, 1774 (monotypy) 

Segmentina avecenninae Izzatullaev, 1980 F 

Izzatullaev, 1980: 406, fig. 1, А, Б, а, б, в. 

● Type locality: Turkmen SSR, Kerkichi, right bank of Amu-Darja. 



Segmentina caucasica Kruglov et Soldatenko, 1997 F 

Kruglov, Soldatenko, 1997: 126, figs. 4D, 7D, 11B. 

● Type locality: northern Caucasus, Northern Osetia Republic, road to Alagir, puddle. 


● Distribution: northern Caucasus, in temporary waterbodies (Kruglov, Soldatenko, 1997). 

Segmentina clessini (Westerlund, 1873) F 

Planorbis clessini Westerlund, 1873: 613-614. 

● Type locality: “Svewrige pе Öland i en back söder om byn Kohlstad nära Borgholm”. 


● Distribution: Europe, except for extreme north-east, northern Kazakhstan, Altai, probably south of Western 

Siberia (Kruglov, Soldatenko, 1997).

Segmentina distinguenda (Gredler, 1859) F 

Planorbis nitidus var. distinguenda Gredler, 1859: 217. 

● Type locality: "Breslau". 

● Types: not traced. According to Dance’s (1986) information, the Gredler’s type materials are housed partly 

in Fernandeum at Innsbruck, partly at the Bozen Gymnasium (Austria). 

● Distribution: Europe, except for extreme north, northern Kazakhstan, Altai, south of Western Siberia, in 

shallow permanent waterbodies, south of Central Asia (Izzatullaev, 1980; Kruglov, Soldatenko, 1997; Starobogatov 

et al., 2004). 

Segmentina inniae Kruglov et Soldatenko, 1997 F 

Kruglov, Soldatenko, 1997: 117-118, fig. 3C. 

● Type locality: Latviya, Skrundskyi District, fishery farm Rita Ausma, pond 33. 


● Distribution: Baltic Sea basin, south of Western Siberia (within the Irtysh River basin) (Starobogatov et al., 

2004; Karimov, 2005). 

Segmentina lineata Kruglov et Soldatenko, 1997 F 

Kruglov, Soldatenko, 1997: 121-122, figs. 3F, 6D, 10A. 

● Type locality: Smolensk Region, Dorogobyzhskyi Area, village Ushakovo, pond. 


● Distribution: Latvia, European Russia, in shallow permanent waterbodies (Kruglov, Soldatenko, 1997). 

Segmentina malkae Kruglov et Soldatenko, 1997 F 

Kruglov, Soldatenko, 1997: 126-128, figs. 4E, 8A, 11C. 

● Type locality: northern Caucasus, Kabardino-Balkaria, Malka River near village Ekaterinogradskaya. 


● Distribution: northern Caucasus, temporary waterbodies (Kruglov, Soldatenko, 1997). 

Segmentina molytes (Westerlund, 1885) F 

Planorbis nitidus var. molytes Westerlund, 1885a: 86. 

● Type locality: "Deutschland bei Heidelberg u. Vegesack". 


● Distribution: Europe, except extreme north, northern Kazakhstan, probably Altai (Kruglov, Soldatenko, 


Segmentina montgazoniana Bourguignat in Servain, 1881 F 

Servain, 1881: 89-90. 

● Type locality: "dans les mares des villages de la Villenueve, la Ville-au-Bois, la Loge-aux-Chèvres, etc. 

(Département de l'Aube)"(France). 


● Distribution: Europe, except south, in shallow permanent waterbodies (Kruglov, Soldatenko, 1997; Starobogatov 

et al., 2004). 

Segmentina nitida (Müller, 1774) F 

[= Planorbis microcephalus Moquin-Tandon, 1855] 

Planorbis nitidus Müller, 1774: 163-164. 



● Distribution: Europe, except for extreme north-east, Ural River (Pirogov et al., 1994; Starobogatov et al., 

2004). 

Segmentina oelandica (Westerlund, 1885) F 

Planorbis nitidus var. oelandicus Westerlund, 1885a: 86.

● Type locality: "Schweden, Borgholm". 

● Types: not traced, possibly in NHG. 

● Distribution: Ukraine, European Russia except for extreme north, south of Western Siberia (within the 

Irtysh River basin), in shallow permanent waterbodies (Kruglov, Soldatenko, 1997; Starobogatov et al., 2004; 

Karimov, 2005). 

Segmentina petropolitana Kruglov et Soldatenko, 1997 F 

Kruglov, Soldatenko, 1997: 124-126, figs. 4C, 7C, 11A. 

● Type locality: Ukraine, vicinities of Vinnitsa. 


● Distribution: Europe, except for extreme north-east, in shallow permanent waterbodies (Kruglov, 

Soldatenko, 1997). 

Segmentina servaini Bourguignat in Servain, 1881 F 

Servain, 1881: 86-87. 

● Type locality: "lac Balaton, ... environs de Krapina (Croatie)". 


● Distribution: Europe (except for north-east), south of Western Siberia (within the Irtysh River basin), in 

shallow permanent waterbodies (Kruglov, Soldatenko, 1997; Starobogatov et al., 2004; Karimov, 2005). 

Segmentina starobogatovi Kruglov et Soldatenko, 1997 F 

Kruglov, Soldatenko, 1997: 122-124, figs. 4B, 7B, 10C. 

● Type locality: Kalinigrad District, Rybachyi settlement, puddle on a meadow. 


● Distribution: Baltic Sea basin, central Volga basin, in shallow permanent and semipermanent waterbodies 

(Kruglov, Soldatenko, 1997). 

Trochorbis Benson, 1855 

Type species: Trochorbis trochoideus Benson, 1855 (OD). 

Trochorbis anastasiae Izzatullaev, 1973 F 

Izzatullaev, 1973a: 67-68, fig. В (1, 2, 3). 



● Distribution: Tajikistan, Vakh River (Izzatullaev, 1973a; Starobogatov et al., 2004). 

Trochorbis calathus (Benson, 1850) F 

Planorbis calathus Benson, 1850: 348-349. 

● Type locality: “in lacubus Bhimtal et Neini Tal, regionis Kemaonensis Himalayanae; necnon in stagnis 

nonnulis agri Rohilllani, prope urbem Morahabad, Indiae septentrionalis”. 


● Distribution: Vaksh River, near natural reserve “Tigrovaya balka”, Tajikistan (Izzatullaev, 1972). 

Trochorbis tadzhikistanica Izzatullaev, 1973 F 

Trochorbis tadzhikistanical [lapsus calami] Izzatullaev, 1973a: 66-67, fig. Б (1, 2, 3). 



● Distribution: Tajikistan, Vakh River (Izzatullaev, 1973a; Starobogatov et al., 2004). 

PHYSIDAE Fitzinger, 1833 

(Kantor Yu. I., Vinarki M.V., Sysoev A.V.)

APLEXINAE Starobogatov, 1967 

Aplexa Fleming, 1820 

Type species: Bulla hypnorum Linnaeus, 1758 (SD Herrmannsen, 1846) 

Aplexa amurensis Starobogatov et Prozorova in Starobogatov, Prozorova et Zatrawkin, 1989 F 

Starobogatov, Prozorova, Zatrawkin, 1989: 65, fig. 1 4, 2 4. 

● Type locality: Khabarovsk Territory [Far East], swamp near settlement Konstantinovka. 

● Holotype: ZIN, No 1. in systematic catalogue. 

● Distribution: Amur basin and Primorje, Balkal and upper Yenissei (northeastern Tuva) basins, in temporary 


Aplexa aphallica Starobogatov et Zatrawkin in Starobogatov, Prozorova et Zatrawkin, 1989 F 

Starobogatov, Prozorova, Zatrawkin, 1989: 66, fig. 1 7. 

● Type locality: :Primorje Region [Far East], Khanka District, a channel in the rice field in vicinity of Vladimiro-Petrovka 

village. 


● Distribution: Khanka lowland, probably Amur basin, in temporary waterbodies (Starobogatov et al., 2004). 

● Remark: Taylor (2003) considered this species to be a synonym of Aplexa amurensis. 

Aplexa ataxiaca (Fagot, 1883) F 

Physa ataxiaca Fagot, 1883: 222-223. 

● Type locality: "Campagne-les-Bains, entre Quillan et Limoux (Aude)" (France). 


● Distribution: southern part of eastern Europe, south of Western Siberia, Kazakhstan, in temporary waterbodies 


● Remark: Taylor (2003) considered this species to be a synonym of Paraplexa cornea (Massot, 1845). 

Aplexa hypnorum (Linnaeus, 1758) F 

Bulla hypnorum Linnaeus, 1758: 727-728. 

● Type locality: "in Europae". 

● Types: not traced, not in LSL (Taylor, 2003). 

● Distribution: Europe. south of Western Siberia, Kazakhstan, eastward to Altai, in temporary waterbodies 


Aplexa moskvichevae Starobogatov et Zatrawkin in Starobogatov, Prozorova et Zatrawkin, 1989 F 

Starobogatov, Prozorova, Zatrawkin, 1989: 66, fig. 1 (6), 2 (6). 

● Type locality: Artemovka River, [Khabarovsk Territory, Far East]. 


● Distribution: Primorje, probably Amur basin, in temporary waterbodies (Starobogatov et al., 2004). 


Aplexa orientalis Starobogatov et Prozorova in Starobogatov, Prozorova et Zatrawkin, 1989 F 


● Type locality: Khabarovsk Territory [Far East], swamp near settlement Konstantinovka. 


● Distribution: Amur basin, Primorje, in temporary waterbodies (Starobogatov et al., 2004). 


Aplexa turrita (Müller, 1774) F 

Planorbis turritus Müller, 1774: 169.


● Holotype: ZMUC. 

● Distribution: Europe, in temporary waterbodies (Starobogatov et al., 2004). 

● Remark: Taylor (2003) considered this species to be a synonym of Aplexa hypnorum. 

Sibirenauta Starobogatov et Streletzkaja, 1967 

Type species: Physa kultukiana B. Dybowski, 1913 (OD) 

Sibirenauta aenigma (Westerlund, 1876) F 

Physa (?) aenigma Westerlund, 1876a: 101, 104-105. 

● Type locality: "Podkamennojes Tunguskas in flode i Jenissei". 

● Type: SMNH No. 1.652. 

● Distribution: south of Central and Eastern Siberia (northward to Yakutsk and mouth of Podkamennaya 

Tunguska River), Altai, in temporary waterbodies (Starobogatov et al., 2004). 

● Remark: According to Taylor (2003), the type specimen of S. aenigma does not belong to Physidae and 

most probably represents a sinistral specimen of Cochlicopa lubrica (terrestrial snail). Thus, the use of the 

Westerlund’s binomen for designation of a species belonging to the family Physidae is problematic. 

Sibirenauta elongata (Say, 1821) F 

[= Physa hypnorum var. depressior Middendorff, 1851; Physa hypnorum var. polaris Westerlund, 1876; 

Physa kultukiana B. Dybowski, 1913] 

Physa elongata Say, 1821: 171. 

● Type locality: shores of Illinois. 

● Syntypes: ANSP No. 21208. 

● Distribution: from north or Western Siberia to Chukotka, upper Amur basin (Starobogatov, Streletzkaya, 


● Remark: according to the Taylor’s (2003) opinion, S. elongata is a strictly North American species and it 

does not occur in Siberia. Taylor records for Siberia two another species of Sibirenauta, namely, S. pictus 

(Krause, 1883) from the Chukotka Peninsula, and S. (?) depressior (Middendorff, 1851) found in the Taimyr 

waterbodies. S. pictus has previously been mentioned by Starobogatov & Budnikova (1976). However, later 

on Starobogatov et al. (1989) regarded S. pictus as a junior synonym of S. sibirica. 

Sibirenauta sibirica (Westerlund, 1876) F 

Physa sibirica Westerlund, 1876a: 55-56, fig. 13. 

● Type locality: "Jenissei, Sopotschnaja Korga (lat. 71º40')". 

● Syntypes: SMNH No. 1651, Westerlund collection 12:31. 

● Distribution: subarctic and arctic areas of Asia, from Yenissei mouth to Chukotka (Starobogatov et al., 

2004). 

Sibirenauta tuvaensis Starobogatov et Zatrawkin in Starobogatov, Prozorova et Zatrawkin, 1989 F 

Starobogatov, Prozorova, Zatrawkin, 1989: 68, figs. 1 9, 2 8. 

● Type locality: Tuva ASSR, Toorakhem settlement [at the border of Mongolia]. 

● Holotype: ZIN, No. 1 in systematic catalogue.. 

● Distribution: Tuva, in temporary waterbodies (Starobogatov et al., 2004). 

● Remark: Taylor (2003: 57) wrote: “This species does not belong to Sibirenauta as indicated by the enormous 

sarcobellum (W/L about .6) and the penis that widens in its distal half to more than twice the width of 

the proximal half, ending in a blutn rounded tip”. Thus, the proper systematic position of “Sibirenauta” tuwaensis 

needs to be revised. 

PHYSINAE Fitzinger, 1833

Costatella Dall, 1870 

Type species: Physa costata Newcomb, 1861 (OD) 

Costatella acuta (Draparnaud, 1805) F 

Physa acuta Draparnaud, 1805: 55. 

● Type locality: "dans la Garonne et les rivières qui s'y jettent". 

● Syntype: NHMV No. 1820/XXVI/45. 

● Distribution: Ciscaucasia, Transcaucasia, Central Asia, Volga River delta, southern Primorje (Izzatullaev, 


● Remark: in Western European literature (Falkner et al., 2001; Glöer, 2002) this species is referred to as 

Physella acuta. Taylor (2003) placed the species in the genus Haitia Clench et Aguayo, 1932. 

Costatella integra (Haldeman, 1841) F 

Physa (Diastropha) integra Haldeman, 1841: cover p. 3. 

● Type locality: Indiana. 


● Distribution: introduced from north America, Volga River delta, Ciscaucasia, southern Primorje, also in 

cooling basins of the electric plants (Starobogatov et al., 2004). 

● Remark: in Western European literature (Falkner et al., 2001; Glöer, 2002) this species is referred to as 

Physella integra. Taylor (2003) placed the species in the genus Haitia Clench et Aguayo, 1932. 

Physa Draparnaud, 1801 

Type species: Bulla fontinalis Linnaeus, 1758 (SD Children, 1823; ICZN Opinion 94, 1958) 

Physa adversa (Da Costa, 1778) F 

Turbo adversus Da Costa, 1778: 96-98, pl. 5, fig. 6. 

● Type locality: the Thames from Wandsworth to Windsor; ponds and stagnant waters about London; Middlesex; 

Surrey; river Wandle (England). 

● Types: unknown, most probably lost. 

● Distribution: entire Europe and Siberia eastward to Yenissei (Pirogov et al., 1994; Starobogatov et al., 

2004). 

● Remark: Taylor (2003) considered Turbo adversus to be a synonym of Physa fontinalis. 

Physa arachleica Starobogatov et Prozorova in Starobogatov, Prozorova et Zatrawkin, 1989 F 


● Type locality: Chita Region, Arakhlej Lake [SE Siberia]. 



Physa bulla (Müller, 1774) F 

Planorbis bulla Müller, 1774: 167-168. 


● Types: ZMUC (fide Taylor, 2003). 

● Distribution: western and southern part of eastern Europe, southern part of Western Siberia, Kazakhstan 


● Remark: Taylor (2003) considered Planorbis bulla to be a synonym of Physa fontinalis. 

Physa chukchensis Starobogatov et Budnikova, 1976 F 

Starobogatov, Budnikova, 1976: 84. fig. 5 I. 

● Type locality: Ust’-Chaun [Chukchi Peninsula]. 


● Distribution: northern Far East from Chukchi Peninsula to lower reaches of Kolyma, in thermokarst waterbodies 

(Starobogatov et al., 2004).

● Remark.: Taylor (2003) believed that Ph. chukchensis is a synonym of Beringiophysa jennessii (Dall, 

1919). 

Physa dalmatina Küster, 1844 F 

Küster, 1844 (1841-1850): 17, Taf. 2, figs. 17-19. [the date of publication of the page 17 is given according to 

Smith & England, 1937]. 

● Type locality: Dalmatia (Croatia). 

● Holotype: Presumably destroyed in SMF during World War II (fide Taylor, 2003). 

● Distribution: South Europe (Balkans) to Western Siberia (Taylor, 2003). Taylor & Sitnikova (2004) record 

this species from the Hövsgöl Lake, Mongolia. 

● Remark: Taylor (2003) mentioned Physa sartlandinensis Mozley, 1934 as a junior synonym of Ph. dalmatina. 

Starobogatov et al. (1989, 2004) do not include Ph. dalmatina in the list of freshwater snails of Russia 

and adjacent territories and do not discuss possible relationships between Ph. dalmatina and another species 

of the genus Physa recorded from the former USSR territory. 

Physa fontinalis (Linnaeus, 1758) F 

Bulla fontinalis Linnaeus, 1758: 727. 

● Type locality: "in Lacuum". 

● Types: not traced, not in LSL (Taylor, 2003). 

● Distribution: basin of Baltic Sea (Starobogatov et al., 2004), Western Siberia (Vinarski et al., 2007). 

● Remark: basing on Linnaeus (1746), Taylor (2003) accepted vicinity of Uppssala (Sweden) as a type locality 

of this species. 

Physa hankensis Starobogatov et Prozorova in Starobogatov, Prozorova et Zatrawkin, 1989 F 

Starobogatov, Prozorova, Zatrawkin, 1989: 72, fig. 3 5, 4 5,6 

● Type locality: Khabarovsk, swamp near Kaktokovskoye Lake [Far East]. 


● Distribution: Amur basin and southern Primorje (Starobogatov et al., 2004). 

Physa jarochnovitschae Starobogatov et Zatrawkin in Starobogatov, Prozorova et Zatrawkin, 1989 F 


● Type locality: delta of the Kolyma River (NE Siberia). 



● Remark: Taylor (2003) considered Ph. jarochnovitchae as a synonym of Ph. hankensis. 

Physa jennessii Dall, 1919 F 

Dall, 1919: 3A, 4A, 12A, 18A, 20A, pl. 2, fig. 2. 

● Type locality: ponds near Bernard harbour (Canada). 

● Types: NMC Nos. 4019, 4020. 

● Distribution: Chukchi Peninsula ans westward to lower Kolyma, in thermokarst waterbodies (Starobogatov 

et al., 2004). 

Physa khabarovskiensis Starobogatov et Zatrawkin in Starobogatov, Prozorova et Zatrawkin, 1989 F 

Starobogatov et al., 1989: 73, fig. 3 6. 

● Type locality: Khabarovsk, swamp near Kaktokovskoye Lake (Far East). 



● Remark: Taylor (2003) considered Ph. khabarovskiensis as a synonym of Ph. hankensis. 

Physa kuvaevi Starobogatov et Prozorova in Starobogatov, Prozorova et Zatrawkin, 1989 F 

Starobogatov et al., 1989: 74, fig. 3 8, 4 8. 

● Type locality: lake near airport Anadyr (Chukchi Peninsula).



● Remark: Taylor (2003) believed that Ph. kuvaevi is a synonym of Beringiophysa jennessii (Dall, 1919). 

Physa skinneri Taylor, 1954 F 

Taylor, 1954: 4, 9-10. 

● Type locality: "SE. cor. sec. 6, T.5 N., R. 28 E., Beaver Co., Oklahoma" (USA, Pleistocene). 

● Holotype: Museum of Zoology, University of Michigan No. 181292. 

● Distribution: Chaun lowland river basin (northeast of Siberia), in thermokarst waterbodies (Starobogatov et 

al., 2004). 

● Remark: Taylor and Sitnikova (2004) believed that Ph. skinneri is widely distributed in the former USSR. 

They mentioned findings of this species from Ukraine (Zhitomyr Region), Kazakhstan, Western and Eastern 

Siberia. 

Physa streletzkajae Starobogatov et Budnikova, 1976 F 

Starobogatov, Budnikova, 1976: 84, fig. 5 II. 

● Type locality: Anadyrskij district, Vakarevo, lake No. 6, station 5 [Chukchi Peninsula] 


● Distribution: basins of Yana and Anadyr rivers (northern Far East), in lakes (Starobogatov et al., 2004). 

Physa tei Starobogatov et Prozorova in Starobogatov, Prozorova et Zatrawkin, 1989 F 

Starobogatov et al., 1989: 74-75, fig. 3 9. 

● Type locality: lake near airport Anadyr (Chukchi Peninsula). 


● Distribution: lower Anadyr River (Chukchi Peninsula), in thermokarst waterbodies (Starobogatov et al., 

2004). 

● Remark: Taylor (2003) believed that Ph. tei is a synonym of Beringiophysa jennessii (Dall, 1919). 

Physella Haldeman, 1843 

Type species: Physa globosa Haldeman, 1841 (monotypy) 

Physella nuttallii (Lea, 1864) F 

Physa nuttallii Lea, 1864: 116. 

● Type locality: Lewis' River, Oregon. 

● Holotype: USNM No. 170770. 

● Distribution: Krest Bay coast (Chukchi Peninsula), in lakes (Starobogatov et al., 2004). 

ELLOBIOIDEA L. Pfeiffer, 1854 (1822) 


CARYCHIIDAE Jeffreys, 1830 

Carychium Müller, 1774 

Type species: Carychium minimum Müller, 1774 (monotypy) 

Carychium cymatoplax Pilsbry, 1901 T 

Pilsbry, 1901c: 23-24. 

● Type locality: Yaeyama Island [Loo Choo Islands, Japan]. 

● Lectotype (Baker, 1963): ANSP No. 80959a. 

● Distribution: southern Primorye (Prozorova, 2006; Prozorova, Kavun, 2007b).

Carychium lederi O. Boettger, 1880 T 

[= Carychium primitivum Schileyko, 1967] 

Boettger, 1880b: 383. 

● Type locality: not stated (Talysh – from the title). 


● Distribution: Lenkoran lowland and mountain forests of Talysh (Azerbaijan) (Likharev, Schileyko, MS). 

Carychium minimum Müller, 1774 T 

Müller, 1774: 125. 

● Type locality: Fridrichsdal (vicinities of Copenhagen). 


● Distribution: widely distributed in European forest zone (mainly in southern subzone of taiga and in mixed 

and broad-leaved forests), reaching Arkhangelsk in the north (Velichkovsky, 1927), Crimea and Caucasus 

inclusive in the south, and forest areas of South Ural (Krestyaninov, 1973) and northern, central, and eastern 

Kazakhstan (Uvalieva, 1981) in the east. 

● Remark: Uvalieva described 2 new species of Carychium, C. montanum and C. kasachstanicum, holotypes 

of which were said to be stored in ZIN. Examination of the "holotypes" has shown that (1) they do not correspond 

to descriptions and illustrations of the author; (2) they were collected two years after the publication, 

i.e. cannot be types; (3) they are typical C. minimum, and, therefore, both names are junior synonyms of this 

species. 

Carychium pessimum Pilsbry, 1901 T 

Pilsbry, 1901d: 562. 

● Type locality: Tane-ga-shima, Osumi [Japan]. 

● LectOtype (Baker, 1963): ANSP No. 82481a. 

● Distribution: Primorye, Sakhalin (Prozorova, 2006; Prozorova, Kavun, 2007b). 

Carychium sibiricum Westerlund, 1897 T 

[= Carychium pessimum Pilsbry, 1902; Carychium gerstfeldti Schlesch, 1938] 

Westerlund, 1897b: 124 

● Type locality: "Amur". 


● Distribution: deciduous and mixed forests of the Amur basin, Primorye, southern Sakhalin, Kurile Islands, 

southern Kamchatka (Likharev, Schileyko, MS). 

Carychium tridentatum (Risso, 1826) T 

Saraphia tridentata Risso, 1826: 84-85. 

● Type locality: not stated (Alpes Maritimes (southern France) – from title). 

● Types: probably lost (Arnaud, 1978). 

● Distribution: common in East Europe, mainly in mixed and deciduous forests (including Carpathians), 

spreading in the north to St.-Petersburg, in the south (through steppe zone along non-climax formations) – to 

mountain-forest areas of Crimea inclusive. Widely distributed in mountain forests of Caucasus, but absent in 

forests of Lenkoran lowland and Talysh (Likharev, Schileyko, MS). 

STYLOMMATOPHORA A. Schmidt, 1855 


SUCCINEIDAE Beck, 1837

SUCCINEINAE Beck, 1837 

Novisuccinea Pilsbry, 1948 

Type species: Succinea ovalis Say, 1817 (OD) 

Novisuccinea altaica (Martens, 1871) T 

Succinea altaica Martens, 1871: 46, 50. 

● Type locality: "Buchtarminsk" (later – Ust-Bukhtarma, now flooded by Bukhtarminskoe Water Reservoir; 

south-western Altai). 

● Lectotype (Kilias, 1973): ZMB No. 20616a. 

● Distribution: Tien-Shan (Kirgiz, Zailijskij, Kungei, Terskei ridges), Altai, Sayan, Tuva (Likharev, 

Schileyko, MS). 

Novisuccinea diserta Schileyko et Likharev, 1986 T 

Schileyko, Likharev, 1986: 215, figs 15, 16. 

● Type locality: Transbaikalia, Khamar-Daban Ridge, valley of Solzan River. 


● Distribution: coastal areas of southern Baikal (Likharev, Schileyko, MS). 

Novisuccinea evoluta (Martens, 1879) T 

[= Succinea granulosa Lindholm, 1927] 

Succinea evoluta Martens, 1879: 126. 

● Type locality: «Kuldsha» (Ining (Kuldja), northwest China). 


● Distribution: Tien-Shan (Kirgiz, Fergana, Talassky ridges), Altai, Transbaikalia (Likharev, Schileyko, MS). 

Novisuccinea lyrata (Gould, 1859) T 

[= Succinea horticola Reinhardt, 1877] 

Succinea lyrata Gould, 1859: 40. 

● Type locality: "Loo Choo" (Okinawa Island, Ryukyu Archipelago). 


● Distribution: South Kurile Islands, South Sakhalin (Likharev, Schileyko, MS). 

Novisuccinea martensiana (Nevill, 1878) T 

[= Succinea martensiana var. gigas Martens, 1882] 

Succinea martensiana Nevill, 1878: 5, pl. 1, figs. 30, 31. 

● Type locality: "Yarkand: Sasak Taka, Pasrobat", north-western China. 

● Syntypes: ZMB No. 30142. 

● Distribution: Tien-Shan (Likharev, Schileyko, MS). 

Novisuccinea strigata (L. Pfeiffer, 1855) T 

[= Succinea chrysis Westerlund, 1883] 

Succinea strigata Pfeiffer L., 1855: 297. 

● Type locality: "Port Clarence", Alaska. 


● Distribution: Kamchatka (Koryak National Area), Chukchi Peninsula (Likharev, Schileyko, MS). 

Pamirsuccinea Schileyko et Likharev, 1986 

Type species: Pamirsuccinea eximia Schileyko et Likharev, 1986 (OD) 

Pamirsuccinea eximia Schileyko et Likharev, 1986 T 

Schileyko, Likharev, 1986: 202, figs. 3, 4. 

● Type locality: Pamir, vicinities of Roshtkala village, south-eastward of Khorog.


● Distribution: species recorded in three localities in East Pamir: vicinities of Roshtkala, Yashikul, and Kudara 

villages on left bank of the Talimas River (Likharev, Schileyko, MS). 

Succinea Draparnaud, 1801 

Type species: Helix putris Linnaeus, 1758 (= Succinea amphibia Draparnaud, 1801) (SD Gray J.E., 1847) 

Succinea gladiator Schileyko et Likharev, 1986 T 


● Type locality: Tuva, mountain pass in Aradansky Ridge, on Kyzyl road, 190 km from Minusinsk. 



Succinea insularis Mousson, 1887 T 

Mousson, 1887: 20-21, pi. 1, fig. 5. 

● Type locality: "Ujsut" Island in the Amur River mouth. 

● Syntype-. ZMZ No. 519288. 


Succinea lauta Gould, 1859 T 

Gould, 1859: 422. 

● Type locality: «Hakodati» (Hakodate, southern Hokkaido, Japan). 

● Lectotype (Johnson, 1964): USNM No. 24129. 

● Distribution: Amur Region, southern Khabarovsk Territory, Primorye Territory, South Sakhalin, South Kurile 

Islands (Likharev, Schileyko, MS). 

Succinea putris (Linnaeus, 1758) T 

Helix putris Linnaeus, 1758: 774. 



● Distribution: nearly transpalearctic range; the species is replaced by a similar oriental species S. lauta in the 

Amur Region, southern Khabarovsk Territory, Primorye Territory, South Sakhalin, and South Kurile Islands 

(Likharev, Schileyko, MS). 

● Remarks: Due to high interpopulation variability, a lot of species and varieties of S. putris group have been 

described in the late 19th - early 20th centuries. The resolution of problem of their status requires an examination 

of type material or, at least, material from type localities. Since virtually all these forms were described 

from West Europe, at present it seems reasonable to refrain from ultimate judgements on their status 

in the absence of sufficient material. In any case, the material studied has shown the presence of a broad 

range of interpopulation variability of shell characters at constant anatomy. 

Succinella Mabille, 1870 

Type species: Succinea oblonga Draparnaud, 1801 (monotypy) 

Succinella oblonga (Draparnaud, 1801) T 

Succinea oblonga Draparnaud, 1801: 56. 

● Type locality: not stated (France – from title). 

● Syntype: NHMV, MNHN. 

● Distribution: Carpathians (Sverlova, 2006), East-European Plain, Crimea, Caucasus; spreading to Yenisei 

and West Altai in the east. Northern border of the range approximately coincides with that of mixed and 

broad-leaved forests (Likharev, Schileyko, MS). 

OXYLOMINAE Schileyko et Likharev, 1986

Oxyloma Westerlund, 1885 

Type species: Succinea hungarica Hazay, 1881 (= Succinea dunkeri L. Pfeiffer, 1865) (monotypy) 

Oxyloma ajanica Schileyko et Likharev, 1986 T 


● Type locality: vicinities of Ayan settlement on the Okhotsk Sea coast. 


● Distribution: coast of Okhotsk and northern Japan seas, Kurile Islands (except southern) (Starobogatov et 

al., 2004), Iturup Island (Gusarov, 1999); Sakhalin (Prozorova, Berezhok, 2004). 

Oxyloma dunkeri (L. Pfeiffer, 1865) T 

[= Succinea hungarica Hazay in Kobelt, 1880] 

Succinea dunkeri Pfeiffer L., 1865: 101. 

● Type locality: bog in delta of Danube. 

● Syntypes: ZMB. 

● Distribution: Ukraine – Kherson Region and vicinities of Melitopol (Sverlova, 2006); vicinities of Astrakhan, 

mountain Daghestan, Lenkoran lowland (Likharev, Schileyko, MS). 

Oxyloma elegans (Risso, 1826) T 

[= Succinea pfeifferi Rossmässler, 1835] 

Succinea elegans Risso, 1826: 59. 

● Type locality: not stated (Alpes Maritimes (southern France) – from title). 

● Syntypes: MNHN. 

● Distribution: Palearctic (Likharev, Schileyko, MS). 

Oxyloma hirasei (Pilsbry, 1901) T 

Succinea hirasei Pilsbry, 1901a: 348. 

● Type locality: Tsuchiura, Hitachi, eastern Hondo, Japan. 


● Distribution: Sakhalin (Prozorova, Berezhok, 2004). 

Oxyloma retusa (Lea, 1834) T 

Succinea retusa Lea, 1834: 117. 

● Type locality: vicinities of Cincinnati, Ohio, USA. 


● Distribution: several records in Kamchatka, in Azabachiya River valley (Likharev, Schileyko, MS); Iturup 

Island (Gusarov, 1999). 

Oxyloma sarsi (Esmark in Esmark et Hoyer, 1886) T 

[= Oxyloma lejavai Hudec, 1966] 

Succinea pfeifferi var. sarsi Esmark – Esmark, Hoyer, 1886: 108, pl. 5, fig. 2. 

● Type locality: not stated (northern Norway – from title). 


● Distribution: most part of Palearctic, eastward to Yakutsk (Likharev, Schileyko, MS). 

Oxyloma starobogatovi Schileyko et Likharev, 1986 T 


● Type locality: Gorno-Altai Autonomous Region, Alagansky District, Surulukol lake, northern shore. 


● Distribution: type locality.

Oxyloma stellifera (Schileyko, 1967) T 

Succinoides stelliferus Schileyko, 1967: 24, fig. 1. 

● Type locality: vicinities of Lenkoran. 


● Distribution: Lenkoran lowland (Likharev, Schileyko, MS). 

Succinoides Schileyko, 1967 

Type species: – Succinoides stelliferus Schileyko, 1967 (OD) 

Order GEOPHILA Férussac, 1821 

COCHLICOPIDAE Hesse, 1922 

Cochlicopa Férussac, 1821 

Type species: Helix lubrica Müller, 1774 (SD Westerlund, 1902) 

Cochlicopa collina (Drouët, 1855) T 

[= ?Cochlicopa lubrica var. ovata Jeffreys, 1862; ?Cochlicopa lubrica var. viridula Jeffreys, 1862; Zua lubrica 

var. olivea Locard, 1880; Cochlicopa lubrica var. lacteola Lindholm, 1911] 

Achatina collina Drouët, 1855: 46-47. 

● Type locality: «près de Lyon, а Mouy-de-l’Oise, а Liézey, dans les Vosges» (France). 


● Distribution: Russia: Moscow, Leningrad, Irkutsk regions; Ukraine: Podolia and mountain Crimea (Likharev, 

Schileyko, MS; Starobogatov, 1996b). 

Cochlicopa curta Clessin, 1908 T 

Cochlicopa lubrica var. curta Clessin, 1908: 8. 

● Type locality: not stated (vicinities of Regensburg, Austria – from title). 


● Distribution: Russia (southern Krasnoyarsk Region), Ukraine (Podolia and Zaporozhye Region), Georgia 

(Borzhomi) (Starobogatov, 1996b). 

Cochlicopa dushanbensis Starobogatov, 1996 T 

Starobogatov, 1996b: 119, fig. 4 F, G. 

● Type locality: Dushanbe, garden of Institute of Zoology and Parasitology of Tajikistan Academy of Sciences. 


● Distribution: mountain areas of Central Asia from southern Tajikistan (unknown from Pamir) to Ferghana 

depression (Starobogatov, 1996b). 

Cochlicopa hachijoensis Pilsbry, 1902 T 

Cochlicopa lubrica hachijoensis Pilsbry, 1902: 57. 

● Type locality. Hachijo-jima, Izu, Japan. 


● Distribution: Sakhalin, South Kurile Islands, Moneron Island (Prozorova, Berezhok, 2004; Prozorova, 

Kavun, 2007b). 

Cochlicopa heptapotamica Starobogatov, 1996 T 

Starobogatov, 1996b: 116-117, fig. 4B. 

● Type locality: Karatal River bank at Kesken settlement, Alma-Ata Region, Kazakhstan.


● Distribution: mountain regions of western Central Asia from Pamir to Zailijskij Alatau Ridge (Starobogatov, 

1996b). 

Cochlicopa izzatullaevi Starobogatov, 1996 T 

Starobogatov, 1996b: 118-119, fig. 4 D, E. 

● Type locality: floodland of Sorbo River, Romit Wildlife Reservation, Tajikistan. 


● Distribution: from southern Tajikistan (unknown from Pamir) to Zailijskij Alatau Ridge (Starobogatov, 

1996b). 

Cochlicopa kamchatica Starobogatov, 1996 T 

Starobogatov, 1996b: 119-120, fig. 5 A, B. 

● Type locality: Kamchatka, Milkovo, forest on the first terrace. 


● Distribution: Kamchatka (Starobogatov, 1996b); North Kurile Islands (Pearce et al., 2002). 

Cochlicopa kurenkovi Starobogatov, 1996 T 

Starobogatov, 1996b: 120-121, fig. 5C. 

● Type locality: Kamchatka, Milkovo, forest on the first terrace. 


● Distribution: Kamchatka (Starobogatov, 1996b); North Kurile Islands (Pearce et al., 2002). 

Cochlicopa kurilensis Starobogatov, 1996 T 

Starobogatov, 1996b: 122-123, fig. 5F. 

● Type locality: Shikotan Island (Kurile Islands), vicinity of Krabovaya (formerly Anama) inlet. 


● Distribution: Shikotan, Iturup, Kunashir, Zeleny islands (Kurile Islands), southern Sakhalin (Starobogatov, 

1996b). 

Cochlicopa likharevi Starobogatov, 1996 T 

Starobogatov, 1996b: 121, fig. 5D. 

● Type locality: Primorye Territory, Suputinsky Wildlife Reservation. 


● Distribution: Sikhote-Alin mountain system (Starobogatov, 1996b). 

Cochlicopa lubrica (Müller, 1774) T 

[= Bulimus nitidissimus Krynicki, 1833; Cionella columna Clessin, 1875; Cochlicopa repentina Hudec, 1960] 

Helix lubrica Müller, 1774: 104-105. 



● Distribution: forest and forest-steppe zones from West Europe to Irkutsk Region (Central Siberia) and to 

mountain regions of Central Asia (Likharev, Schileyko, MS); Kurile Islands (Shikotan, Paramushir, Atlasova) 

(Pearce et al., 2002). 

Cochlicopa lubricella (Ziegler in Porro, 1838) T 

[= Cochlicopa exigua Menke, 1830 (nom. nud.); Achatina lubrica var. fusiformis Picard, 1840; Glandina azorica 

Albers, 1852; Cionella pulchella Mousson, 1873; Cionella lubrica var. polita Westerlund, 1887, non 

Porro, 1838; Zua lubrica f. lacteola Lindholm, 1911] 

Bulimus lubricus var. lubricella Ziegler – Porro, 1838: 53-54. 

● Type locality: Comasca Province (Italy). 


● Distribution: Russia: European part, Siberia, Kamchatka, Primorye Territory (Ternei); Ukraine (Polesye to

steppe zone), Georgia (Borzhomi, Lagodekhi), Azerbaijan, Uzbekistan (mountain regions), Tajikistan (mountain 

regions) (Starobogatov, 1996b). 

Cochlicopa lubricoides (Potiez et Michaud, 1838) T 

[= Zua boisii Dupuy, 1850; Achatina nilssoni Malm, 1851; Bulimus maderensis Lowe, 1852; Bulimus subcylindricus 

var. exigua Moquin-Tandon, 1855; Bulimus lubricus var. lubricella Stabile, 1846, non Porro, 1838; 

Cochlicopa lubrica var. columna Clessin, 1875; Ferussacia crassula Fagot, 1879] 

Achatina lubricoides Potiez, Michaud, 1838: 129. 

● Type locality: «les Appennins, en Italie а Rimini». 

Types unknown. 

● Distribution: Russia: European part, Caucasus (Starobogatov, 1996b). 

Cochlicopa maacki Starobogatov, 1996 T 

Starobogatov, 1996b: 121-122, fig. 5E. 

● Type locality: Amur River near mouth of Sungary River. 


● Distribution: Amur territory and Sikhote-Alin mountain system, Sakhalin (Starobogatov, 1996b; Prozorova, 

Berezhok, 2004). 

Cochlicopa major (Bourguignat, 1864) T 

Ferussacia subcylindrica var. major Bourguignat, 1864a: 36. 

● Type locality: Alger and Oran (Algeria). 


● Distribution: Russia (Orenburg and Irkutsk regions), Ukraine (Poltava, Rovno, Chernigov, Kiev, Vinnitza 

regions) (Starobogatov, 1996b). 

Cochlicopa minima (Siemaschko, 1847) T 

[= Cionella lubrica var. litaunica Westerlund, 1887; Cionella lubrica var. pfeiffeti sensu Westerlund, 1874, 

non Weinland, 1874] 

Achatina minima Siemaschko, 1847: 111, pl. 1, fig. 4. 

● Type locality: "bei Riga" (Latvia). 


● Distribution: Ukraine (Ternopol Region) (Starobogatov, 1996b). 

Cochlicopa mukhitdinovi Starobogatov, 1996 T 

Starobogatov, 1996b: 117-118, fig. 4C. 

● Type locality: Kuramin Range near Bobokir village, Asht District, Khodjend Region, Tajikistan. 


● Distribution: from Pamir to Kuramin Range, probably also in mountain ranges of Tien-Shan system in 

north Tajikistan (Starobogatov, 1996b). 

Cochlicopa nitens (Gallenstein, 1852) T 

Achatina nitens Gallenstein, 1852: 75. 

● Type locality: Klagenfurt, Karintia (Austria). 


● Distribution: East-European Plain, Crimea, Caucasus, and southern Siberia (Likharev, Schileyko, MS). 

Cochlicopa pfeifferi (Weinland, 1874) T 

Cionella pfeifferi Weinland, 1874: 36, textfig. 

● Type locality: «in montibus, quos dicunt «Albem Suevicam» prope Hohen-Wittlingen» (Baden- 

Würtemberg, Germany). 


● Distribution: Russia (Orenburg Region), Ukraine (Ternopol Region) (Starobogatov, 1996b).

Cochlicopa pseudonitens Uvalieva, 1967 T 

Uvalieva, 1967: 215, figs. 2, 3. 

● Type locality: South Altai Mountains, floodplain of Malaya Narymka River. 


● Distribution: species found in three localities in South Altai: in Malaya Narymka river valley, on bank of 

Bolshaya Narymka river, and in vicinities of Verkhne-Katunsk village (Likharev, Schileyko, MS). 

● Remark: Records of the species from mountains of Uzbekistan and Tajikistan are based on misidentification 

(Starobogatov, 1996b). 

Cochlicopa shikotanica Starobogatov, 1996 T 

Starobogatov, 1996b: 122, fig. 5F. 

● Type locality: Shikotan Island (Kurile Islands), vicinity of Krabovaya (formerly Anama) inlet. 


● Distribution: Shikotan, Iturup, Kunashir, Zeleny islands (Kurile Islands), southern Sakhalin (Starobogatov, 

1996b). 

ORCULIDAE Steenberg, 1925 

ORCULINAE Steenberg, 1925 

Orcula Held, 1837 

Type species: Pupa dolium Draparnaud, 1801 (SD Herrmannsen, 1847) 

Orcula dolium (Draparnaud, 1801) T 

Pupa dolium Draparnaud, 1801: 58-59. 


● Syntype: NHMV. 

● Distribution: species not found on East European Plain, but may probably occur in Transcarpathia (Likharev, 


Orculella Steenberg, 1925 

Type species: Pupa orientalis L. Pfeiffer, 1861 (OD) 

Orculella bulgarica (Hesse, 1915) T 

Orcula bulgarica Hesse, 1915: 62. 

● Type locality: Bulgaria, vicinities of Varna. 


● Distribution: Caucasus: probably near Anapa (?), debris of Araks River near Darosham (Nakhichevan), and 

eastern coast of Sevan Lake near mouth of Masrik River. The species has never been found alive (Likharev, 


Orculella pfeifferi Hausdorf, 1996 T 

Hausdorf, 1996a: 28-29, textfig. 12, pi. 2, figs. 23-24. 

● Type locality. "V. Van: Insel Achtamar, 1750 m" (NE Turkey). 


● Distribution: single finding in Armenia ("Puraghan, N Ecmiadzin") (Hausdorf, 1996a). 

Orculella ruderalis Akramowski, 1947 T 

Orculella scyphus ruderalis Akramowski, 1947: 179, figs. 1, 2. 

● Type locality: Armenia, Ekhegnadzor district, Gnishik village, "Khozanner" area.


● Distribution: several localities near Gnishik village, Armenia (Likharev, Schileyko, MS). 

Pilorcula Germain, 1912 

Type species: Pupa raymondi Bourguignat, 1863 (monotypy) 

Pilorcula (?) aspinosa Hausdorf, 1996 T 

Hausdorf, 1996a: 56-57, pl. 6, fig. 74. 

● Type locality: "Georgien: 3 km E Tkvarceli". 


● Distribution: type locality (Hausdorf, 1996a). 

Pilorcula pusilla Hausdorf, 1996 T 

Hausdorf, 1996a: 57, pl. 6, fig. 75. 

● Type locality: "Georgien: Novyj Afon, Nadelwald am Psurrcha-Bach". 


● Distribution: type locality (Hausdorf, 1996a). 

Pilorcula trifilaris trifilaris (Mousson, 1863) T 

Pupa trifilaris Mousson, 1863: 391. 

● Type locality: Redutkaleh (northward of Poti). 

● Lectotype (Hausdorf, 1996a): ZMZ No. 514413a. 

● Distribution: Colchis lowland, western part of Great Caucasus northward to Maikop, along northern slopes 

of Great Caucasus to Vladikavkaz in the east (Likharev, Schileyko, MS). 

Pilorcula trifilaris longior Hausdorf, 1996 T 

[= Pupa (Orcula) doliolum var. intermedia Retowski, 1883, non Morelet, 1851; Pupa (Orcula) Raimondi var. 

bifilaris f. longior Retowski, 1889; Orcula raimondi O. Boettger, 1905, part.] 

Hausdorf, 1996a: 60-62, textfig. 28, figs. 82-83. 

● Type locality: "V. Zonguldak: Abas (ca. 10 km W Zonguldak)" (Turkey). 

•Holotype: SMF No. 311154. 

● Distribution: Crimea (Retowski, 1883, as Pupa (Orcula) doliolum var. intermedia), possibly Adzharia 

(deposits of the river Chorokh near Batumi) (Hausdorf, 1996a). 

Pilorcula trifilaris quadrifilaris (Rosen, 1905) T 

[= Pupa hifilaris sensu Retowski, 1883, non Mousson, 1873; Pupa hifilaris sensu O. Boettger, 1884, non 

Mousson, 1873; Pupa raimondi var. hifilaris sensu Rosen, 1905, part, non Mousson, 1873] 

Pupa Raimondi f. quadrifilaris Rosen, 1905: 57. 

● Type locality: "Im Psekupsauswurfe" (Krasnodar Territory). 

● Syntype: SMF No. 4593. 

● Distribution: maritime areas of Krasnodar Territory and Abkhazia (Hausdorf, 1996a). 

Schileykula Gittenberger, 1983 

Type species: Orcula batumensis sensu Hesse, 1924 (OD) 

Schileykula batumensis (Retowski, 1889) T 

[= Pupa kaznakovi Rosen, 1914] 

Pupa (Orcula) doliolum var. batumensis Retowski, 1889: 254. 

● Type locality: debris of the Chorokh river near Batumi. 

● Lectotype (Zilch in Neubert, 1993): SMF 3607a. 

● Distribution: Adzharia (a single doubtless finding in debris of the Chorokh river near Batumi) (Likharev,


Sphyradium Charpentier, 1837 

Type species: Bulimus doliolum Bruguière, 1792 (SD Martens in Albers, 1860) 

Sphyradium doliolum (Bruguière, 1792) T 

[= Pupa bifilaris Mousson, 1873] 

Bulimus doliolum Bruguière, 1792: 351. 

● Type locality: vicinities of Paris. 


● Distribution: Crimea, Caucasus (Likharev, Schileyko, MS); western and central Ukraine (Sverlova, 2006). 

PAGODULININAE Pilsbry, 1924 

Pagodulina Clessin, 1876 

Type species: Pupa pagodula Des Moulins, 1830 (monotypy) 

Pagodulina lederi (O. Boettger, 1886) T 

Pagodina pagodula var. lederi Boettger, 1886a: 305, pl. 3, fig. 8. 

● Type locality: Talysh. 


● Distribution: Elburz Range, including Talysh mountains (Likharev, Schileyko, MS). 

Pagodulina pagodula (Des Moulins, 1830) T 

Pupa pagodula Des Moulins, 1830: 158, figs. 1-3. 

● Type locality: Périgord (southwestern France, western part of Massif Centrale). 


● Distribution: probably occurs in Transcarpathian area (Likharev, Schileyko, MS). 

LAURIINAE Steenberg, 1925 

Euxinolauria Lindholm, 1924 

Type species: Pupa (Charadrobia) pulchra Retowski, 1883 (OD) 

Euxinolauria caucasica (L. Pfeiffer, 1857) T 

Pupa caucasica Pfeiffer, 1857: 88. 

● Type locality: Kazbek. 


● Distribution: broad-leaved forests of central and eastern areas of southern slopes of Great Caucasus (Likharev, 


Euxinolauria glomerosa Suvorov et Schileyko, 1991 T 

Suvorov, Schileyko, 1991: 76, figs. 6, 7. 

● Type locality: vicinities of Tzoniarisi village on Meskheti Ridge, Adzharia. 


● Distribution: type locality; found in Botanical Garden of Batumi (Egorov, Greke, 2005). 

Euxinolauria honesta Suvorov et Schileyko, 1991 T 

Suvorov, Schileyko, 1991: 75, fig. 5. 

● Type locality: mountain pass Beshumi on Arsiyan Ridge, Adzharia (2250 m above sea level).



Euxinolauria mica Schileyko, 1998 T 

Schileyko, 1998a: 77, fig. 81 (nom. nov. pro Euxinolauria zonifera sensu Schileyko, 1975c, 1984, non Lauria 

zonifera Pilsbry, 1934) 

● Type locality: Caucasian Reserve, vicinities of Guzeripl. 


● Distribution: Known only from several findings in type locality (near Guzeripl, western part of Great Caucasus) 


Euxinolauria nemethi (Hausdorf, 1996) T 

Leiostyla nemethi Hausdorf, 1996b: 111-112, fig. 1. 

● Type locality: Northwestern Caucasus: Mamta valley at km 26/33 in the gorge in direction to Krasnaya 

Poljana. 


● Distribution: Mamta and Hosta valleys (vicinities of Sochi, Krasnodar Territory) (Hausdorf, 1996b); vicinity 

of Sukhumi (Abkhazia) (Egorov, Greke, 2005). 

Euxinolauria paulinae (Lindholm, 1913) T 

[= Lauria paulinae unicolumellaris Lindholm, 1914; Leiostyla adolfi Pokryszko, 1991] 

Lauria paulinae Lindholm, 1913: 62-63. 

● Type locality: «Notanebi, 37 Werst nördlich von Batumi». 

● Lectotype (Schileyko, 1984): ZIN. 

● Distribution: maritime regions of Adzharia, 40-50 km around Batumi (Likharev, Schileyko, MS). 

Euxinolauria pulchra (Retowski, 1883) T 

[= Pupa pulchra f. bilabiata Retowski, 1887; Lauria pulchra var. nitens O. Boettger, 1888; Pupa pontica 

Retowski, 1889] 

Pupa (Charactrobia) pulchra Retowski, 1883: 57-58. 

● Type locality: Sudak (southern coast of Crimea). 


● Distribution: south-western and western parts of Great Caucasus, western part of Lesser Caucasus (Likharev, 


Euxinolauria rectidentata Schileyko, 1975 T 

Schileyko, 1975: 1769, figs. 1(1-2), 6(1). 

● Type locality: valley of the Machakheli-tzkali river in Adzharia. 


● Distribution: type locality, vicinities of Khelvachauri (Adzharia) (Egorov, Greke, 2005). 

Euxinolauria silicea Schileyko, 1975 T 

[= Lauria pulchra sensu Hudec et Lezhawa, 1969 non Pupa pulchra Retowski, 1883] 

Schileyko, 1975c: 1777, fig. 6, 8. 

● Type locality: Tekhuri river valley, south-western part of Great Caucasus, upstream of Taleri village. 


● Distribution: two findings in type locality. 

Euxinolauria sinangula Schileyko, 1975 T 

Schileyko, 1975c: 1775, fig. 3 (3-4), 6, 7. 

● Type locality: Batumi Botanical Garden. 


● Distribution: two findings in type locality.

Euxinolauria superstructa (Mousson, 1876) T 

[= Pupa superstructa var. lederi O. Boettger, 1883; Pupa superstructa var. unibasalis O. Boettger, 1886; 

Lauria zonata mut. albina O. Boettger, 1889] 

Pupa (Pupilla) superstructa Mousson, 1876a: 37-38, pl. 2, fig. 7. 

● Type locality: «Lailasch, gouvernement de Koutaпs» (Svaneti Ridge). 

● Syntypes: ZMZ Nos. 514508, 514509. 

● Distribution: Black Sea maritime area of Caucasus, Novorossiysk to Adzharia (Likharev, Schileyko, MS). 

Euxinolauria tenuimarginata (Pilsbry, 1922) T 

Lauria tenuimarginata Pilsbry, 1922: 76, pl. 10, figs. 1-2. 

● Type locality: Notanebi, 40 km N of Batumi. 

● Lectotype (Baker, 1963): ANSP No. 130116. 

● Distribution: species found in three localities of south-western Transcaucasia: near Notanebi station, 40 km 

northward of Batumi; near Chakvistavi village, 20 km eastward of Batumi; and in Batumi Botanical Garden 

at Zelenyi Mys, in Colchis forest grove (Likharev, Schileyko, MS). 

Euxinolauria vitrea (Schileyko, 1989) T 

Speleodentorcula vitrea Schileyko, 1988a: 1303, fig. 1 (I). 

● Type locality: West Caucasus, Verkhnekazachjebrodskaya Cave on River Mzymta. 


● Distribution: type locality; north-western vicinity of Adler (Black Sea coast), right side of Mzymta ravine, 

Ashtyr Cave (Egorov, Greke, 2005). 

Euxinolauria zonifera (Pilsbry, 1934) T 

[= Pupa superstructa var. zonata O. Boettger, 1883 non Pupa zonata Gassies, 1869] 

Lauria zonifera Pilsbry, 1934: 139 (nom. nov. pro Pupa zonata O. Boettger, 1883). 

● Type locality: Tzebelda on River Kodori, Abkhazia. 


● Distribution: western regions of Great Caucasus, Colchis lowland, and Black Sea maritime part of Lesser 

Caucasus (Likharev, Schileyko, MS). 

Lauria Gray, 1840 

Type species: Pupa umbilicata Draparnaud, 1801 (= Turbo cylindracea Da Costa, 1778) (SD Herrmannsen, 

1847) 

Lauria cylindracea (Da Costa, 1778) T 

[= Pupa umbilicata Draparnaud, 1801; Pupa sempronii Charpentier, 1837; Pupa dilucida Rossmässler, 1837; 

Lauria cymmetrica Puzanov, 1925] 

Turbo cylindraceus Da Costa, 1778: 89. 

● Type locality: Middlesex and Surrey, Lincolnshire, Morsley, Northamptonshire, Flintshire (England). 


● Distribution: mountain-forest Crimea, Transcaucasia, possibly Daghestan and West Kopetdagh (Likharev, 


ARGNINAE Hudec, 1965 

Argna Cossmann, 1899 

Type species: Pupa ferrari Porro, 1838 (OD) 

Argna bielzi (Rossmässler, 1859) T 

Pupa bielzi Rossmässler, 1859: 109, pl. 85, fig. 942. 

● Type locality: "Nagy Falu, unweit Bethlen" (southern Transylvania).

● Lectotype (Zilch, 1985): SMF No. 51716/2. 

● Distribution: mountain and foothill areas of Ukrainian (Forested) Carpathians (Likharev, Schileyko, MS). 

STROBILOPSIDAE Pilsbry, 1918 

Eostrobilops Pilsbry, 1927 

Type species: Strobilops hirasei Pilsbry, 1927 (OD) 

Eostrobilops coreana Pilsbry, 1927 T 

Strobilops coreana Pilsbry, 1927b: 43, pl. 10, figs. 8-10. 

● Type locality: vicinities of Pyongyang ((North Korea). 


● Distribution: Kedrovaya Pad Natural Reserve (southern Primorye Territory) (Likharev, Schileyko, MS). 

VALLONIIDAE Morse, 1864 

ACANTHINULINAE Steenberg, 1918 

Acanthinula Beck, 1847 

Type species: Helix aculeata Müller, 1774 (monotypy) 

Acanthinula aculeata (Müller, 1774) T 

Helix aculeata Müller, 1774: 81-82. 



● Distribution: most part of European territory of Russia, eastward to about 40º E, northward to latitude of 

Petrozavodsk; mountain Crimea, Caucasus, highland regions of Central Asia (not found in Kopetdagh) (Likharev, 

Schileyko, MS); western, central and northern Ukraine (Sverlova, 2006). 

Spermodea Westerlund, 1902 

Type species: Helix lamellata Jeffreys, 1830 (monotypy) 

Spermodea lamellata (Jeffreys, 1830) T 

Helix lamellata Jeffreys, 1830: 333. 

● Type locality: Scarborough, York, eastern England. 


● Distribution: Kaliningrad Region (Likharev, Schileyko, MS); probably western Ukraine (Sverlova, 2006).. 

Zoogenetes Morse, 1864 

Type species: Helix harpa Say, 1824 (monotypy) 

Zoogenetes harpa (Say, 1824) T 

[= Helix amurensis Gerstfeldt, 1859] 

Helix harpa Say, 1824: 256, pl. 15, fig. 1. 

● Type locality: North America. 


● Distribution: Circumboreal-alpine species. Russia: north of European part, Siberia, Kamchatka, Commander 

and Kurile islands, Sakhalin; to Sikhote-Alin along eastern coast of Asia; single record in Great Caucasus 

(near Teberda, 2200 m above sea level) (Likharev, Schileyko, MS; Prozorova, Berezhok, 2004).

VALLONIINAE Morse, 1864 

Planogyra Morse, 1864 

Type species: Helix asteriscus Morse, 1857 (monotypy) 

Planogyra asteriscus (Morse, 1857) T 

Helix asteriscus Morse, 1857: 128. 

● Type locality: Bethel, Oxford Co., Maine, USA. 

● Types; unknown. 

● Distribution: Russian Far East (Khabarovsk Territory, South Sakhalin, Moneron Island and southern Kurile 

Islands) (Prozorova, Kavun, 2007b as Planogyra sp.). 

Vallonia Risso, 1826 

Type species: Vallonia rosalia Risso, 1826 (= Helix pulchella Müller, 1774) (monotypy) 

● Remark: Morelet (1858) described Helix flocculus from vicinities of Petropavlovsk, Kamchatka. This species 

could belong to Vallonia or Punctum, but its status is uncertain, whereas the type material was not found 

(Gerber, 1996). 

Vallonia asiatica (Nevill, 1878) T 

[= Vallonia ladacensis sensu Schileyko, 1984, part] 

Helix (Vallonia) costata var. asiatica Nevill, 1878: 4. 

● Type locality: «Wakhan» (Afghanistan). 

● Lectotype (Gerber, 1996): Zoological Survey of India (Calcutta) No. M23209. 

● Distribution: Armenia, mountain regions of Central Asia (Gerber, 1996). 

Vallonia chinensis Suzuki, 1944 T 

[= Vallonia chinensis Tokunaga et Naora, 1939, nom. nud.] 

Suzuki, 1944: 375, pl. 33, figs. 1-4, 9, 10. 

● Type locality: «Kuhsiangtung, Harbin, Manchouko» (China, Manchuria, Haerhpin [Harbin]; fossil). 

● Holotype: «Sigenkagaku Kenskyusyo, Tokyo» (after Suzuki, 1944); apparently lost (Gerber, 1996). 

● Distribution: the species is known from a single Recent shell collected by Maak on «Amur» (Gerber, 

1996). 

Vallonia costata (Müller, 1774) T 

[= Vallonia rosalia Risso, 1826] 

Helix costata Müller, 1774: 31. 



● Distribution: Holarctic (Likharev, Schileyko, MS). 

Vallonia enniensis (Gredler, 1856) T 

[= Helix costellata Sandberger, 1875] 

Helix pulchella var. enniensis Gredler, 1856: 56. 

● Type locality: «Talgrund des Talfer-Baches bei Bozen» (Trentino-Alto Adige, northern Italy). 

● Lectotype (Gerber, 1996): SMF No. 193139a. 

● Distribution: Latvia, Transcarpathia, Ciscarpathia, Moscow Region, vicinities of Novorossijsk (Likharev, 


Vallonia excentrica Sterki in Pilsbry, 1893 T 

Pilsbry, 1893: 249-251, pl. 32, figs. 6-9. 

● Type locality: USA, New York, Staten Island. 

● Lectotype (Pilsbry, 1948): ANSP No. 10080.

● Distribution: species widely but locally distributed over Holarctic territory; presence in Siberia requires 

confirmation (Likharev, Schileyko, MS); Kunashir (Kurile Islands) (Pearce et al., 2002). 

Vallonia kamtschatica Likharev, 1963 T 

Vallonia cyclophorella kamtschatica Likharev, 1963: 70, fig. 2. 

● Type locality: Kamchatka, vicinities of Milkovo. 


● Distribution: Kamchatka, vicinities of Ayan (Okhotsk Sea coast) (Likharev, Schileyko, MS); Siberia 

(Daursky Ridge, Irkutsk, Krasnoyarsk, Sayan Ridge), Kunashir, Simushir (Kurile Islands) (Gerber, 1996); 

Altai (Porkyszko, Horsák, 2007). 

Vallonia ladacensis (Nevill, 1878) T 

[= Vallonia miserrima Gude, 1914] 

Helix (Vallonia) ladacensis Nevill, 1878: 4. 

● Type locality: «Mataian in the Dras valley (Ladak)» (India, Kashmir). 

● Lectotype (Rajagopal, Subba Rao, 1972): Zoological Survey of India (Calcutta) No. M18328. 

● Distribution: mountain regions of Central Asia (Tien-Shan) (Likharev, Schileyko, MS; Gerber, 1996; 

Porkyszko, Horsák, 2007). 

Vallonia mionecton mionecton (O. Boettger, 1889) T 

Helix (Vallonia) adela var. mionecton Boettger, 1889: 941, pl. 26, fig. 11 a-d. 

● Type locality: «Transcaspien. Auf dem Gipfel des Agh-dagh im Kopet-dagh» (northern Iran). 


● Distribution: Tajikistan: Zeravshan and Hissar Ridges (Gerber, 1996). 

Vallonia mionecton schamhalensis Rosen, 1892 T 

Vallonia mionecton var. schamhalensis Rosen, 1892: 125. 

● Type locality: «Schamhala in Chorassan» (northern Iran). 

● Lectotype (Gerber, 1996): SMF No. 3742a. 

● Distribution: Tajikistan/Uzbekistan: Chasret-Sultan Mountains (Gerber, 1996). 

Vallonia patens patens Reinhardt, 1883 T 

[= Vallonia costata var. amurensis Sterki in Pilsbry, 1893] 

Reinhardt, 1883: 43. 

● Type locality: «Powantschan in China (Prov. Chili)» (China, Hebei: «Bao-hwa-shan»). 

● Lectotype (Yen, 1939): SMF No. 42466. 

● Distribution: besides two records in the Amur valley, the species was also found in forests of Primorye Territory, 

in vicinities of Vladivostok and on many islands in the Peter the Great Bay (Likharev, Schileyko, MS, 

as Vallonia amurensis; Gerber, 1996). 

Vallonia peteri Schileyko, 1984 T 

Vallonia tenuilabris peteri Schileyko, 1984: 169, fig. 90 (VI). 

● Type locality: Prymorye Territory, Peter the Great Bay, Kozlov Island. 


● Distribution: Subspecies found on following islands of Peter the Great Bay: Kozlov, Dva Brata, Verkhovsky, 

Pakhtusov, Vera (Likharev, Schileyko, MS). 

Vallonia pulchella (Müller, 1774) T 

[= Helix adela Westerlund, 1874] 

Helix pulchella Müller, 1774: 30. 



● Distribution: Holarctic (Likharev, Schileyko, MS).

Vallonia pulchellula tenerrima Gerber, 1996 T 

Gerber, 1996: 136-137, figs. 2h, 3r, 48d, 52a-c. 

● Type locality: Vladivostok, SSE part of Egersheld Peninsula, ca. 200 m from refueling station toward Cape 

Tokarevskogo. 

● Holotype: Zoologische Staatssammlung, München No. 1885/1. 

● Distribution: Vladivostok (Gerber, 1996). 

Vallonia tenuilabris (Al. Braun, 1843) T 

Helix tenuilabris Braun, 1843: 143. 

● Type locality: «im Löss von Wiesbaden» (Germany, Hessen). 


● Distribution: Siberia, mountain regions of Central Asia, Amur basin (Likharev, Schileyko, MS). 

Vallonia zaru Almuhambetova, 1979 T 

Almuhambetova, 1979: 32, fig. 3. 

● Type locality: Central Asia, Zailijskij Ridge, canyon of Turgen River, Tau-Turgen sanatorium. 

● Holotype: lost. 


PUPILLIDAE Turton, 1831 

Gibbulinopsis Germain, 1919 

Type species: Orthogibbus pupula Deshayes, 1863 (OD) 

Gibbulinopsis cryptodon (Heude, 1880) T 

[= Pupa heudeana Moellendorff, 1884; Pupilla heudeana grandis Moellendorff, 1901] 

Pupa cryptodon Heude, 1880a: 77, pl. 18, fig. 20. 

● Type locality: «environs de la ville de Houai-ugan» (south-eastern China). 

● Syntypes: SMF No. 4601. 

● Distribution: islands of Peter the Great Bay (Likharev, Schileyko, MS). 

Gibbulinopsis gracilis (Izatullaev, 1970) T 

Pupilla gracilis Izzatullaev, 1970a: 53, fig. 1. 

● Type locality: Tajikistan, Hissar Ridge, debris of Sardai-Mion River, near Romit settlement. 



● Remark: Species was described from a single shell, so its validity requires confirmation. It cannot be excluded 

that it is a high form of G. signata, because a reduction of upper palatal fold occurs in some populations 

of the latter species. Arkhangelsky (1933) reported findings of Pupilla sp. in Samarkand vicinities, on 

the River Siab banks, in Urgut and Sary-Mazar. He wrote that it "differs from other Pupilla in size (up to 5 

mm), brown color, and oblique, well seen striation". Arkhangelsky distinguished this form from G. signata, 

because this species was mentioned separately. Probably, the author dealt with G. gracilis. 

Gibbulinopsis interrupta (Reinhardt in Martens, 1876) T 

Bulimus (Pupilla) interrupta Reinhardt – Martens, 1876: 367. 

● Type locality: Borzhomi. 


● Distribution: Central part of Nort Caucasus, Daghestan, central and eastern Transcaucasia (Likharev, 

Schileyko, MS).

Gibbulinopsis nanosignata Schileyko et Izatullaev, 1980 T 

Schileyko, Izzatullaev, 1980: 283, fig. 

● Type locality: Hissar Ridge, right bank of Iskanderdarja River. 


● Distribution: northern foothills of eastern Turkestan Ridge (Vorukh settlement; also found in debris of Isfara 

River); Hissar Ridge, near lakes Iskanderkul and Varenkul, on banks of Iskanderdarja River (Likharev, 


Gibbulinopsis signata (Mousson, 1873) T 

[= ? Pupa armeniaca Issel, 1865; Pupa cristata Martens, 1874; Pupa signata var. parvula Mousson, 1876; 

Pupa signata var. cylindrica Mousson, 1876; Pupa diecki Gredler, 1889; Pupa signata var. cyclostoma Westerlund, 

1893; Pupa signata var. debilis Westerlund, 1893] 

Pupa signata Mousson, 1873: 211-212, pl. 8, fig. 7. 

● Type locality: debris of Araks River. 


● Distribution: Eastern Transcaucasia, southern Central Asia, south-eastern Kazakhstan (Likharev, Schileyko, 

MS). 

Pupilla Leach, 1828 

Type species: Pupa marginata Draparnaud, 1801 (= Turbo muscorum Linnaeus, 1758) (monotypy) 

Pupilla alabiella Schileyko, 1984 T 

Schileyko, 1984: 191, figs. 110 (II), 111. 

● Type locality: Verkhovsky Island, Peter the Great Bay. 


● Distribution: two small rocky islands in the Peter the Great Bay (Verkhovsky and Dva Brata) (Likharev, 


Pupilla altaica Meng et Hoffmann, 2008 T 

Meng, Hoffmann, 2008: 230, fig. 5. 

● Type locality: Central Altay, SE of Inya, dry steppe, carbonate rocks, on the edge of Chuya-Katun meadowland, 

824 above sea level, 50º23.933’N, 86º40.576’E. 

● Holotype: Zoological Museum Dresden, MTD Moll 53311. 

● Distribution: Central Altay (Meng, Hoffmann, 2008). 

Pupilla alluvionica Meng et Hoffmann, 2008 T 

Meng, Hoffmann, 2008: 231, 233-234, fig. 6. 

● Type locality: South-Eastern Altay, Saylyugem, open and active alluvions, in the flood plain of the river in 

spate, 2224 above sea level, 50º01.048’N, 89º14.177’E. 

● Holotype: Zoological Museum Dresden, MTD Moll 53315. 

● Distribution: Central and Eastern Altay (Meng, Hoffmann, 2008). 

Pupilla anzobica Izzatullaev, 1970 T 

Izzatullaev, 1970: 54, fig. 2. 

● Type locality: Hissar Ridge, Anzob mountain pass (3450-3500 m above sea level). 



Pupilla bigranata (Rossmässler, 1839) T 

Pupa bigranata Rossmässler, 1839: 27, fig. 645. 



● Distribution: West Ukraine, valleys of some tributaries of the Volga (Moscow River, Oka); southern parts

of Ural with adjacent territories of Kazakhstan and West Siberia; mountain areas of Central Asia and southern 

Kazakhstan; several records in East and North Caucasus (Likharev, Schileyko, MS). 

Pupilla bipapulata Akramowski, 1947 T 

Akramowski, 1947: 255. 

● Type locality: Armenia, Abovyan Region, Gegart. 


● Distribution: South and south-eastern Armenia: Azat River basin to Daralagez to Kafan and Megrin regions 


Pupilla gallae Tzvetkov, 1940 T 

Tzvetkov, 1940a: 384. 

● Type locality: South Kazakhstan, Zailijskij Ridge, Issyk ravine, 50 km eastward of Alma-Ata. 


● Distribution: mountain systems of south-eastern Kazakhstan and north-eastern Kirgizia: Zailijskij, Terskei, 

and, probably, Kungei ridges (Likharev, Schileyko, MS). 

Pupilla inequidenta Schileyko et Almuhambetova in Almuhambetova, 1979 T 


● Type locality: floodplain of Aksai River, Zailijskij Ridge. 


● Distribution: several localities in Zailijskij Ridge (Likharev, Schileyko, MS). 

Pupilla inops (Reinhardt, 1877) T 

[= Pupilla muscorum var. caucasica O. Boettger, 1879] 

Pupa triplicata var. inops Reinhardt, 1877a: 79, pl. 3, fig. 3. 



● Distribution: East Ciscaucasia and Transcaucasia (Likharev, Schileyko, MS). 

Pupilla limata Schileyko, 1984 T 

Schileyko, 1984: 193, figs. 110 (I), 112. 

● Type locality: Chukchi Peninsula, vicinities of Markovo. 


● Distribution: two localities in Chukchi Peninsula: vicinities of Markovo and bank of the Amguema River 

along Egvekinot-Iultin road (Likharev, Schileyko, MS). 

Pupilla muscorum (Linnaeus, 1758) T 

[= Pupa marginata Draparnaud, 1801; Pupa unidentata C. Pfeiffer, 1821 non Studer, 1820; ? Pupa cupa Jan, 

1832; Pupa muscorum f. albina Menke, 1830; Pupa muscorum f. edentula Menke, 1830; Pupa muscorum 

var. pratensis Clessin, 1871; Pupa muscorum var. lundstromi Westerlund, 1876; Pupa muscorum var. caucasia 

O. Boettger, 1880; Pupa muscorum var. glis Westerlund, 1893; Pupa muscorum asiatica Moellendorff, 

1901; Pupilla muscorum var. milaschevitschi Lindholm, 1911] 

Turbo muscorum Linnaeus, 1758: 767. 

● Type locality: Sweden. 

● Syntype: BMNH. 


Pupilla sterri (Voith in Fürnrohr, 1840) T 

[= Pupa cupa auct., non Jan, 1832; Pupa cupa var. carpathica Kimarowicz, 1890; Pupilla wolfae Tzvetkov, 

1940] 

Pupa sterri Voith – Fürnrohr, 1840: 469. 

● Type locality: Regensburg, Bayern.


● Distribution: Caucasus (vicinities of Tbilisi); Carpathians (Baidashnikov, 1985); Central Asia (Likharev, 


Pupilla striopolita Schileyko, 1984 T 

Schileyko, 1984: 188, fig. 106. 

● Type locality: Tien-Shan, Chatkal Ridge, Kasankai River bank near Ala-Buka village. 


● Distribution: Central Asia: Chatkal, Talas, Kirgiz, Zailijskij ridges (Likharev, Schileyko, MS). 

Pupilla triplicata (Studer, 1820) T 

[= ? Pupa triplicata var. luxurians Reinhardt, 1877; Pupa triplicata var. suboviformis O. Boettger, 1879; 

Pupa triplicata var. cylindrata O. Boettger, 1879] 

Glischrus (Pupa) triplicata Studer, 1820: 89. 

● Type locality: "Kt. Waadt bei Bex-Vieux", Switzerland. 


● Distribution: Carpathians; Crimea; Nicolaev and DOnetzk regions of Ukraine (Sverlova, 2006); isolated 

populations in basins of the Don (Novocherkassk), Volga, and Oka; North Caucasus and Transcaucasia; Central 

Asia, southern Kazakhstan, southern Altai; most eastern record – on Baikal shore, near Cape Krestovsky 


Pupilla turcmenica O. Boettger, 1889 T 

Pupa (Pupilla) cupa var. turcmenica Boettger, 1889: 958, pl. 26, fig. 3a-c. 

● Type locality: Kopetdagh, Ag-Dagh Mountain; 


● Distribution: Central Asia (Kopetdagh, Hissar, Kirgiz, Zailijskij ridges) (Likharev, Schileyko, MS). 

Pupoides L. Pfeiffer, 1854 

Type species: Bulimus nitidulus L. Pfeiffer, 1839 (SD Kobelt, 1880) 

Pupoides coenopictus (Hutton, 1834) T 

Pupa coenopicta Hutton, 1834: 85. 

● Type locality: India, Bombay. 


● Distribution: species found in debris of the Vakhsh River near "Tigrovaya Balka" Reserve; probably occurring 

in basin of middle or upper reaches of this river (southern Tajikistan) (Izzatullaev, 1970b; Likharev, 


GASTROCOPTIDAE Pilsbry, 1918 

Gastrocopta Wollaston, 1878 

Type species: Pupa acarus Benson, 1856 (OD) 

Gastrocopta huttoniana (Benson, 1849) T 

[= Pupa theeli Westerlund, 1898, non Westerlund, 1877] 

Pupa huttoniana Benson, 1849: 126. 

● Type locality: Simla (Himalayas, North India). 


● Distribution: species repeatedly found in debris of Kafirnigan River near Dushanbe. Apparently, mollusks 

live in this river valley, between Hissar and Karategin ridges (Likharev, Schileyko, MS).

Gastrocopta theeli (Westerlund, 1876) T 

[= Pupa denudata Mousson, 1887; Gastrocopta coreana Pilsbry, 1916] 

Pupa theeli Westerlund, 1876a: 102, fig. 4. 

● Type locality: Mikulino, vicinities of Yeniseisk. 

● Lectotype (Schileyko, 1984): SMNH No. 1657. 

● Distribution: species distributed patchily: South Primorye, vicinities of Yeniseisk and Chelyabinsk, South 

Altai, Kirgiz Ridge; basins of Kura and Rioni rivers (found in debris); highland of Daghestan (Botlikh) and 

North Caucasus (middle reaches of Chegem River) (Likharev, Schileyko, MS). 

VERTIGINIDAE Fitzinger, 1833 

Vertigo Müller, 1774 

Type species: Vertigo pusilla Müller, 1774 (monotypy) 

Vertigo antivertigo (Draparnaud, 1801) T 

[= Turbo sexdentatus Montagu, 1803; Alaea palustris Jeffreys, 1830; Vertigo sinuata Mousson, 1873] 

Pupa antivertigo Draparnaud, 1801: 57. 


● Types: probably in NHMV. 

● Distribution: Most part of Palearctic: eastward to Transbaikalia inclusive, northward to zone of tundra 


Vertigo circumlabiata Schileyko, 1984 T 

Schileyko, 1984: 208, fig. 129. 

● Type locality: Kunashir Island, vicinities of Sernovodsk City (South Kurile Islands). 


● Distribution: Kunashir Island (South Kurile Islands), Okhotsk Sea coast near Ayan, Sakhalin (Likharev, 

Schileyko, MS; Prozorova, Berezhok, 2004). 

Vertigo eogea Pilsbry, 1919 T 

Pilsbry, 1919: 151-152, pl. 14, figs. 1-3. 

● Type locality: Akkeshi, Kushiro, Yesso (Hokkaido, Japan). 


● Distribution: Paramushir, Shumshu, Atlasova Islands (Kurile Islands) (Kuroda, Koba, 1933). 

Vertigo extima (Westerlund, 1876) T 

[= Vertigo modesta ultima Pilsbry, 1919] 

Pupa (Vertigo) arctica var. extima Westerlund, 1876a: 42. 

● Type locality: Baklanikha on the Yenisei. 

● Lectotype (Waldén, 1986): SMNH Westerlund collection No. 8: 145. 

● Distribution: Yenisei (vicinities of Baklanikha) and Ob (Pokur settlement, Surgut District, Tyumen Region) 

basins, Chukchi Peninsula (Amguema and Lavrentiya rivers) (Likharev, Schileyko, MS); Karelia (von 

Proschwitz, 2004). 

Vertigo hirasei Pilsbry, 1901 T 

Pilsbry, 1901b: 128. 

● Type locality: «Yanagawa, prov. Chikugo, Kiusiu» (Kyushu, Japan). 

● Lectotype (Pilsbry, 1919): ANSP No. 79738a. 

● Distribution: Shikotan Island (South Kurile Islands) (Likharev, Schileyko, MS). 

Vertigo hydrophila (Reinhardt, 1877) T 

Pupa (Vertigo) hydrophila Reinhardt, 1877b: 323, pl. 11, fig. 6.

● Type locality: «Hakotade insulae Yesso» (Hokkaido, Japan). 

● Syntype: ZMB No. 28178. 

● Distribution: South Kurile Islands (Iturup, Kunashir) (Pearce et al., 2002). 

Vertigo japonica Pilsbry et Hirase, 1904 T 

[= Vertigo coreana Pilsbry, 1919; Vertigo tosana Pilsbry, 1919] 

Pilsbry, Hirase, 1904a: 118. 

● Type locality: Ikushagawa, Oshima Islands, Japan. 

● Holotype: ANSP No. 85746a. 

● Distribution: South Primorye, South Kurile Islands, coast of Tatar Strait (vicinities of Vanino) and Okhotsk 

Sea (Ayan) (Likharev, Schileyko, MS). 

Vertigo kushiroensis Pilsbry et Hirase, 1905 T 

Vertigo hirasei kushiroensis Pilsbry, Hirase, 1905: 718. 

● Type locality: Akkeshi, Kushiro (Hokkaido, Japan). 


● Distribution: Shikotan Island (South Kurile Islands) (Likharev, Schileyko, MS). 

Vertigo microsphaera Schileyko, 1984 T 

Schileyko, 1984: 210, fig. 132. 

● Type locality: Podutesnaya Bay, Bering Island. 


● Distribution: Bering Island (Commander Islands), Shikotan and Kunashir Islands (Kurile Islands) 

(Schileyko, 1984; Pearce et al., 2002); Altai (Pokryszko, Horsák, 2007). 

Vertigo modesta modesta (Say, 1824) T 

[= Pupa hoppii Møller, 1842; Pupa shuttleworthiana L. Pfeiffer, 1847; Pupa decora Gould, 1848; Pupa 

genesii Gredler, 1856; Pupa borealis Morelet, 1858; Vertigo arctica Wallenberg, 1858; Pupa ronnebyensis 

Westerlund, 1871; Pupa inermis Westerlund, 1877 non Deshayes, 1863; Vertigo geyeri Lindholm, 1925; Vertigo 

westerlundi Pilsbry, 1922] 

Pupa modesta Say, 1824: 259, pl. 15, fig. 5. 

● Type locality: vicinities of Lake Superior (NW North America). 


● Distribution: north of Eurasia including West and East Siberia (Likharev, Schileyko, MS); Kurile Islands 

(Pearce et al., 2002); Sakhalin (Prozorova, Berezhok, 2004). 

Vertigo modesta alpestris Alder, 1838 T 

Vertigo alpestris Alder, 1838: 340. 

● Type locality: Northumberland, England. 


● Distribution: Crimea (Chatyrdagh), Siberia, Sikhote-Alin with foothills (Likharev, Schileyko, MS). 

Vertigo moulinsiana (Dupuy, 1849) T 

[= Pupa laevigata Kokeil in Gallenstein, 1852; Vertigo ventrosa Heynemann, 1862; Pupa lilljeborgi Westerlund, 

1868; Pupa küsteriana Westerlund, 1875] 

Pupa moulinsiana Dupuy, 1849: [3], No. 284. 



● Distribution: East-European Plain, to Lithuania in the north, Volga River in the east and southern Armenia 

in the South (Likharev, Schileyko, MS). 

Vertigo pseudosubstriata Ložek, 1954 T 

[= Vertigo laevis Uvalieva, 1967]

Ložek, 1954: 327, pl. 1, fig. 1. 

● Type locality: Czechia, Moravia region, excavation of Paleolithic settlement in Dolni Vestonice above 

River Dyi. 

● Holotype: Geological Institute, Prague. 

● Distribution: South Altai and Zailijskij, Jungar, Terskei ranges (Likharev, Schileyko, MS); Sakhalin (Prozorova, 

Berezhok, 2004; Prozorova, Kavun, 2007b). 

Vertigo pusilla Müller, 1774 T 

[= Vertigo heterostropha Leach in Turton, 1831] 

Müller, 1774: 124. 



● Distribution: East-European Plain, eastward to the Volga, northward to approximately 60º N; Caucasus, 

North Kazakhstan, Altai (Likharev, Schileyko, MS); Carpathians (Sverlova, 2006). 

Vertigo pygmaea (Draparnaud, 1801) T 

[= Alaea vulgaris Jeffreys, 1830] 

Pupa pygmaea Draparnaud, 1801: 57. 


● Syntypes: NHMV. 

● Distribution: East-European Plain, to Lithuania in the north, Volga River in the east and southern Armenia 

in the south (Likharev, Schileyko, MS), Altai (Pokryszko, Horsák, 2007), Carpathians (Sverlova, 2006). 

Vertigo sieversi (O. Boettger, 1879) T 

[= Pupa (Vertigo) sieversi var. punctum O. Boettger, 1880; Pupa (Vertigo) sieversi var. subalpestris O. 

Boettger, 1880; Pupa (Vertigo) sieversi var. substriata Likharev et Rammelmeyer, 1952] 

Pupa (Vertigo) sieversi Boettger, 1879b: 407-408, pl. 10, figs. 6, 7. 

● Type locality: «Tabizhuri» (vicinities of Tabatzkuri Lake, South Georgia, Trialeti Ridge) 

● Holotype: unknown. 

● Distribution: Species widely but locally distributed in central and western parts of North Caucasus and 

Transcaucasia (Likharev, Schileyko, MS). 

Vertigo substriata (Jeffreys, 1830) T 

[= Pupa curta Held, 1837; Pupa striata Reeve, 1863; Pupa pygmaea var. nitidula Mousson, 1876; Pupa substriata 

var. mitis O. Boettger, 1880; Alaea substriata f. viridana Lindholm, 1910] 

Alaea substriata Jeffreys, 1830: 515. 

● Type locality: England, Burnstaple. 


● Distribution: East-European Plane, except nothernmost areas, North Caucasus and Georgia; an isolated part 

of range in Altai (Likharev, Schileyko, MS), Carpathians (Sverlova, 2006). 

Vertilla Moquin-Tandon, 1855 

Type species: Vertigo plicata A. Müller, 1838 (= Vertigo angustior Jeffreys, 1830) (SD Pilsbry, 1929) 

Vertilla angustior (Jeffreys, 1830) T 

[= Vertigo hamata Held, 1837; Vertigo plicata A. Müller, 1838] 

Vertigo angustior Jeffreys, 1830: 361. 

● Type locality: "Marino near Swansea" (England). 


● Distribution: Lithuania, Estonia, Russia (Leningrad, Moscow, Nizhnyi Novgorod, Kirov regions, Tatarstan), 

western Byelorussia and Ukraine, mountain Crimea, North Caucasus and Transcaucasia (Likharev, 

Schileyko, MS); southern Krasnoyarsk Region (Udaloi, Novikov, 2005a).

TRUNCATELLINIDAE Steenberg, 1925 

Columella Westerlund, 1878 

Type species: Pupa inornata Clessin, 1872 (= Pupa edentula Draparnaud, 1805) (SD Westerlund, 1887) 

Columella acicularis Almuhambetova, 1979 T 


● Type locality: Zailijskij Ridge, Tau-Turgen Valley. 



Columella aspera Waldén, 1966 T 

Waldén, 1966: 53, fig. 1. 

● Type locality: Västmanland, Linde Kirchspiel, im Nadewald ö Siggebohyttan [Sweden]. 

● Holotype: NMG. 

● Distribution: Valdai Hills (Shikov, 1979); south Siberia near Baikal Lake (Prozorova et al., 2007). 

Columella columella (G. Martens, 1830) T 

[= Pupa gredleri Clessin, 1872; Pupa regularis Westerlund, 1898] 

Pupa columella Martens, 1830: 171. 

● Type locality: «Württemberg, zwischen Heslach und Kalten Thal bei Stuttgart». 

● Lectotype (Forcart, 1959a): NMB No. 5900a. 

● Distribution: Ukraine (Carpathians, Podolsk Hills, Kiev Region, mountain Crimea) (Sverlova, 2006), Transcaucasia 

(vicinities of Kadjaran, Armenia and Kaputdjukh Mount, Lesser Caucasus), mountain systems of 

Central Asia, Siberia (Likharev, Schileyko, MS); Sakhalin (Prozorova, Kavun, 2007b). 

Columella edentula (Draparnaud, 1805) T 

[= Vertigo lepidula Held, 1837; Pupa inornata m. turritella Westerlund, 1875; Pupa edentula f. nana O. 

Boettger, 1880] 

Pupa edentula Draparnaud, 1805: 59-60, pl. 3, figs. 28, 29. 


● Lectotype (Pokryszko, 1990; as «Typus»): NHMV. 

● Distribution: East-European Plain (except for steppe area), Caucasus and Transcaucasia, Central Asia (Likharev, 

Schileyko, MS); Russian Far East including Sakhalin, Kurile Islands and Kamchatka (Pearce et 

al., 2002; Prozorova et al., 2007); Altai (Pokryszko, Horsák, 2007). 

Columella intermedia Schileyko et Almuhambetova in Schileyko, 1984 T 

Schileyko, 1984: 219, fig. 139 (III). 

● Type locality: Talasskij Ridge, Bakhrausu ravine. 


● Distribution: type locality; Altai (Pokryszko, Horsák, 2007). 

Truncatellina Lowe, 1852 

Type species: Pupa linearis Lowe, 1852 (monotypy) 

Truncatellina callicratis (Scacchi, 1833) T 

[= Pupa rivierana Benson, 1854; Pupa laevistriata Retowski, 1888] 

Turbo callicratis Scacchi, 1833: 11. 

● Type locality: Naples, Botanical Garden [Italy]. 


● Distribution: North Caucasus, Transcaucasia, Kopetdagh, mountain areas of eastern Central Asia, South 

Altai (Likharev, Schileyko, MS).

Truncatellina claustralis (Gredler, 1856) T 

[= Pupa claustralis var. anodus Gredler, 1856; Pupa clavella Reinhardt, 1877] 

Pupa claustralis Gredler, 1856: 116, pl. 2, fig. 1a-c. 

● Type locality: Sallegg [Tirol, Austria]. 


● Distribution: Ciscarpathia, Crimea, Central Transcaucasia (Likharev, Schileyko, MS). 

Truncatellina costulata (Nilsson, 1822) T 

[= Pupa ascaniensis Schmidt, 1849] 

Pupa costulata Nilsson, 1822: 51. 

● Type locality: «Ad Esperöd Scaniae». 


● Distribution: East-European Plain: eastward to the Volga, northward approximately to Estonia-Moscow- 

Uljyanovsk line; Crimea, Transcaucasia, Central Asia (Likharev, Schileyko, MS). 

Truncatellina cylindrica (Férussac, 1807) T 

[= Pupa minutissima Hartmann, 1821; Vertigo pupila Held, 1837; Pupa micula Mousson, 1876; Truncatellina 

cylindrica var. costigerella Lindholm, 1926; Truncatellina tauricola Lindholm, 1926] 

Vertigo cylindrica Férussac, 1807: 52. 



● Distribution: East-European Plain: northward to Estonia, Moscow Region, eastward to South Ural; southward 

to Carpathians, Crimea, North Caucasus, Transcaucasia (Likharev, Schileyko, MS); Altai (Pokryszko, 

Horsák, 2007). 

CHONDRINIDAE Steenberg, 1925 

Chondrina Reichenbach, 1828 

Type species: Bulimus avenaceus Bruguière, 1792 (SD Reichenbach, 1836) 

Chondrina amphorula Schileyko, 1984 T 

Schileyko, 1984: 231, figs. 148 (III), 150. 

● Type locality: vicinities of Verkhnee Inkhelo village, near Karata settlement, Daghestan. 


● Distribution: Daghestan, North Caucasus (valleys of rivers Chegem and Terek), Armenia (Idzhevan District). 

Chondrina clienta clienta (Westerlund, 1883) T 

[= Glischrus hordeus Studer, 1820, nom. oblit.; Pupa avenacea var. paucidens Westerlund, 1887; Pupa subhordeum 

Westerlund, 1887] 

Pupa avenacea var. clienta Westerlund, 1883a: 60. 

● Type locality: «Choè im Tatra Gebirge». 

● Lectotype (Gittenberger, 1973): NMG No. 2174. 

● Distribution: Carpathians (Likharev, Schileyko, MS). 

Chondrina clienta caucasica Ehrmann, 1931 T 

Ehrmann, 1931: 19, pl. 1, fig. 4. 

● Type locality: Lechkhum Ridge (south slopes of Great Caucasus). 

● Lectotype (Gittenberger, 1973): SMF No. 44091a. 

● Distribution: mountain Crimea and Caucasus (Likharev, Schileyko, MS).

Chondrina granum (Draparnaud, 1801) T 

[= Pupa profuga Westerlund, 1898] 

Pupa granum Draparnaud, 1801: 59. 


● Lectotype (Gittenberger, 1973): NHMV No. 77707. 

● Distribution: East Transcaucasia, West Kopetdagh, Great Balkhan (Likharev, Schileyko, MS). 

Chondrina rhodia taurica (Kessler, 1860) T 

Pupa taurica Kessler, 1860: 226. 

● Type locality: vicinities of Yalta, Crimea. 


● Distribution: mountain Crimea (mostly southern slopes) (Likharev, Schileyko, MS). 

Granaria Held, 1837 

Type species: Pupa frumentum Draparnaud, 1801 (SD Herrmannsen, 1847) 

Granaria frumentum (Draparnaud, 1801) T 

[= Pupa frumentum var. minor Rossmässler, 1837; Pupa frumentum var. elongata Rossmässler, 1837] 

Pupa frumentum Draparnaud, 1801: 59. 

● Type locality: not stated (France-- from title). 

● Lectotype (Gittenberger, 1973): NHMV No. 77703. 

● Distribution: Ukraine: Carpathians, Ciscarpathia (Likharev, Schileyko, MS). 

PYRAMIDULIDAE Kennard et Woodward, 1914 

Pyramidula Fitzinger, 1833 

Type species: Helix rupestris Draparnaud, 1801 (monotypy) 

Pyramidula rupestris (Draparnaud, 1801) T 

[= Helix saxatilis Hartmann, 1821; Euryomphalia umbilicata Beck, 1837; Helix spinula Villa, 1841; Helix 

rupestris var. trochoides Moquin-Tandon, 1855; Pyramidula przevalskii Lindholm, 1922] 

Helix rupestris Draparnaud, 1801: 71-72. 



● Distribution: Volyno-Podolsk Hills, East Carpathians, mountain Crimea, Caucasus, Central Asia (Likharev, 

Schileyko, MS); Far East of Russia (Sikhote Alin) (Prozorova, Kavun, 2007a). 

ENIDAE Woodward, 1903 

PSEUDONAPAEINAE Schileyko, 1978 

Akramowskiella Schileyko, 1984 

Type species: Buliminus umbrosus Mousson, 1873 (OD) 

Akramowskiella andronakii (Lindholm, 1913) T 

Buliminus (Ena) andronakii Lindholm, 1913: 20. 

● Type locality: Artvin, northeastern Turkey. 


● Distribution: ? Adzharia (Likharev, Schileyko, MS).

Akramowskiella schuschaensis (Kobelt, 1902) T 

Buliminus (Ena) schuschaensis Kobelt, 1902: 727, pl. 107, figs. 9, 10. 

● Type locality: vicinities of Shusha, Nagornyi Karabakh. 

● Syntypes: SMF No. 10702 ("lectotype"), 10703/10 ("paralectotypes"). 

● Distribution: eastern part of Lesser Caucasus (Likharev, Schileyko, MS). 

Akramowskiella umbrosa (Mousson, 1873) T 

[= Buliminus talyschanus O. Boettger, 1880; Buliminus obscurus var. humberti Rosen, 1901; Buliminus hyrcanus 

Lindholm, 1915] 

Buliminus umbrosus Mousson, 1873: 205-206, pl. 8, fig. 1. 



● Distribution: North Caucasus and Transcaucasia (Likharev, Schileyko, MS). 

Clausilioides Lindholm, 1925 

Type species: Buliminus biplicatus Retowski, 1889 (OD) 

Clausilioides filifer (Lindholm, 1913) T 

Buliminus (Brephulus) filifer Lindholm, 1913: 21-22. 

● Type locality: Chorokh River debris, near Kapandidi village, ca. 7 km from Batumi (Adzharia, Georgia). 



Differena Schileyko, 1984 

Type species: Differena leucostoma Schileyko, 1984 (OD) 

Differena leucostoma Schileyko, 1984 T 

Schileyko, 1984: 316, figs. 225, 226. 

● Type locality: Kyapaz Mount, north part of Mrovdagh Ridge in eastern part of Lesser Caucasus. 



Geminula Lindholm, 1925 

Type species: Buliminus didymodus O. Boettger, 1880 (= Bulimus isseliana Bourguignat in Issel, 1865) (OD) 

Geminula continens (Rosen, 1892) T 

[= Chondrus natalensis Samadov, 1952] 

Buliminus (Amphiscopus) continens Rosen, 1892: 125-126. 

● Type locality: «Schamhala und Kasandshik» (northwestern Kopetdagh). 

● Syntypes: SMF No. 64121. 

● Distribution: Great Balkhan, Kopetdagh, Zeravshan, Nurat and Hissar ridges (Likharev, Schileyko, MS). 

Geminula isseliana (Bourguignat in Issel, 1865) T 

[= Bulimus ghilanensis Bourguignat in Issel, 1865; Chondrus nucifragus Mousson, 1873 non L. Pfeiffer, 

1848; Buliminus didymodus O. Boettger, 1880; Buliminus callilabris O. Boettger, 1889; Buliminus tardigyrus 


Bulimus isselianus Bourguignat – Issel, 1865: 421-422, pl. 2, figs. 37-40. 

● Type locality: «Lago Goktscha in Armenia». 


● Distribution: north slopes of Lesser Caucasus, to Khanlar in the west; middle reaches of Araks River valley 

to Dzhvezh in the west; mountain steppes of Talysh (Zuvand); Kopetdagh (Likharev, Schileyko, MS).

Imparietula Lindholm, 1925 

Type species: Bulimus leucodon L. Pfeiffer, 1846 (OD) 

Imparietula brevior (Mousson, 1876) T 

[= Chondrula brevior var. viator Westerlund, 1897] 

Buliminus brevior Mousson, 1876a: 34-35, pl. 2, fig. 5. 

● Type locality: upper reaches of Araks River. 


● Distribution: Armenia (near Lichk village and vicinities of Gyumri (former Leninakan)), ? northern slopes 

of Lesser Caucasus (Likharev, Schileyko, MS). 

Imparietula schelkovnikovi (Rosen, 1914) T 

[= Buliminus königi Retowski, 1914] 

Buliminus schelkovnikovi Rosen, 1914: 188, pl. 2, fig. 9. 

● Type locality: Boz-Dagh Mount near Evlakh. 


● Distribution: Boz-Dagh Mount and vicinities of Khudat (northwestern Great Caucasus) (Likharev, 


Laevozebrinus Lindholm, 1925 

Type species: Buliminus urgutensis Kobelt, 1902 (OD) 

Laevozebrinus eremita (Reeve, 1849) T 

[= Buliminus kuschakewitzi Ancey, 1886; Buliminus kuschakewitzi var. candisata Ancey, 1886; Buliminus 

bonvallotianus Ancey, 1886; Buliminus potaninianus Ancey, 1886; Buliminus eremita var. hepatica Ancey, 

1887; Buliminus eremita var. procera Ancey, 1887; Buliminus eremita var. germabensis O. Boettger, 1889; 

Buliminus (Petraeus) walteri O. Boettger, 1889; Buliminus kuschakewitzi var. funki Ancey, 1893; Buliminus 

eremita var. solivagus Westerlund, 1896; Buliminus bonvallotianus var. colorata Ancey, 1902] 

Bulimus eremita Reeve, 1849: pl. 78, sp. 573. 

● Type locality: Afghanistan, «on ... march from the Bolun Pass to Cabul». 


● Distribution: Kopetdagh, western Tajikistan, Ferghana Valley and adjacent ridges (Likharev, Schileyko, 

MS). 

Laevozebrinus guttula (Muratov, 1992) T 

Pseudonapaeus guttula Muratov, 1992: 39-42, fig. 2. 

● Type locality: southwestern Kopetdagh, between Sumbar and Chandyr rivers, 10 km south of Kara-Kala. 


● Distribution: Kopetdagh, basins of Sumbar and Chandyr rivers (Likharev, Schileyko, MS). 

Laevozebrinus lenis Schileyko, 1984 T 

Schileyko, 1984: 305, figs. 215 (III), 218. 

● Type locality: Karatau Ridge, 50 km northeast of Turkestan City. 


● Distribution: Karatau Ridge (Likharev, Schileyko, MS). 

Laevozebrinus ujfalvyanus (Ancey, 1886) T 

[= Subzebrinus prochorovi Lindholm, 1927) 

Buliminus ufjalvyanus [sic – lapsus calami] Ancey, 1886a: 38-40. 

● Type locality: «Khanat de Kokhand» (Ferghana Valley). 


● Distribution: Ferghana Valley, Ferghana, Chatkal and Alai ridges (Likharev, Schileyko, MS).

Laevozebrinus urgutensis (Kobelt, 1902) T 

Buliminus ? urgutensis Kobelt, 1902: 21, pl. 248, fig. 1606. 

● Type locality: Urgut, Samarkand Region. 

● Lectotype (Zilch, 1959-1960): SMF No. 150811. 

● Distribution: Urgut, Samarkand Region (Likharev, Schileyko, MS). 

Ljudmilena Schileyko, 1984 

Type species: Chondrus sieversi Mousson, 1873 (OD) 

Ljudmilena sieversi (Mousson, 1873) T 

[= Buliminus hoplites Westerlund, 1890; Buliminus adjaricus Retowski, 1914; Buliminus araxena Lindholm, 

1923; Buliminus acampsica Lindholm, 1923] 

Chondrus sieversi Mousson, 1873: 207-208, pl. 7, fig. 6. 

● Type locality: debris of Araks River. 


● Distribution: Adzharia, South Armenia, Nakhichevan (Likharev, Schileyko, MS). 

Ljudmilena tricollis (Mousson, 1876) T 

[= Buliminus tricollis var. excellens Retowski, 1889; Buliminus armeniacus Ancey, 1893 non Mortillet, 1854; 

Chondrula excellens var. duodecimgyrata Lindholm, 1923; Chondrula carseana Lindholm, 1923] 

Chondrus tricollis Mousson, 1876b: 141, pl. 5, fig. 2. 

● Type locality: «Azhuz» (Atzkuri), Kura River valley. 


● Distribution: local records in southern Transcaucasia except its eastern areas (Likharev, Schileyko, MS). 

Mastoides Westerlund, 1896 

Type species: Buliminus albocostatus Westerlund, 1896 (monotypy) 

Mastoides albocostatus (Westerlund, 1896) T 

[= Buliminus albocostatus var. distortus Westerlund, 1896] 

Buliminus (Mastoides) albocostatus Westerlund, 1896: 192. 

● Type locality: Arslanbob, Ferghana Ridge. 


● Distribution: vicinities of Arslanbob, Ferghana Ridge (Likharev, Schileyko, MS). 

Mastoides obeliscus Schileyko, 2007 T 

Schileyko, 2007: 85-87, fig. 2. 

● Type locality: Dzhaz-ichu River valley, Kara-Unghur River basin, Ferghana Range. 



Mastoides orloffensis (Kobelt, 1905) T 

[= Buliminus albocostatus Ancey, 1902 non Westerlund, 1896] 

Buliminus (Mastoides) albocostatus orloffensis Kobelt, 1905: 19, fig. 1972 (nom. nov. pro Buliminus albocostatus 

Ancey, 1902 non Westerlund, 1896). 

● Type locality: Karagoin ravine, Talas River valley. 

● Syntypes: SMF No. 238552 (including “lectotype”); possible syntypes ZMMU Nos. Lc-21493, Lc-14565. 

● Distribution: Talas River valley (Likharev, Schileyko, MS). 

Ottorosenia Muratov, 1992 

Type species: Buliminus (Subzebrinus) varenzovi Rosen, 1893 (OD)

Ottorosenia varenzovi (Rosen, 1893) T 

[= ?Buliminus miser var. misellus Westerlund, 1896] 

Buliminus (Subzebrinus) varenzovi Rosen, 1893: 179. 

● Type locality: Kopetdagh, Gaudan mountain pass. 

● Lectotype (Muratov, 1992): ZIN. 

● Distribution: Kopetdagh and (?) Turkestan Ridge (Likharev, Schileyko, MS). 

Pseudochondrula Hesse, 1933 

Type species: Buliminus florenskii Rosen, 1914 (= Pupa seductilis Rossmässler, 1837) (OD) 

Pseudochondrula lederi (O.Boettger, 1883) T 

[= Buliminus lederi var. limis Westerlund, 1887] 

Buliminus lederi Boettger, 1883: 177, pl. 7, fig. 1. 

● Type locality: vicinities of Esheri village, north of Sukhumi, Svanetia, Georgia. 


● Distribution: locally on Great Caucasus, except for eastern part, and northern slopes of Lesser Caucasus 

(Bakuriani) (Likharev, Schileyko, MS). 

Pseudochondrula seductilis (Rossmässler, 1837) T 

[= Buliminus seductilis var. armeniaca Mortillet, 1854; Chondrus scapus var. destitutus Mousson, 1873; 

Chondrus sagax var. destituta Mousson, 1873; Buliminus komarowi O. Boettger, 1880; Buliminus (Chondrula) 

incertus Retowski, 1883; Buliminus florenskii Rosen, 1914; Buliminus (Chondrula) lindholmi Retowski, 

1915] 

Pupa seductilis Rossmässler, 1837: 10, pl. 23, figs. 306, 307. 

● Type locality: Dalmatia. 

● Syntypes: SMF Nos. 237088 ("lectotype"), 237089/1 ("paralectotype"). 

● Distribution: southwestern Georgia (Batumi) and Armenia (Likharev, Schileyko, MS). 

Pseudochondrula sinistrorsa Kokotschashvili et Schileyko in Schileyko, 1984 T 

Schileyko, 1984: 294, figs. 205 (V), 208 (IV). 

● Type locality: western Georgia, southwestern Suram Ridge, Lashe village. 



Pseudochondrula tetrodon (Mortillet, 1854) T 

[= Buliminus (Chondrula) diffusus Mousson, 1876; Buliminus (Chondrula) kollyi Retowski, 1889] 

Buliminus tetrodon Mortillet, 1854: 11, pl. 1, fig. 3. 

● Type locality: Ispir, vilayet Erzurum, Turkey. 


● Distribution: Georgia (Tbilisi, Kutaisi, Borzhomi) and southern Armenia (Likharev, Schileyko, MS). 

Pseudochondrula tuberifera (O. Boettger, 1879) T 

[= Buliminus lederi var. limis Westerlund, 1887] 

Buliminus (Chondrula) tuberifer Boettger, 1879a: 22-23, pl. 1, fig. 9. 

● Type locality: Kazbek and Kobi. 

● Syntype: ZMB No. 31251 

● Distribution: Kazbek, vicinities of Kobi, Suram Ridge, North Osetia (Zrug ravine), western Great Caucasus 

(Pregradnaya settlement on Urup River) (Likharev, Schileyko, MS). 

Pseudonapaeus Westerlund, 1887 

Type species: Buliminus asiaticus Martens, 1881 (SD Lindholm, 1922)

Pseudonapaeus albiplicatus (Martens, 1874) T 

[= Buliminus przevalskii Ancey, 1886; Buliminus ferghanensis Kobelt, 1890; Buliminus retteri Rosen, 1897; 

Buliminus albiplicatus f. implicata Westerlund, 1898; Subzebrinus flammulatus Lindholm, 1927] 

Buliminus albiplicatus Martens, 1874: 20, pl. 2, fig. 15. 

● Type locality: Tashkent. 


● Distribution: entire Tien-Shan (Likharev, Schileyko, MS). 

Pseudonapaeus aptychus (Ancey, 1886) T 

[= Buliminus aptychus var. capusiana Ancey, 1886] 

Buliminus aptychus Ancey, 1886b: 334. 

● Type locality: «Wjernoje (Semiretschinsk)» (Alma-Ata, Kazakhstan). 


● Distribution: Zailijskij Ridge (Kazakhstan) (Likharev, Schileyko, MS). 

Pseudonapaeus asiaticus (Martens, 1881) T 

[= Buliminus vamberyi Ancey, 1886; Buliminus komarovi Kobelt, 1890 non O. Boettger, 1880; Buliminus 

turanicus Ancey, 1893] 

Buliminus asiaticus Martens, 1881 (1880-1881): 29, pl. 6, figs. 12-14. 



● Distribution: northeastern and eastern Central Asia and East Kazakhstan (Likharev, Schileyko, MS). 

Pseudonapaeus bacillus Kuznetsov, 1999 T 

Kuznetsov, 1999: 108-109, fig. 5. 

● Type locality: Kazakhstan, Alma-Ata Region, Kaskelen District, Zailijskij Alatau Ridge, left side of Aksai 

River valley, 300 m upstream from pioneer camp «Leninetz», lower part of eastern slope above earth road. 


● Distribution: northern slope of Zailijskij Alatau Ridge (northern Tien-Shan) (Kuznetsov, 1999). 

Pseudonapaeus chodschendicus (Mukhitdinov, 1976) T 

Chondrulopsina chodschendicus Mukhitdinov, 1976: 39, fig. 1A, 2A. 

● Type locality: Kuraminskij Ridge, vicinities of Asht village. 


● Distribution: Kuraminskij Ridge (Likharev, Schileyko, MS). 

Pseudonapaeus diplus (Westerlund, 1896) T 

Buliminus (Pseudopetraeus) diplus Westerlund, 1896: 191. 

● Type locality: Ferghana Ridge, Arslanbob. 

● Lectotype (Schileyko, 1984): NMG No. 2018. 

● Distribution: Ferghana Ridge (Arslanbob and Kyzyl-Ungur) (Likharev, Schileyko, MS). 

Pseudonapaeus dissimilis (Martens, 1882) T 

[= Buliminus ferghanensis Kobelt, 1890; Buliminus retteri Rosen, 1897; Buliminus albiplicatus f. implicata 

Westerlund, 1898; Subzebrinus flammulatus Lindholm, 1927] 

Buliminus dissimilis Martens, 1882b: 106. 

● Type locality: Arasan-Bulak (Alma-Ata Region, Kazakhstan). 


● Distribution: foothills of Zailijskij, Kungei, Terskei ridges, upper reaches of Naryn River (Likharev, 


Pseudonapaeus drymaeus (Westerlund, 1896) T 

Buliminus (Chondrulopsis) drymaeus Westerlund, 1896: 194.

● Type locality: Ferghana Ridge, Arslanbob. 


● Distribution: western Ferghana Ridge (Likharev, Schileyko, MS). 

Pseudonapaeus eleonorae (Tzvetkov, 1940) T 

Ena (Subzebrinus) eleonorae Tzvetkov, 1940a: 386-387. 

● Type locality: Kazakhstan, Alma-Ata Region, Dzhambul District, Kastek Ridge, Uzun-Agach River gorge, 

Alayak pasture. 

● Lectotype (Kuznetsov, 1999): ZMMU No. Lc-24068. 

● Distribution: western parts of Kyungei Ala-Too and Kastek ridges, eastern Kirgiz Ala-Too Ridge (northern 

Tien-Shan) (Kuznetsov, 1999). 

Pseudonapaeus entodon (Martens, 1882) T 

[= Buliminus entodon var. dextroversa Ancey, 1886] 

Buliminus entodon Martens, 1882b: 106. 

● Type locality: near Vernyi City (= Alma-Ata), foothills of Zailijiskij Alatau Ridge. 


● Distribution: northern slopes of Zailijskij, Kastek, Kirgiz Ala-Too ridges (northern Tien-Shan) (Kuznetsov, 

1999). 

Pseudonapaeus entoptyx (Lindholm, 1925) T 

Buliminus entoptyx Lindholm, 1925a: 32. 

● Type locality: Kara-Su, Ferghana Ridge. 


● Distribution: Ferghana, western Talas (Aksu-Djabagly natural reserve), Pskem and Ugam ridges (Likharev, 


Pseudonapaeus errans (Westerlund, 1896) T 

[= Buliminus costatus Clessin, 1894; Buliminus merzbacheri Weber, 1913] 

Buliminus (Mastoides) errans Westerlund, 1896: 193. 

● Type locality: along Kugart River, near Taranbazar. 


● Distribution: Ferghana Ridge: near Taranbazar and Nara-Alma River (tributary of Kugart [= Koek-Art] 

River) valley (Likharev, Schileyko, MS). 

Pseudonapaeus galinae (Tzvetkov, 1940) T 

Ena galinae Tzvetkov, 1940a: 386. 

● Type locality: foothills of Zailijskij Alatau Ridge near Alma-Ata, slope of river near State farm of NKVD 

(Kazakhstan). 

● Lectotype (Schileyko, 1984): ZMMU No. Lc-3773. 

● Distribution: foothills of Zailijskij Ridge (Kazakhstan) (Likharev, Schileyko, MS). 

● Remark: Izzatullaev (1991) recorded a Chinese species, Buliminus saccatus Moellendorff, 1901, from Terskei 

Ridge. However, the description and figure much better correspond to P. galinae than to P. saccatus, and 

the described species is most probably a form of variability of P. galinae. 

Pseudonapaeus goldfussi (Kobelt, 1893) T 

Buliminus (Chondrula?) goldfussi Kobelt, 1893: 82, pl. 171, fig. 1102. 

● Type locality: Alai Ridge. 


● Distribution: Ferghana and Alai ridges (Likharev, Schileyko, MS). 

Pseudonapaeus herzensteini (Ancey, 1886) T 

[= Buliminus herzensteini var. strophostoma Ancey, 1886; Buliminus herzensteini var. pellucens Ancey,

1886] 

Buliminus herzensteini Ancey, 1886a: 270 (May); 1886b: 23-24 (July). 

● Type locality: not given (l’Asie centrale Russe – from the title). 

● Syntypes: 2 syntypes NMWZ NMW.1955.158.24106 (illustrated by Wood, Gallichan, 2008, pl. 7, fig. 5, 

vi). 

● Remark: Ancey (1886b) listed more specific type locality of ‘Partie orientale du Turkestan russe’ at the end 

of the section on Buliminus herzensteini, but it is after a discussion on the variety pellucens so it is unclear 

if this applies to the species at all (Wood, Callichan, 2008). 

Pseudonapaeus izzatullaevi Kuznetsov, 1999 T 

Kuznetsov, 1999: 102-103, fig. 1 A, B, C. 

● Type locality: Uzbekistan, Surkhan-Darya Region, southern slope of Hissar Ridge, Khopzada River gorge. 


● Distribution: type locality (Kuznetsov, 1999). 

Pseudonapaeus kasnakowi (Westerlund, 1898) T 

Buliminus (Brephulus) kasnakowi Westerlund, 1898: 163. 

● Type locality: Vakhsh valley. 


● Distribution: Hissaro-Darvaz ridge system (Likharev, Schileyko, MS). 

Pseudonapaeus latilabris (Lindholm, 1927) T 

Ena (Pseudonapaeus) latilabris Lindholm, 1927c: 101. 

● Type locality: Type locality; "Ferghana-Gebiet, am Oberlauf des Fl. Turduk" [Fergana district, 

upper part of Turduk River]. 


● Distribution; Chatkal and Ferghana ridges (Kirgizia). 

Pseudonapaeus leucopleurus (Lindholm, 1927) T 

?Subzebrinus leucopleurus Lindholm, 1927d: 258. 

● Type locality: Chatkal Ridge, vicinities of Sary-Chelek Lake. 

● Syntypes: ZIN, ZMMU No. Lc-15132. 


Pseudonapaeus leucoptychus (Ancey, 1886) T 

[= Buliminus costatus Clessin, 1894; Buliminus merzbacheri Weber, 1913] 

Buliminus leucoptychus Ancey, 1886b: 336-337. 

● Type locality: «fleuves Fekkes et Naryn-Kol» (rivers Tekes and Narynkol). 

● Syntypes: ZIN, RBINS No. 10591. 

● Distribution: Chu River valley, vicinities of Issyk-Kul Lake, East Kirgizia (Likharev, Schileyko, MS). 

Pseudonapaeus lindholmi Kuznetsov, 1999 T 


● Type locality: Kirgizia, Chu Region, Alamedi District, Kirgiz Alo-Too Ridge, northwest of Chon-Kurchak 

village, 200 m northwest of automobile bridge, floodland of left bank of Chon-Kurchak River. 


● Distribution: Kirgiz Ala-Too, Zailijskij Alatau, Kyungei Ala-Too ridges (northern Tien-Shan) (Kuznetsov, 

1999). 

Pseudonapaeus miser (Martens, 1874) T 

Buliminus miser Martens, 1874: 21-22, pl. 2, fig. 17. 

● Type locality: Auchi-Dagana valley on the road from Ura-Tyube to mountain pass leading to Zaravshan

valley (northern foothills of Turkestan Ridge). 


● Distribution: Turkestan, Hissar, Alai, Ferghana, Chatkal, Kirgiz, Zailijskij ridges; Tarbagatai, western Altai 


Pseudonapaeus naykatikus Pazylov in Pazylov, Kuchbaev et Daminova, 2001 T 

Pazylov et al., 2001: 57-58, fig. 2. 

● Type locality: Atai Ridge, Karabulak Ravine, near Naukata settlement. 

● Holotype: Gulistan State University. 


Pseudonapaeus otostomus (Westerlund, 1898) T 

Buliminus (Brephulus) otostomus Westerlund, 1898: 164. 

● Type locality: Totkul, Vakhsh valley. 


● Distribution: Hissaro-Darvaz ridge system (Likharev, Schileyko, MS). 

Pseudonapaeus regelianus (Ancey, 1886) T 

[= Buliminus cylindroconus Ancey, 1886] 

Buliminus asiaticus var. regeliana Ancey, 1886a: 26-27. 

● Type locality: «Wjernoje, dans la Songarie» (Alma-Ata, Kazakhstan). 

● Possible syntype: RBINS.10591. 

● Distribution: Zailijskij and Dzungar ridges and south and west of Issyk-Kul Lake (Likharev, Schileyko, 

MS). 

Pseudonapaeus retrodens (Martens, 1879) T 

[= Buliminus retroplicatus Westerlund, 1894] 

Buliminus retrodens Martens, 1879: 126. 



● Distribution: Zailijskij, Kungei, Terskei, Djetim-Bel, Naryn, At-Bashi ridges, north-eastern Ferghana Ridge 


Pseudonapaeus schileykoi Izzatullaev, 1991 T 

Izzatullaev, 1991: 138, fig. 

● Type locality: Karateginskij Ridge, Komarou ravine, vicinities of Rvoz village. 


● Distribution: Karateginskij, Darvazskij, Peter the First ridges (Likharev, Schileyko, MS). 

Pseudonapaeus schnitnikovi (Lindholm, 1922) T 

Buliminus (Pseudopetraeus) schnitnikovi Lindholm, 1922b: 274. 

● Type locality: vicinities of Alma-Ata (Kazakhstan). 


● Distribution: Zailijskij Ridge, near Alma-Ata (Kazakhstan) (Likharev, Schileyko, MS). 

Pseudonapaeus secalinus (Martens, 1881) T 

[= Buliminus semenovi Ancey, 1893; Buliminus biformis Westerlund, 1896; Buliminus torquatus Westerlund, 

1896; Ena secalina var. maibulaki Skvortzov, 1940] 

Buliminus secalinus Martens, 1881 (1880-1881): 27, pl. 6, figs. 8, 9. 



● Distribution: Tien-Shan: Zailijskij, Kirgiz, Kungei, Terskei, ?Ferghana, Alai ridges (Likharev, Schileyko, 

MS).

Pseudonapaeus sogdianus (Martens, 1874) T 

[= Buliminus oxianus Martens, 1876; Buliminus oxianus var. brevior Martens, 1876; Buliminus labiellus var. 

kokandensis Martens, 1882; Buliminus larvatus Ancey, 1902; Buliminus korshinskii Lindholm, 1922] 

Buliminus sogdianus Martens, 1874: 19, pl. 2, fig. 14a-c. 

● Type locality: upper reaches of Zeravshan valley, Kuli-Kalan. 


● Distribution: mountain areas of Turkmenia, Uzbekistan, western Kirgizia (Likharev, Schileyko, MS). 

Pseudonapaeus stabilis stabilis Schileyko, 1984 T 

Schileyko, 1984: 262, figs. 172, 173. 

● Type locality: West Tien-Shan, Pskem River valley, 15 km upstream of Nanai settlement. 


● Distribution: West Tien-Shan, Pskem and Ugam ridges, Pskem River valley and lower reaches of Ugam 

River (Kuznetsov, 1999). 

Pseudonapaeus stabilis chatkalicus Kuznetsov, 1999 T 

Kuznetsov, 1999: 110, fig. 6 A, B, C. 

● Type locality: Uzbekistan, Tashkent Region, Bostanlyk District, Keksui Ridge, 5.5 km south of Burchmulla 

settlement, right side of Chatkal River valley, lower third of western slope of Paltau Mount, 50 m above road, 

1200 m above sea level. 


● Distribution: Keksui Ridge (West Tien-Shan), lower reaches of Chatkal Ugam River (Kuznetsov, 1999). 

Pseudonapaeus submucronatus (Lindholm, 1927) T 

Ena (Pseudonapaeus) submucronatus Lindholm, 1927c: 100. 

● Type locality: Chatkal Ridge, middle reaches of Uzun-Akhmat River, 1200-1400 m above sea level. 


● Distribution: two localities in the Naryn River Basin: along one of right tributaries of the Naryn in Chatkal 

Ridge and in foothils of Ferghana Ridge, along the same river (Likharev, Schileyko, MS). 

Pseudonapaeus subobscurus (Ancey, 1886) T 

Buliminus subobscurus Ancey, 1886b: 330-331. 

● Type locality: Daraty-Bulak, «Fekkès» (Tekes) River valley. 

● Syntypes: 1 syntype NMWZ NMW.1955.158.24112 (illustrated by Wood, Gallichan, 2008, pl. 8, fig. 2, ii), 

1 syntype RBINS.105891. 

● Distribution: East Kirgizia (Sary-Djaz Ridge, Tekes River valley) (Likharev, Schileyko, MS). 

Pseudonapaeus trigonochilus (Ancey, 1886) T 

[= Buliminus trigonochilus f. expansilabris Westerlund, 1896; Buliminus trigonochilus f. planicollis Westerlund, 

1896; Buliminus trigonochilus f. pachychyla Westerlund, 1896; Buliminus castaneus Westerlund, 1897; 

Buliminus maracandensis Rosen, 1901; Buliminus latilabris Lindholm, 1927] 

Buliminus trigonochilus Ancey, 1886a: 270 (May); 1886b: 28-29 (July). 

● Type locality: not given, Samarkand, Turkestan in later reference (Ancey, 1902). 

● Syntypes: 2 syntypes NMWZ NMW.1955.158.24170 (illustrated by Wood, Gallichan, 2008, pl. 7, fig. 6, 

vii). 

● Distribution: southern slopes of Chatkal, Ferghana and Alai ridges, vicinities of Samarkand (Likharev, 


Pseudonapaeus zeravshanicus Pazylov et Daminova in Pazylov, Kuchbaev et Daminova, 2001 T 


● Type locality: Aeravshan Ridge, vicinities of Bashir village. 

● Holotype: Gulistan State University.


Subzebrinus Westerlund, 1887 

Type species: Buliminus labiellus Martens, 1881 (SD Moellendorff, 1901) 

Subzebrinus labiellus (Martens, 1881) T 

[= Buliminus labiellus var. strophostoma Ancey, 1886; Buliminus pullaster Ancey, 1886] 

Buliminus labiellus Martens, 1881 (1880-1881): 24, pl. 6, figs. 1, 2. 

● Type locality: Tarbagatai (Dzungar), «zwischen den Seen Balchasch und Dsaisan». 


● Distribution: Zailijskij, Dzungar, Tarbagarai ridges (Likharev, Schileyko, MS). 

Turanena Lindholm, 1922 

Type species: Buliminus herzi O. Boettger, 1889 (OD) 

Turanena albolimbata (Lindholm, 1927) T 

Ena (Turanena) albolimbata Lindholm, 1927c: 98. 

● Type locality: Chatkal Ridge, upper reaches of Turduk River (Naryn River Basin). 


● Distribution: Chatkal, ? Talas, Kirgiz ridges (Likharev, Schileyko, MS). 

Turanena boamica Kuznetsov, 1999 T 


● Type locality: Kirgizia, Chu Region, Kemin District, Kirgiz Ala-Too Ridge, Oktorkoi Mountains, left side 

of Chu River gorge (= Boam gorge), southwestern part of Kyz-Kuioo village, middle of east-south-eastern 

slope of mount, small ravine. 



Turanena cognata (Lindholm, 1927) T 

Ena (Turanena) cognata Lindholm, 1927c: 99. 

● Type locality: Chatkal Ridge, lower reaches of Turduk River. 


● Distribution: Chatkal, ? Talas, Kirgiz ridges (Likharev, Schileyko, MS). 

Turanena conicula (Martens, 1882) T 

Buliminus coniculus Martens, 1882a: 23-24, pl. 3, fig. 3. 



● Distribution: Alai (Chauvai village) and Zeravshan (near Marguzor Lake) ridges (Likharev, Schileyko, 

MS). 

Turanena herzi (O. Boettger, 1889) T 

Buliminus (Pseudonapaeus) herzi Boettger, 1889: 950, pl. 17, fig. 14. 

● Type locality: Iran, Elburs, vicinities of Shakhrud. 


● Distribution: ? mountains of southwestern Turkmenia (Likharev, Schileyko, MS). 

Turanena inversa Schileyko et Moisseeva, 1995 T 

Schileyko, Moisseeva, 1995: 45-46, figs. 1, 2. 

● Type locality: Tien-Shan, Chatkal Range, upper part of the valley of the Aflatun River (left tributary of the 

Chatkal River). 

● Holotype: ZMMU No. Lc-22029.


Turanena leptogyra (Lindholm, 1927) T 

Ena (Turanena) leptogyra Lindholm, 1927c: 99. 

● Type locality: Chatkal Ridge, Balgut village between Mart and Kumbel mountain passages. 


● Distribution: Alai, Chatkal, Pskem, ? Ugam, Talas ridges (Likharev, Schileyko, MS). 

Turanena margaritae Schileyko et Moisseeva, 1989 T 

Schileyko, Moisseeva, 1989: 139, fig. 1. 

● Type locality: Tien-Shan, Kichik-Alai Ridge, Kirgiz-Ata Canyon, bank of Mazar-Sai River upstream from 

Eski-Naukat settlement. 



Turanena martensiana (Ancey, 1886) T 

[= Buliminus liostracus Ancey, 1886] 

Buliminus martensiana Ancey, 1886a: 45-46. 

● Type locality: mountains near Shakhimardan, foothills of Altai. 


● Distribution: Turkestan, Zeravshan, Hissar ridges through Alai, Zaalai, Ferghana ridges to southwestern 

slopes of Kungei Ridge (Likharev, Schileyko, MS). 

Turanena meshkovi Schileyko, 1984 T 

Schileyko, 1984: 279, fig. 186 (VII, VIII). 

● Type locality: West Tien-Shan, Pskem River valley, 15 km upstream of Nanai village. 


● Distribution: West Tien-Shan, Pskem River valley (Likharev, Schileyko, MS). 

Turanena scalaris (Naegele, 1902) T 

Buliminus (Pseudonapaeus) scalaris Naegele, 1902: 6-7. 

● Type locality: «in Monte Razoki, Urmia» (northwestern Iran). 


● Distribution: South Armenia (Gnishik village, Dzafarigyukh area) (Likharev, Schileyko, MS). 

Turanena stschukini (Lindholm, 1927) T 

Ena (Turanena) stschukini Lindholm, 1927c: 100. 

● Type locality: Dzuvan-Syuget River valley, right tributary of Uzun-Akhmat River (Naryn River Basin). 


● Distribution: Chatkal and Ferghana ridges (Likharev, Schileyko, MS). 

Turanena tenuispira Schileyko, 1984 T 

Schileyko, 1984: 282, figs. 186 (V, VI), 196, 197. 

● Type locality: Kara-Kostek River valley near Alma-Ata. 


● Distribution: Zailijskij and Ferghana ridges (Likharev, Schileyko, MS). 

CHONDRULOPSININAE Schileyko, 1978 

Chondrulopsina Lindholm, 1925 

Type species: Buliminus haberhaueri Ancey, 1886 (= Buliminus intumescens var. fedtschenkoi Ancey, 1886) 

(OD)

Chondrulopsina fedtschenkoi (Ancey, 1886) T 

[= Buliminus intumescens var. inermis Ancey, 1893; Buliminus anomphalus Westerlund, 1897; Buliminus 

bisinuatus Westerlund, 1898; Buliminus rennenkampfi Rosen, 1901; Chondrulopsina trisinuata Mukhitdinov, 

1976] 

Buliminus intumescens var. fedtschenkoi Ancey, 1886a: 51. 

● Type locality: «Samarkand, dans la vallée du Sarafschan». 

● Syntype: NMWZ NMW.1955.158.01676 (illustrated by Wood, Gallichan, 2008, pl. 8, fig. 3, iii). 

● Distribution: mountain and submountain areas of Samarkand Region: Ferghana, Alai, Kuramin, Turkestan, 

Zaalai, Hissar ridges (Likharev, Schileyko, MS). 

Chondrulopsina intumescens (Martens, 1874) T 

[= Buliminus annenkovi Ancey, 1893; Buliminus (Chondrula) anomphalus f. infans Westerlund, 1897; Buliminus 

intumescens var. sultanus Rosen, 1901; Subzebrinus likharevi Mukhitdinov, 1976] 

Buliminus (Chondrula) intumescens Martens, 1874: 22, pl. 2, fig. 18. 

● Type locality: Chupanata near Samarkand. 

● Holotype: ZMB No. 22459. 

● Distribution: mountain regions of Central Asia except for northern Tien-Shan and Dzungar Ridge (Likharev, 


Siraphoroides Schileyko, 1977 

Type species: Siraphorus moltschanovi Likharev et Rammelmeyer, 1952 (OD) 

Siraphoroides moltschanovi (Likharev et Rammelmeyer, 1952) T 

Siraphorus moltschanovi Likharev, Rammelmeyer, 1952: 207, figs. 121, 122. 

● Type locality: Arslanbob, Ferghana Ridge. 


● Distribution: Ferghana Ridge (Arslanbob), Yangyskyr Mountains near eastern Ferghana Ridge, upper 

reaches of Pchan River (right tributary of Ala-Buka, Naryn River Basin), Naryn River valley in 10 km of At- 

Bashi hydroelectric power plant (Likharev, Schileyko, MS). 

MERDIGERINAE Schileyko, 1984 

Merdigera Held, 1837 

Type species: Helix obscura Müller, 1774 (SD Herrmannsen, 1847) 

Merdigera invisa Kijashko, 2006 T 

Kijashko, 2006: 89–91, fig. 1. 

● Type locality: Northwest Caucasus, Adygeja, Lagonaki Mountains, southern foot of Lagonaki 

Ridge, right riverbank of Tsitse (right tributary of the river Psheha), (~ 1200 m above sea level), 

lime rocks 

● Holotype: ZIN 1/527–2006 

● Distribution: type locality (Kijashko, 2006). 

Merdigera obscura (Müller, 1774) T 

Helix obscura Müller, 1774: 103. 



● Distribution: Baltic countries, western Ukraine, Byelorussia (Vitebsk Region), Russia (Tver, Smolensk, 

Yaroslavl regions, vicinities of Moscow and St.-Petersburg), mountain Crimea, northwestern Caucasus, Uzbekistan 

(West Tien-Shan), Kazakhstan (Dzhungaria) (Kuznetsov, 1999).

BULIMINUSINAE Kobelt, 1880 

Adzharia Hesse, 1933 

Type species: Adzharia renschi Hesse, 1933 (monotypy) 

Adzharia renschi Hesse, 1933 T 

Hesse, 1933: 158, fig. 4. 

● Type locality: «Chula» (Khulo) mountain pass, Adzharo-Imereti Ridge. 



Buliminus Beck, 1837 

Type species: Buliminus labrosus Olivier, 1804 (SD Gray, 1847) 

Buliminus urmianus O. Boettger, 1898 T 

[= Buliminus valentini Kobelt, 1902] 

Buliminus (Petraeus) halepensis var. urmiana Boettger, 1898: 26-27. 

● Type locality: «Urmia in Kurdistan, Nordwest-Persien» (vicinities of Rezaiyeh (Urmia) City, northwestern 

Iran). 

● Syntypes: "Lectotype" SMF No. 14506, "paralectotypes" SMF No. 14507/15, 150761/13. 

● Distribution: Nakhichevan area (mouth of Alindzha-chai River, 5 km west of Dzhulfa), Azerbaijan (Zangilan 

Region) (Likharev, Schileyko, MS). 

ANDRONAKIINAE Schileyko, 1998 

Andronakia Lindholm, 1914 

Type species: Chondrula (?) catenulata Lindholm, 1913 (monotypy) 

Andronakia catenulata (Lindholm, 1913) T 

Chondrula (?) catenulata Lindholm, 1913: 22-23. 

● Type locality: «Ortschaft Wasrija am Pass nach Kwarzschana (Gouvernement Batum» (northwestern Turkey). 


● Distribution: Adzharia: Mtirala Mount in 28-30 km east of Batumi and Khelvachauri district near Chirnali 

village in Chirnali River valley (Likharev, Schileyko, MS). 

RETOWSKIINAE Schileyko, 1978 

Retowskia O. Boettger, 1881 

Type species: Chondrus schlaeflii Mousson, 1863 (monotypy) 

Retowskia schlaeflii (Mousson, 1863) T 

[= Buliminus schlaeflii var. ingens O. Boettger, 1888] 

Chondrus schlaeflii Mousson, 1863: 390. 

● Type locality: «Rédutkaleh» (vicinities of Poti). 


● Distribution: mountain areas of western Great Caucasus (including upper reaches of Belaya River in the 

east) and Transcaucasia (including Suram Ridge in the east) (Likharev, Schileyko, MS).

ENINAE Woodward, 1903 

Brephulopsis Lindholm, 1925 

Type species: Chondrus attenuatus Krynicki, 1833 (= Bulimus bidens Krynicki, 1833) (OD) 

Brephulopsis bidens (Krynicki, 1833) T 

[= Chondrus attenuatus Krynicki, 1833; Bulimus theodosianus Bourguignat, 1876; Buliminus retowskianus 

Clessin, 1881; Buliminus (Brephulus) bidens natio turriformis Puzanov, 1925; Buliminus (Brephulus) bidens 

natio pygmaea Puzanov, 1925; Buliminus (Brephulus) bidens natio Cimmerii Puzanov, 1926; Buliminus (Brephulus) 

bidens natio gracilis Puzanov, 1926; Buliminus (Brephulus) bidens natio costulatus Puzanov, 1926] 

Chondrus bidens Krynicki, 1833: 401-402, pl. 3, fig. 3a-e. 

● Type locality: Simferopol (Crimea). 


● Distribution: Crimea (mountains and steppe) (Likharev, Schileyko, MS). 

Brephulopsis cylindrica (Menke, 1828) T 

[= Chondrus lineatus Krynicki, 1833; Chondrus turgidus Krynicki, 1833; Chondrus fuscilabris Krynicki, 

1833; Chondrus concolor Krynicki, 1833; Bulimus unicolor Andrzejowski in Krynicki, 1833; ?Bulimus turritella 

Andrzejowski in Krynicki, 1833; Bulimus illibatus Rossmässler, 1837; Bulimus tauricus L. Pfeiffer, 

1848; Chondrus duboisi Mousson, 1863; Bulimus subacuminatus Retowski, 1888; Buliminus cylindricus inflatus 

Pusanov, 1925; Buliminus [Zebrina) cylindricus inflatus Pusanov, 1925; Buliminus (Zebrind) cylindricus 

minor Puzanov, 1925; Buliminus (Zebrina) cylindricus natio cupressiformis Puzanov, 1925] 

Bulimus cylindrica Menke, 1828: 77. 



● Distribution: Crimea (mountains and steppe); Odessa, Nikolaev, Kherson, Zaporo-zhje, Dnepropetrovsk, 

Donetzk regions of Ukraine; probably introduced records from Moldova, Kiev, Lvov, Novorossiisk, Anapa 

and Sukhumi (Likharev, Schileyko, MS; Sverlova, 2006). 

Caucasicola Hesse, 1917 

Type species: Buliminus raddei Kobelt, 1880 (monotypy) 

Caucasicola raddei (Kobelt, 1880) T 

Buliminus raddei Kobelt, 1880: 50, figs. 2008, 2009 

● Type locality: «? Avarien» (mountain Daghestan). 


● Distribution: horseshoe-shaped range starts in North Caucasus, approximately at Maikop (one record near 

Nalchik), passing westward to Novorossiisk and Gelendzhik and including western regions of southern slopes 

of Great Caucasus, to the Kodor River in the east. As for East Caucasus, Kobelt himself was not certain about 

the reliability of the type locality, and there are no reliable records of the species from Daghestan (Likharev, 


Chondrula Beck, 1837 

Type species: Helix tridens Müller, 1774 (SD Martens in Albers, 1860) 

Chondrula bielzi (Kimakowicz, 1890) T 

Dentistomus (Amphitrorsus) bielzi Kimakowicz, 1890: 222. 

● Type locality: Transylvania. 


● Distribution: Ukraine (Lesistye Carpathians) (Likharev, Schileyko, MS).

Chondrula caucasica (L. Pfeiffer, 1852) T 

[= Buliminus ponticus Retowski, 1887] 

Bulimus caucasica L. Pfeiffer, 1852: 94. 

● Type locality: «in Caucaso». 


● Distribution: Novyi Afon (Abkhazia), Crimea between Feodosiya and Sudak (Likharev, Schileyko, MS). 

Chondrula microtraga (Parreyss in Rossmässler, 1839) T 

Pupa microtragus Parreyss – Rossmässler, 1839: 30-31, pl. 49, fig. 651. 

● Type locality: Greece. 


● Distribution: the species could be found in Adzharia near border of Turkey and near the border of Romania 

(Likharev, Schileyko, MS); one probably introduced population in Odessa (Sverlova, 2006). 

Chondrula sunzhica Steklov, 1962 T 

Chondrula microtraga sunzhica Steklov, 1962: 141, pl. 2, figs. 3, 4. 

● Type locality: Akchagyl and «Subakchagyl» deposits (Pliocene) of Sunzha River near Alda village, Ciscaucasia. 

● Holotype: PIN. 

● Distribution: East Georgia (Telavi District, Alvani village; Kvareli District, Eniseli-Tarsagalavani village; 

Gremi village, Mount Chrdili) (Likharev, Schileyko, MS). 

Chondrula tridens (Müller, 1774) T 

[= Chondrus major Krynicki, 1833; Chondrus elatus Andrzejowski in Krynicki, 1833; Pupa tricallosa Parreyss 

in Krynicki, 1833; ?Chondrus microstomus Andrzejowski in Krynicki, 1833; Pupa tridens var. eximia 

Rossmässler, 1835; Buliminus albolimbatus L. Pfeiffer, 1848; Buliminus bayeri L. Pfeiffer, 1848; Buliminus 

tridens var. attenuatus Issel, 1865; Chondrus galiciensis Clessin, 1879; Buliminus tridens var. kubanensis O. 

Boettger, 1879; Buliminus tridens var. caucasicus Martens, 1880; Buliminus tridens var. major f. marcida O. 

Boettger, 1886; Chondrula tridens var. tenuilabiata Lindholm, 1901; Buliminus tridens var. exiguus Retowski, 

1914; Buliminus tridens var. terkensis Retowski, 1914] 

Helix tridens Müller, 1774: 106-107. 

● Type locality: «in Italia». 


● Distribution: East-European Plain (Ryazan Region, steppe zone to the Ural River in the east), Crimea, Caucasus, 

Carphahians (Likharev, Schileyko, MS; Sverlova, 2006). 

Chondrus Cuvier, 1817 

Type species: Bulimus zebra Olivier, 1801 (SD Gray, 1847) 

Chondrus zebrula (Férussac, 1821) T 

[= Bulimus zebra Olivier, 1801 non Bruguière, 1792; Bulimus tantalus L. Pfeiffer, 1868; Buliminus olympicus 

Kobelt, 1877; Brephulus bithynicus Galland, 1884; Brephulus zebra var. anatolica Galland, 1884; Brephulus 

brardus Galland, 1884; Brephulus zebropsis Galland, 1884; Buliminus euxinus Retowski, 1889] 

Helix (Cochlogena) zebrula Férussac, 1821: 58, 70 (nom. nov. pro Bulimus zebra Olivier, 1801). 

● Type locality: «Gemleck, les Dardanelles» (Gemelek, Turkey). 

● Lectotype (Schileyko, 1998b): MNHN. 

● Distribution: empty shells found in sea debris in Crimea (near Sudak) and on the sea shore in Poti (between 

mouths of Rioni and small river flowing from Paleostom Lake) (Likharev, Schileyko, MS). 

Ena Turton, 1831 

Type species: Bulimus montanus Draparnaud, 1801 (SD Herrmannsen, 1847)

Ena montana (Draparnaud, 1801) T 

Bulimus montanus Draparnaud, 1801: 65. 

● Type locality: «dans les montagnes des Cévennes et de la Savoie» (France). 


● Distribution: western regions of Ukraine, Latvia, Estonia, Valdai Hills, vicinities of Moscow and St.- 

Petersburg, ? South Ural (Likharev, Schileyko, MS). 

Georginapaeus Schileyko, 1988 

Type species: Bulimus hohenackeri L. Pfeiffer, 1848 (OD) 

Georginapaeus hohenackeri (L. Pfeiffer, 1848) T 

[= Bulimus hohenackeri Krynicki, 1837, nom. nud.; Buliminus hohenackeri var. intermedius Mousson, 1863; 

Buliminus hohenackeri f. leucolaemis Lindholm, 1922] 

Bulimus hohenackeri Pfeiffer, 1848: 223. 

● Type locality: Georgia. 


● Distribution: central and eastern parts of North Caucasus, Transcaucasia (Likharev, Schileyko, MS). 

Peristoma Krynicki, 1833 

Type species: Peristoma merduenianum Krynicki, 1833 (monotypy) 

Peristoma boettgeri (Clessin, 1883) T 

[= Bulimus assimilis Rossmässler, 1837; Buliminus kusnetzowi Kobelt, 1907; Buliminus brauneri Lindholm, 

1908] 

Buliminus boettgeri Clessin, 1883: 50, pl. 2, fig. 15. 



● Distribution: Georgia: Kutaisi, Tekhuri River valley, vicinities of Tzageri, Borzhomi, Teberda Natural Reserve 


Peristoma lanceum Schileyko, 1984 T 

Schileyko, 1984: 343, fig. 250. 

● Type locality: Bzyb River valley near Pitzunda. 



● Remark: records of Peristoma merduenianum from some localities of West Caucasus – Borzhomi, Kutaisi 

(Mousson, 1873; Boettger, 1881a, as Buliminus tener) – probably refer to this species. 

Peristoma merduenianum Krynicki, 1833 T 

[= Bulimus tener Rossmässler, 1837; ? Buliminus fragilis Anton, 1839] 

Krynicki, 1833: 421-423, pl. 9, fig. 7a-d. 

● Type locality: «Merdven and along Aila» (mountain Crimea). 


● Distribution: mountain Crimea (Likharev, Schileyko, MS). 

Peristoma rupestre (Krynicki, 1833) T 

[= Bulimus assimilis Rossmässler, 1837; Buliminus kusnetzowi Kobelt, 1907; Buliminus brauneri Lindholm, 

1908] 

Buliminus rupestris Krynicki, 1833: 413-415, pl. 3, fig. 5a-d. 

● Type locality: Merdven village (mountain Crimea). 


● Distribution: mountain Crimea (Likharev, Schileyko, MS).

Ramusculus Lindholm, 1925 

Type species: Bulimus subulatus Rossmässler, 1837 (OD) 

Ramusculus subulatus (Rossmässler, 1837) T 

Bulimus subulatus Rossmässler, 1837: 48, fig. 393. 

● Type locality: «? probably Crimea». 



Thoanteus Lindholm, 1925 

Type species: Bulimus gibber Krynicki, 1833 (OD) 

Thoanteus ferrarii Hausdorf, 1994 T 

Hausdorf, 1994:b354-355, figs. 1-3. 

● Type locality: rock opposite the pioneer-camp «Artek-Lazurnyi@, 10-20 m above the lower road between 

Artek and Gurzuf, Yalta District, Crimea. 



Thoanteus gibber (Krynicki, 1833) T 

[= Bulimus revolutus Parreyss in Krynicki, 1833; Bulimus revolutus Rossmässler, 1837; Bulimus phorcus 

Bourguignat, 1855] 

Bulimus gibler [laps. cal., corrected by Krynicki himself in 1837] Krynicki, 1833: 416, pl. 3, fig. 6a-e. 

● Type locality: «between Merdven and Skel’yu village» (Crimea). 



Zebrina Held, 1837 

Type species: Helix detritus Müller, 1774 (SD Gray, 1847) 

Zebrina detrita (Müller, 1774) T 

[= Bulimus radiatus Bruguière, 1789] 

Helix detritus Müller, 1774: 101-102. 

● Type locality: «in Italia & in Saxonia». 


● Distribution: Ukraine (flat areas of Transcarpathia, Mala Kopanya village), probably southwestern Transcaucasia 


MULTIDENTULINAE Schileyko, 1978 

Euchondrus O. Boettger, 1883 

Type species: Pupa chondriformis Mousson, 1861 (monotypy) 

Euchondrus acutior (Lindholm, 1922) T 

Chondrula acutior Lindholm, 1922c: 358-359. 

● Type locality: “im Genist des Fl. Tschoroch bei Kapandidi (unweit Batum)“. 


● Distribution: Chorokh River valley. 

Euchondrus lamelliferus (Rossmässler, 1859) T 

[= ? Chondrus phasianus Mousson, 1863; Buliminus clessini Retowski, 1883; Buliminus lamelliferus var. 

angustatus Retowski, 1887]

Bulimus lamellifera Rossmässler, 1859: 95, pl. 83, fig. 919. 

● Type locality: Syria. 


● Distribution: there are no reliable records supported by anatomical studies (the shell of the species is hardly 

distinguishable from Improvisa pupoides) but the species is probably present in Adzharia near the border of 

Turkey (Likharev, Schileyko, MS). 

Improvisa Schileyko, 1978 

Type species: Chondrus pupoides Krynicki, 1833 (OD) 

Improvisa pupoides (Krynicki, 1833) T 

[= Buliminus angustior Retowski, 1888] 

Chondrus pupoides Krynicki, 1833: 410. 

● Type locality: Mashuk Mount near Pyatigorsk (North Caucasus). 


● Distribution: North Caucasus and Transcaucasia (Likharev, Schileyko, MS). 

Pentadentula Suvorov, 2006 

Type species: Pentadentula balandinae Suvorov, 2006 (OD) 

Pentadentula balandinae Suvorov, 2006 T 

Suvorov, 2006: 93-96, figs. 1-2. 

● Type locality: NW Caucasus, Russia, Krasnodar Territory, valley of Mzymta river, surroundings of 

Monastyr settlement, Akh-Tsu canyon, in the soil in ledges of lime rocks, near the bottom of canyon. 



Senaridenta Schileyko, 1978 

Type species: Chondrula nachicevanjensis Hudec, 1972 (OD) 

Senaridenta nachicevanjensis (Hudec, 1972) T 

Chondrula nachicevanjensis Hudec, 1972: 217, figs. 2, 5. 

● Type locality: vicinities of Stepanakert, Nagornyi Karabakh. 



CLAUSILIIDAE Gray, 1855 

SERRULININAE Ehrmann, 1927 

Caspiophaedusa Lindholm, 1924 

Type species: Clausilia perlucens O. Boettger, 1877 (OD) 

Caspiophaedusa perlucens (O. Boettger, 1877) T 

[= Serrulina signifera Westerlund, 1897] 

Clausilia perlucens Boettger, 1877a: 69. 

● Type locality: «Kaukasus». 

● Lectotype (Zilch, 1954): SMF No. 61559a. 

● Distribution: Talysh, southern slopes of Great Caucasus from Lagodekhi to Khudat, north and northeastern 

parts of Lesser Caucasus and Zangezur (Likharev, Schileyko, MS).

Pontophaedusa Lindholm, 1924 

Type species: Clausilia funiculum Mousson, 1863 (OD) 

Pontophaedusa funiculum (Mousson, 1863) T 

Clausilia funiculum Mousson, 1863: 397-398. 

● Type locality: «Chysirkaleh» (near Poti). 


● Distribution: Black Sea coast from Sochi to Turkey (Likharev, Schileyko, MS). 

● Remark: Egorov (2001) stated that there is the lectotype of C. funiculum in SMF (No. 61488/5). However, 

Zilch (1954) did not mentioned any type specimens of the species, whereas that number belongs to the 

specimen illustrated in the Kobelt's chapter in Rossmässler's "Iconographie". 

Pravispira Lindholm, 1924 

Type species: Clausilia semilamellata Mousson, 1863 (OD) 

Pravispira semilamellata (Mousson, 1863) T 

[= Clausilia semilamellata var. serrulosa Retowski, 1889] 

Clausilia semilamellata Mousson, 1863: 396-397. 

● Type locality: «Rédutkaleh» (north of Poti). 

● Syntypes: ZMZ No. 516635. 

● Distribution: forest areas of western and central Great Caucasus, Black Sea coast from Sochi to Turkey, 

West and South Georgia, Kutkashen District in Azerbaijan (Likharev, Schileyko, MS). 

● Remark: Egorov (2001) stated that there are syntypes of C. semilamellata in SMF (No. 61536a/5). However, 

Zilch (1954) did not mentioned any type specimens of the species. 

Serrulina Mousson, 1873 

Type species: Clausilia sieversi L. Pfeiffer, 1871 (OD) 

Serrulina serrulata serrulata (L. Pfeiffer, 1847) T 

[= Clausilia gracilior Mousson, 1863; Clausilia erivanensis Issel, 1865; Clausilia serrulata var. semiserrata 


Clausilia serrulata Pfeiffer, 1847: 71. 

● Type locality: «Tauria» (Crimea). 


● Distribution: western and central forest areas of North Caucasus and Transcaucasia, Transcarpathian Region 

of Ukraine (Tyachevsky District, between Tereblya and Teresva rivers). 

● Remark: The type locality is evidently erroneous, because the species does not live in Crimea. The type 

could be collected in sea debris. 

Egorov (2001) stated that there is the lectotype of C. serrulata in SMF (No. 61514/2). However, Zilch 

(1954) did not mentioned any type specimens of the species. 

Serrulina sieversi sieversi (L. Pfeiffer, 1871) T 

Clausilia sieversi Pfeiffer, 1871: 70-71. 

● Type locality: «Caucasia meridionalis» (Lenkoran, Azerbaijan). 


● Distribution: South Azerbaijan: forests of Talysh and Lenkoran lowland (Likharev, Schileyko, MS). 

● Remark: Egorov (2001) stated that there is the lectotype of C. serrulata in SMF (No. 61527a/6). However, 

Zilch (1954) did not mentioned any type specimens of the species. 

Serrulina sieversi occidentalis Likharev, 1962 T 

Likharev, 1962a: 118. 

● Type locality: Azerbaijan, Zakatal Region, Tzilban-chai River. 

● Holotype: ZIN.

● Distribution: eastern Georgia and Azerbaijan: mountain desiduous forests of southern slopes of Great Caucasus 

and adjacent lowland (Likharev, Schileyko, MS). 

Serrulinella Nordsieck, 1864 

Type species: Serrulina senghanensis de Morgan in Germain, 1933 (OD) 

Serrulinella senghanensis (de Morgan in Germain, 1933) T 

Serrulina senghanensis de Morgan – Germain, 1933: 391. 

● Type locality: «Le Ghilan: Siah Senghan, Titi, Serd-вb-è-Bala». 

● Lectotype (Nordsieck, 1978, as «holotype»): MNHN. 

● Distribution: Georgia (Egorov, 2001). 

Truncatophaedusa Majoros, Nemeth et Szili-Kovacs, 1994 

Type species: Truncatophaedusa evae Majoros, Nemeth et Szili-Kovacs, 1994 (OD) 

Truncatophaedusa evae Majoros, Nemeth et Szili-Kovacs, 1994 T 

Majoros et al., 1994: 124-125, fig. 1. 

● Type locality: valley of Dagomys River near Nor Luis village, 10 km northward from Dagomys village. 

● Holotype: private collection of Szili-Kovacs. 

● Distribution: West Caucasus adjacent to Black Sea coast (Egorov, 2001). 

ALOPIINAE A. Wagner, 1913 

Alopia H. et A. Adams, 1855 

Type species: Clausilia bielzii L. Pfeiffer, 1848 (SD Westerlund, 1902) 

Alopia glauca (Bielz, 1853) T 

Balea glauca Bielz, 1853: 130. 



● Distribution: probably in Ukrainian Transcarpathia, in areas adjacent to Molodova and Bistritza river basins. 

Cochlodina Férussac, 1821 

Type species: Clausilia bidens Draparnaud, 1801 (= Turbo laminata Montagu, 1803) (SD Pilsbry, 1922) 

Cochlodina cerata (Rossmässler, 1836) T 

[= Cochlodina pareyssi Rossmässler, 1836] 

Clausilia cerata Rossmässler, 1836b: 15, fig. 258. 

● Type locality: «Buccovina» (Ukraine, Chernovtzy Region). 


● Distribution: Ukraine (Chernovtzy and Zakarpatye regions) (Likharev, Schileyko, MS). 

Cochlodina costata (Ziegler in C. Pfeiffer, 1828) T 

[= Cochlodina commutata Rossmässler, 1836] 

Clausilia costata Ziegler – Pfeiffer, 1828: 42, pl. 7, figs. 17, 18. 

● Type locality: «Illyrien». 


● Distribution: Ukraine: Zakarpatye Region (Polevina, 1959; Zdun, 1960). 

● Remark: Sverlova (2006) stated that the species identification by Zdun was erroneous and therefore the 

species presence in Ukraine requires confirmation.

Cochlodina laminata (Montagu, 1803) T 

Turbo laminatus Montagu, 1803: 359-361, pl. 11, fig. 4. 

● Type locality: Lackham wood and Bow Wood (Wiltshire), neighbourhood of Sandwich (Kent) (England). 


● Distribution: East-European Plain, mountain Crimea, Stavropol Region (Likharev, Schileyko, MS); Carpathians 


Cochlodina orthostoma (Menke, 1830) T 

Clausilia orthostoma Menke, 1830: 130. 

● Type locality: vicinities of Leipzig (Germany). 


● Distribution: Carpathians, Baltic countries, northwestern, western and central regions of East-European 

Plain (Likharev, Schileyko, MS). 

MENTISSOIDEINAE Lindholm, 1924 

Acrotoma O. Boettger, 1881 

Type species: Clausilia komarovi O. Boettger, 1881 (OD) 

Acrotoma baryshnikovi Likharev et Schileyko, 2007 T 

Acrotoma (Iliamneme) baryshnikovi Likharev, Schileyko, 2007: 65-67, figs.1-2. 

● Type locality: Georgia, South Ossetia, Dzhavsky district, environs of Akhsardgin (about 2 km of Kvaisi 

settlement), Taling-Leget Cave, 1700 m above sea level, 

● Holotype: ZIN No. 2/554-1983. 


Acrotoma claussi Nordsieck, 1977 T 

Nordsieck, 1977: 91, textfig. 1, pl. 5, fig. 21. 

● Type locality: Abkhazia, Bzyb River valley, upstream of Goluboe Lake [this is a very small lake, ca 200 

square meters on the way to well-known Ritza Lake]. 



Acrotoma gegica Suvorov, 2002 T 

Suvorov, 2002: 161-163, fig. 1 A-E. 

● Type locality: NW Caucasus, Abkhazia, Ritsinsky National Park, valley of Gega River (right tributary of 

Bzyb River), surroundings of Gegsky waterfall. 



Acrotoma juliae Suvorov, 2002 T 

Suvorov, 2002: 165-166, fig. 3 A-E. 

● Type locality: NW Caucasus, Abkhazia, Ritsinsky National Park, valley of Jupshara River (left tributary of 

Gega River), canyon «Jupsharskie vorota». 



Acrotoma komarowi (O. Boettger, 1881) T 

Clausilia (Acrotoma) komarowi O. Boettger, 1881b: 341-342. 

● Type locality: ««Kodor» Transcaucasiae in funibus Ponticus Caucasi» (Abkhazia, middle reaches of Kodor 

River). 

● Holotype: SMF No. 144037.

● Distribution: Abkhazia (Likharev, Schileyko, MS). 

Acrotoma laccata (O. Boettger, 1881) T 

Clausilia laccata O. Boettger, 1881b: 342-343. 

● Type locality: «in parte meridionali Caucasi Pontici». 

● Holotype: SMF No. 144040/1. 


● Remark: as mentioned by Boettger in the description, the species was described from a single shell, therefore 

the «lectotype» in Zilch, 1976a is in fact holotype. 

Acrotoma narzanensis (Rosen, 1901) T 

Clausilia narzanensis Rosen, 1901b: 10. 

● Type locality: Kislovodsk, Kovarstva Castle. 

● Lectotype (Likharev, 1962a): ZIN. 

● Distribution: North Caucasus (vicinities of Kislovodsk, Kabardino-Balkaria) (Likharev, Schileyko, MS). 

Acrotoma semicincta (O. Boettger, 1881) T 

[= Clausilia (Acrotoma) semicincta var. ciscaucasica Rosen, 1901] 

Clausilia (Acrotoma) semicincta O. Boettger, 1881b: 343-344. 

● Type locality: Teberda (North Caucasus). 


● Distribution: western part of North Caucasus in upper reaches of Kuban and Podkumka rivers (Likharev, 


● Remark: the lectotype designation by Zilch (1976a: 204, fig. 2; SMF No. 144037) is invalid because the 

lectotype has been previously designated by Likharev (1962a: 159). 

Acrotoma tunievi Suvorov, 2002 T 

Suvorov, 2002: 163-164, fig. 2 A-E. 

● Type locality: NW Caucasus, Russia, Krasnodar Territory, surroundings of Khosta settlement, Kavkazsky 

State Natural Biosphere Reserve, Khosta yew-box tree wood. 



Akramowskia Norsieck, 1975 

Type species: Euxina akramowskii Likharev, 1962 (OD) 

Akramowskia akramowskii (Likharev, 1962) T 

Euxina akramowskii Likharev, 1962b: 221, figs. 1-3. 

● Type locality: Khustup Mount near Vachagan village, Kafan District, Armenia. 



Akramowskia valentini (Loosjes, 1964) T 

Armenica valentini Loosjes, 1964: 146, figs. 3-5. 

● Type locality: «between Chaikend and Tandzaver» (Kafan District, Armenia). 


● Distribution: Gekhi River basin, Kafan District, Armenia (Likharev, Schileyko, MS). 

Armenica O. Boettger, 1877 

Type species: Clausilia laevicollis Charpentier, 1852 (monotypy) 

Armenica disjuncta armenica Nordsieck, 1975 T 

Nordsieck, 1975: 97.

● Type locality: Armenia, Ekhegnadzor District, Gnishik village. 


● Distribution: Armenia (Egorov, 2001). 

Armenica gracillima (Retowski, 1889) T 

Clausilia (Oligoptychia) gracillima Retowski, 1889: 261. 

● Type locality: debris of Chorokh River near Batumi. 

● Lectotype (Zilch, 1976a): SMF No. 144152. 

● Distribution: Adzharia (Likharev, Schileyko, MS). 

Armenica griseofusca (Mousson, 1876) T 

[= Clausilia commena Retowski, 1889] 

Clausilia griseofusca Mousson, 1876b: 145-146, pl. 5, fig. 3. 

● Type locality: «environs de Tabizhuri» (vicinities of Tabatzkuri Lake, South Georgia). 

Syntype (?): SMF No. 144161. 

● Distribution: western and southeastern Georgia (Egorov, 2001). 

● Remark: Egorov (2001) has designated a lectotype of the species (SMF No. 144161). However, the designation 

is invalid because the respective specimen is merely cited as "lectotype" (ICZN Declaration 44). Zilch 

(1954) with a query listed that specimen as a syntype. 

Armenica likharevi Nordsieck, 1975 T 

Nordsieck, 1975: 85. 

● Type locality: Armenia, Idzhevan District, Laki ravine near Verin Agdan village. 



Armenica unicristata (O. Boettger, 1877) T 

[= Laciniaria lantzi Lindholm, 1924; Armenica brunnea sensu Likharev, 1962a, non Rossmässler, 1839] 

Clausilia unicristata O. Boettger, 1877b(August): 75-76. 

● Type locality: Ekaterinenfeld (now – Bolnisi, South Georgia). 


● Distribution: southern slopes of Great Caucasus between Zakataly and Shemakha, northern and northeastern 

parts of Lesser Caucasus, Zangezur (Likharev, Schileyko, MS). 

Armenica zakatalica Nordsieck, 1977 T 

[= Armenica hueti sensu Likharev, 1962, non Mortillet, 1854] 

Nordsieck, 1977: 93. 

● Type locality: Great Caucasus, Zakatal Natural Reserve, Rychug Ridge, south slope near Gubor-chai River. 


● Distribution: type locality, Shemakha District (Azerbaijan), Samur River valley near Ikhrek village (Daghestan) 

(Egorov, 2002). 

Elia H. et A. Adams, 1855 

Type species: Clausilia moesta Rossmässler, 1839 (OD) 

Elia derasa (Mousson, 1863) T 

[= Clausilia ossetica Schmidt, 1868; Clausilia sandbergeri Mousson, 1873; Clausailia derasa f. suanetica O. 


Clausailia derasa Mousson, 1863: 400-401. 

● Type locality: «Rédutkaleh» (north of Poti), «Koutais» (Kutaisi). 

● Syntypes: ZMZ No. 516718; SMF Nos. 133462, 133456a. 

● Distribution: West Transcaucasia and Central Ciscaucasia (Likharev, Schileyko, MS).

Elia novorossica (Retowski, 1888) T 

[= Clausilia somchetica var. dissophya Westerlund, 1897] 

Clausilia (Euxina) novorossica Retowski, 1888: 284. 

● Type locality: Novorossiisk. 

● Lectotype (Likharev, 1962a): IZP. 

● Distribution: West Caucasus: mountains between Novorossisk and Anapa (Likharev, Schileyko, MS), in 

the riverside woods in the southern part of Lugansk Region, Ukraine (Gural-Sverlova, Martynov, 2009). 

Elia ossetica (Mousson, 1863) T 

[= Clausilia tschetschenica L. Pfeiffer, 1866] 

Clausilia somchetica var. ossetica Mousson, 1863: 399-400. 



● Distribution: central areas of North Caucasus, Inner Daghestan, Georgia (except western regions), North 

Armenia (Likharev, Schileyko, MS). 

Elia somchetica somchetica (L. Pfeiffer, 1846) T 

[= Clausilia kolenatii Siemaschko, 1847; Clausilia eichwaldi Siemaschko, 1847; Clausilia colchica Pareyss 

in L. Pfeiffer, 1857] 

Clausilia somchetica L. Pfeiffer, 1846: 94. 

● Type locality: Somchetia (Kakhetia, East Georgia). 

● Syntype: SMF No. 133500a. 

● Distribution: forest and subalpine zones of Caucasus (Likharev, Schileyko, MS). 

Elia somchetica raddei (Mousson, 1876) T 

Clausilia raddei Mousson, 1876a: 43-44, pl. 4, fig. 3. 

● Type locality: Mount Schambodel, height 6000 feet, south of Achalzich (North Armenia). 


● Distribution: North Armenia, Akstafa-chai River valley (Egorov, 2001). 

● Remark: Egorov (2001) designated the lectotype (SMF No. 133512/4), though the designation is invalid 

because he merely cited a specimen as a lectotype (ICZN Declaration 44). Moreover, Zilch (1954) did not 

mention any specimen of the type series in the SMF. 

Elia tuschetica Likharev et Lezhawa, 1961 T 

Likharev, Lezhawa, 1961: 473, figs. 1-3. 

● Type locality: left bank of Tusheti Alazan, 1600 m above sea level. 


● Distribution: mountain Tushetia (southwestern part of highland Daghestan), Tusheti Alazan basin (Likharev, 


Euxina O. Boettger, 1877 

Type species: Clausilia hetaera L. Pfeiffer, 1848 (OD) 

Euxina gastron Nordsieck, 1995 T 

[= Euxina persica persica sensu Likharev, 1962a (part., non O. Boettger, 1877] 

Nordsieck, 1995: 11, pl. 1, fig. 1. 

● Type locality: «Prov. Gilan, Tal des Shim Rud zwischen Lahijan und Deilaman, Tchalmeh roud» (northern 

Iran). 


● Distribution: Talysh (Vasharu-chai River valley) (Egorov, 2001). 

Euxina talyschana Likharev, 1962 T 

Likharev, 1962a: 173, figs. 97, 98.

● Type locality: Talysh, Nyudis-Kalasi Mount. 


● Distribution: Talysh mountains (Likharev, Schileyko, MS). 

Euxinastra O. Boettger, 1888 

Type species: Clausilia hamata O. Boettger, 1888 (monotypy) 

Euxinastra hamata (O. Boettger, 1888) T 

Clausilia hamata O. Boettger, 1888: 152-153. 

● Type locality: Batumi (Adzharia). 


● Distribution: Georgia (Batumi) (Likharev, Schileyko, MS). 

Filosa O. Boettger, 1877 

Type species: Clausilia filosa Mousson, 1863 (OD) 

Filosa filosa (Mousson, 1863) T 

Clausilia filosa Mousson, 1863: 395-396. 

● Type locality: «Chysirkaleh» (vicinites of Poti, Adzharia). 


● Distribution: Black Sea coast of Adzharia (Georgia) (Likharev, Schileyko, MS). 

● Remark: Egorov (2001) designated the lectotype (SMF No. 145384a), though the designation is invalid 

because he merely cited a specimen as a lectotype (ICZN Declaration 44). Moreover, Zilch (1976a) did not 


Kazancia Neubert, 1992 

Type species: Kazancia monticola Neubert, 1992 (OD) 

Kazancia lindholmi (Kobelt in Lindholm, 1912) T 

Clausilia lindholmi Kobelt – Lindholm, 1912: 202. 

● Type locality: Salolet Mount near Artvin City (Turkey, vilayet Chorokh). 


● Distribution: East Georgia (Lagodekhi) (Likharev, Schileyko, MS). 

Mentissoidea O. Boettger, 1877 

Type species: Clausilia fusorium Mousson, 1876 (= Clausilia rupicola Mortillet, 1854) (SD Kennard, 

Woodward, 1923) 

Mentissoidea rupicola rupicola (Mortillet, 1854) T 

[= Clausilia fusorium Mousson, 1876; Clausilia litotes var. litoderma O. Boettger, 1881] 

Clausilia rupicola Mortillet, 1854: 13, pl. 1, fig. 7. 

● Type locality: Tortum (Turkey, vilayet Erzurum). 


● Distribution: Georgia (Adzharia and Black Sea coast northward to Sukhumi) (Likharev, Schileyko, MS). 

Mentissoidea rupicola litotes (A. Schmidt, 1868) T 

[= Clausilia litotes var. ganeo O. Boettger, 1883; Clausilia litotes var. suanetica O. Boettger, 1883] 

Clausilia litotes Schmidt, 1868: 162. 

● Type locality: Osetia. 


● Distribution: Great Caucasus (forests of western areas of northern part; southern parts eastward to Nukha, 

Azerbaijan); isolated range in North Osetia, Chechnya, Ingushetia; southern Transcaucasia (northern parts of 

Lesser Caucasus from Adzharo-Imereti Ridge to Nagornyi Karabakh) (Likharev, Schileyko, MS).

● Remark: Egorov (2001) designated the lectotype (SMF No. 143740), though the designation is invalid because 

he merely cited a specimen as a lectotype (ICZN Declaration 44). Moreover, Zilch (1976a) did not 


Scrobifera O. Boettger, 1877 

Type species: Clausilia foveicollis Charpentier, 1852 (= Clausilia taurica L. Pfeiffer, 1848) (monotypy) 

Scrobifera taurica taurica (L. Pfeiffer, 1848) T 

[= Clausilia taurica Krynicki, 1837, nom. nud; Clausilia foveicollis Charpentier, 1852; Clausilia vinosa 


Clausilia taurica Pfeiffer, 1848: 412. 

● Type locality: «Kutais» (Georgia). 


● Distribution: all forest areas of Caucasus except for Adzharia and Talysh (Likharev, Schileyko, MS). 

● Remark: Egorov (2001) designated the lectotype (SMF No. 144074a), though the designation is invalid 

because he merely cited a specimen as a lectotype (ICZN Declaration 44) and the speciemens do not belong 

to the type series, because they have been collected in 1881. Zilch (1976a) cited only a syntype in the SMF. 

Scrobifera taurica brjanskii (Rosen, 1911) T 

[= Clausilia foveicollis var. schaposchnikovi Rosen, 1911] 

Clausilia foveicollis var. brjanskii Rosen, 1911: 125, pl. 2, fig. 6a-b. 

● Type locality: Adler (Caucasian Black Sea coast). 


● Distribution: northern (Nalchik) and southwestern (Sochi) Caucasus (Egorov, 2001). 

Strigileuxina Nordsieck, 1994 

Type species: Clausilia (Euxina) reulauxi Boettger, 1887 (OD) 

Strigileuxina reulauxi (O. Boettger, 1887) T 

Clausilia (Euxina) reulauxi Boettger, 1887: 55-56. 

● Type locality: Batumi (Adzharia). 

● Lectotype (Nordsieck, 1975): SMF No. 145389. 

● Distribution: Georgia (Adzharia) (Likharev, Schileyko, MS). 

CLAUSILIINAE Mörch, 1864 

Clausilia Draparnaud, 1805 

Type species: Pupa rugosa Draparnaud, 1801 (SD Turton, 1831) 

Clausilia bidentata (Strøm, 1765) T 

[= Clausilia nigricans Maton et Rackett, 1807; Clausilia bidentata var. septentrionalis A. Schmidt, 1857; 

Clausilia bidentata var. exigua Westerlund, 1871; Clausilia bidentata var. elongata Clessin, 1878] 

Turbo bidentata Strøm, 1765: 436, pl. 6, fig. 17. 

● Type locality: Scandinavia. 


● Distribution: Baltic countries (Kurzeme, Hiumaa and Saaremaa islands, northwestern Estonia), Leningrad 

Region (Primorsk) (Likharev, Schileyko, MS). 

Clausilia cruciata (Studer, 1820) T 

[= Clausilia cruciata f. minima A. Schmidt, 1856; Clausilia cruciata f. gracilis A. Schmidt, 1856] 

Glischrus (Clausilia) cruciata Studer, 1820: 89. 

● Type locality: «Kt. Wallis, Leukerbad» (Switzerland).


● Distribution: Baltic countries (far from sea shore), Leningrad Region, southwestern Karelia (north of 

Ladoga Lake), sporadically in Pskov, Vitebsk, Moscow, Tver, Brest regions and in Carpathians (Likharev, 


Clausilia dubia Draparnaud, 1805 T 

[= Clausilia roscida Studer, 1820; Clausilia dubia var. speciosa A. Schmidt, 1857; Clausilia dubia var. vindobonensis 

A. Schmidt, 1857; Clausilia dubia var. obsoleta A. Schmidt, 1857; Clausilia dubia var. alpicola 

Clessin, 1878; Clausilia dubia var. podolica Bąkowski, 1880; Clausilia dubia f. runensis Tschapeck, 1883] 

Draparnaud, 1805: 70, pl. 4, fig. 10. 


● Lectotype (Bank, Gittenberger, 2000): NHMV. 

● Distribution: Baltic countries, Pskov and Tver regions, near Vitebsk, «Belovezhskaya Pushcha Natural Reserve, 

Carpathians, Volyno-Podolsk Hills (Likharev, Schileyko, MS). 

Clausilia pumila pumila C. Pfeiffer, 1828 T 

Pfeiffer, 1828: 41, pl. 7, fig. 16. 

● Type locality: Illyria. 


● Distribution: Baltic countries (Lithuania, Latvia, southern Estonia), Kaliningrad Region, sporadically in 

Carpathians and Volyno-Podolsk Hills (Likharev, Schileyko, MS). 

Clausilia pumila sejuncta A. Schmidt in Westerlund, 1871 T 

Clausilia sejuncta A. Schmidt – Westerlund, 1871: 75. 

● Type locality: «in Suecia media, in montibus Omberg et Mösseberg». 


● Distribution: Baltic countries (Lithuania, Latvia), Moscow, Yaroslavl, Tver, Smolensk, Kaluga, Tula regions, 

eastward to Kostroma, southward to Orel Region (Likharev, Schileyko, MS). 

Macrogastra Hartmann, 1840 

Type species: Pupa ventricosa Draparnaud, 1801 (monotypy) 

Macrogastra latestriata latestriata (Bielz in A. Schmidt, 1857) T 

Clausilia latestriata Bielz – A. Schmidt, 1857: 27-28, figs. 52-56, 183, 184. 



● Distribution: Ukraine (Carpathians and Volyno-Podolsk Hills) (Likharev, Schileyko, MS). 

Macrogastra latestriata borealis (O. Boettger, 1878) T 

Clausilia latestriata var. borealis Boettger, 1878c: 136. 

● Type locality: «Neukuhren in Samland, Ostprussien» (now – Pionersky settlement, Kaliningrad Region). 


● Distribution: Baltic countries, deciduous forests of Valdai Hills (Shikov, 1982), Byelorussian Polesye, vicinities 

of Vitebsk (Likharev, Schileyko, MS); Ukranian Polesye (Rovno and Zhitomir regions) (Sverlova, 

2006). 

Macrogastra plicatula (Draparnaud, 1801) T 

Pupa plicatula Draparnaud, 1801: 64. 



● Distribution: northwestern, western and western-central areas of Russia; Chatyr-Dagh, mountain Crimea 

(Likharev, Schileyko, MS); northwestern Ukraine (Sverlova, 2006).

Macrogastra tumida (Ziegler in Rossmässler, 1836) T 

Clausilia ventricosa var. tumida Ziegler – Rossmässler, 1836b: 21, fig. 277. 

● Type locality: «Buccovina» (now – Chernovtzy Region, Ukraine). 


● Distribution: Ukraine: Ivano-Frankovsk, Lvov, Ternopol, Zakarpatskaya, Chernovtzy regions (Likharev, 


Macrogastra ventricosa (Draparnaud, 1801) T 

Pupa ventricosa Draparnaud, 1801: 62-63. 


● Syntypes: NHMV, MNHN. 

● Distribution: Baltic countries, sporadically in mixed and desiduous forests of northwestern, western and 

central areas of Russia (Likharev, Schileyko, MS); western and northwestern Ukraine (Sverlova, 2006). 

Ruthenica Lindholm, 1924 

Type species: Clausilia filograna Ziegler in Rossmässler, 1836 (OD) 

Ruthenica filograna (Ziegler in Rossmässler, 1836) T 

Clausilia filograna Ziegler – Rossmässler, 1836b: 17-18, fig. 264. 

● Type locality: «Oesterreich, Illyrien, Bucovina, Croatien». 


● Distribution: Baltic countries, northwestern and central Russia, Byelorussia, Ukrainian Carpathians and 

Ciscarpathia (Likharev, Schileyko, MS); western Podolsk Hills, Volyn Polesye, Kiev region of the Ukraine 


BALEINAE A. Wagner, 1913 

Alinda H. et A. Adams, 1855 

Type species: Turbo biplicata Montagu, 1803 (SD Martens in Albers, 1860) 

Alinda biplicata (Montagu, 1803) T 

Turbo biplicata Montagu, 1803: 361, pl. 11, fig. 5. 

● Type locality: south England. 


● Distribution: Transcarpathia (Ukraine), Kaliningrad Region, Lithuania, Moscow Region (Imre, 1941; 

Egorov, 2002). 

Alinda fallax (Rossmässler, 1836) T 

Clausilia fallax Rossmässler, 1836b: 16-17, fig. 262. 

● Type locality: «Galicien». 


● Distribution: Ukraine (Ivano-Frankovsk, Ternopol, Chernovtzy regions) (Baidashnikov, 1988). 

Alinda stabilis (L. Pfeiffer, 1847) T 

Clausilia stabilis L. Pfeiffer, 1847: 70. 

● Type locality: Banat. 


● Distribution: Ukraine (Carpathians, Coscarpathia to Vinnitza Region) (Likharev, Schileyko, MS; Sverlova, 

2006). 

Balea Gray, 1824 

Type species: Pupa fragilis Draparnaud, 1801 (= Turbo perversus Linnaeus, 1758) (SD Martens in Albers,

1860) 

Balea perversa (Linnaeus, 1758) T 

Turbo perversus Linnaeus, 1758: 767. 


Type: BMNH. 

● Distribution: species found on one islet near the coast of Estonia (Krausp, 1940). 

● Remark: Likharev (1962a) suggested that the species lives also in mountain Crimea. However, the material 

most probably belongs to Mentissa gracilicosta. 

Bulgarica O. Boettger, 1877 

Type species: Clausilia varnensis L. Pfeiffer, 1848 (OD) 

Bulgarica cana (Held, 1836) T 

Clausilia cana Held, 1836: 275. 

● Type locality: Bayern (Germany). 


● Distribution: East-European Plain to Kazan in the east (Likharev, Schileyko, MS); Carpathians (Sverlova, 

2006). 

Bulgarica vetusta (Rossmässler, 1836) T 

Clausilia vetusta Rossmässler, 1836b: 16, Taf. 18, fig. 261. 

● Type locality: Slovenia, Steiner Alpen und Gorjanz. 


● Distribution: northern Moldova, probably western Ukraine (Egorov, 2002). 

Laciniaria Hartmann, 1844 

Type species: Pupa plicata Draparnaud, 1801 (monotypy) 

Laciniaria plicata (Draparnaud, 1801) T 

Pupa plicata Draparnaud, 1801: 63. 

● Type locality: «Mont Jura» (France). 


● Distribution: mixed and deciduous forests of Baltic countries (except Moozund Islands), western, southwestern 

(Transcarpathia, Carpathians, Ciscarpathia, Volyno-Podolsk Hills, northern and central Ukraine, 

river valleys in Northwestern Black Sea Maritime Territory) and central regions of East-European Plain (Likharev, 

Schileyko, MS; Sverlova, 2006). 

Likharevia Nordsieck, 1975 

Type species: Clausilia (Oligoptychia) gustavi O. Boettger, 1880 (OD) 

Likharevia gustavi (O. Boettger, 1880) T 

Clausilia (Oligoptychia) gustavi Boettger, 1880b: 381-382. 

● Type locality: Shindan-kalasi Mount, Talysh. 

● Lectotype (Nordsieck, 1975): SMF No. 144147. 

● Distribution: Talysh, isolated mounts above forest zone and on rocks and rock slides in Diabar depression 

(Zuvant) (Likharev, Schileyko, MS). 

Mentissa H. et A. Adams, 1855 

Type species: Clausilia canalifera Rossmässler, 1836 (SD Martens in Albers, 1860) 

Mentissa canalifera (Rossmässler, 1836) T 

[= Clausilia detersa Rossmässler, 1836; Clausilia acridula Rossmässler, 1836]

Clausilia canalifera Rossmässler, 1836a: 17-18, fig. 183. 

● Type locality: «Taurien» (Crimea). 


● Distribution: nearly entire mountain-forest Crimea (Baidashnikov, 1990a). 

Mentissa gracilicosta gracilicosta (Ziegler in Rossmässler, 1836) T 

[= Clausilia (Mentissa) gracilicosta natio media Puzanov, 1925] 

Clausilia gracilicosta Ziegler – Rossmässler, 1836a: 18, fig. 184. 

● Type locality: «Taurien» (Crimea). 

● Syntype: SMF No. 133625.. 

● Distribution: the range consists of two parts: western, semicircular, covering southwestern mountain Crimea 

from the Belbek River to Sevastopol and farther nearly to Alupka along sea coast; and eastern – to Ai- 

Petri yaila along sea coast, narrowing to maritime zone towards Yalta (Baidashnikov, 1990a, b); introduced 

to Odessa (Sverlova, 2006) 

Mentissa gracilicosta albocostata Baidashnikov, 1990 T 

Baidashnikov, 1990c: 23-24, fig. 4V. 

● Type locality: eastward of Baidarskaya valley, on western slopes of Ai-Petri Yaila, between villages Rodnikovskoe 

and Kolkhoznoe (southern Crimea). 


● Distribution: southern Crimea: type locality and Baidarskaya valley (Egorov, 2002). 

Mentissa gracilicosta orientalis Baidashnikov, 1990 T 

Baidashnikov, 1990c: 25-26, figs. 2A, 4D. 

● Type locality: Kara-Dagh Natural Reserve, Syuryu-Kaya Ridge (southern Crimea). 


● Distribution: eastern part of southern Crimea (Egorov, 2002). 

Mentissa gracilicosta sodalis (A. Schmidt, 1868) T 

Clausilia sodalis A. Schmidt, 1868: 166-167. 

● Type locality: east of Belokamensk near Sevastopol (southern Crimea). 


● Distribution: southern Crimea from vicinities of Sevastopol to Simferopol (Egorov, 2002). 

Mentissa gracilicosta tschatyrdagika Baidashnikov, 1990 T 

Baidashnikov, 1990c: 26, figs. 2B, 3A, 4G. 

● Type locality: slopes of lower plateau of Chatyr-Dagh near Perevalnoe village (southern Crimea). 


● Distribution: southern Crimea: vicinities of Chatyr-Dagh (Egorov, 2002). 

Mentissa velutina Baidashnikov, 1990 T 

Baidashnikov, 1990a: 21, figs. 1 A, B, 2 A, B, 3 A, B, V. 

● Type locality: southern slopes of Crimean mountains between Goluboi settlement and Opolznevoe village. 

● Holotype: Insitute of Zoology of Ukrainian Academy of Sciences. 

● Distribution: Yalta Mountain-Forest Natural Reserve, Ai-Petri and Yalta yailas and western part of Babugan-Yaila 

(Roman-Kosh Mount), at 650-1500 m above sea level (Crimea) (Likharev, Schileyko, MS). 

Micropontica O. Boettger, 1881 

Type species: Clausilia closta O. Boettger, 1881 (monotypy) 

Micropontica annae Kijashko, 2005 T 

Kijashko, 2005: 157–160, figs. 1, 2. 

● Type locality: Northwest Caucasus, Adygeja, Lagonaki Mountains, western part of Lagonaki

Ridge, southern slope of Mt. Zhitnaya (~ 1900 m above sea level). 

● Holotype: ZIN 1/505–2003. 

● Distribution: type locality (Kijashko, 2005). 

Micropontica circassica (O. Boettger, 1888) T 

[= Clausilia circassica var. claviformis Rosen, 1914; Clausilia retowskii var. saltuensis Rosen, 1914] 

Clausilia (Micropontica) circassica Boettger, 1888: 154. 

● Type locality: Oschten-Fischt Mounts (northwestern Great Caucasus). 

● Syntypes: Überseemuseums, Bremen, Germany No. 114/1-4. 

● Distribution: western part of Great Caucasus (Oshten-Fisht mountain range, upper reaches of Kodor River 

south of Klukhor mountain pass); easternmost finding – in upper reaches of Malka River (Terek basin) near 

Shidzha-Tmas Mount (Likharev, Schileyko, MS). 

Micropontica closta (O. Boettger, 1881) T 

Clausilia (Micropontica) closta Boettger, 1881b: 345-346. 

● Type locality: Psyrtzkhij monastery (Novyi Afon, Abkhazia). 


● Distribution: Western Caucasus, vicinities of Gagra and Novyi Afon (Black Sea coast of 

Abkhazia) (Likharev, Schileyko, MS); Krasnodar Territory, surroundings of Khosta settlement, 

Kavkazsky State Natural Biosphere Reserve, Khosta yew-box tree wood; Sotchinskiy National Park, 

Agurskoye ravine (Tuniev, 2002; Kijashko, 2007; Kijashko, 2009). 

Micropontica interjecta (Rosen, 1914) T 

Clausilia retowskii var. interjecta Rosen, 1914: 206, pi. 3, figs. 17a-b. 

● Type locality. Stanitza Belorechenskaya (northwestern Caucasus, Krasnodar Territory). 


● Distribution: type locality and Maikop (Krasnodar Territory) (Egorov, 2002). 

Micropontica retowskii (O. Boettger, 1888) T 

[= Clausilia baleopsis Westerlund, 1897; Clausilia (Micropontica) retowskii var. simulia Westerlund, 1901] 

Clausilia (Micropontica) retowskii Boettger, 1888: 154-155. 

● Type locality: Guk Mount (near Maikop). 


● Distribution: western part of Great Caucasus near Caucasian Natural Reserve and its immediate vicinities 


Mucronaria O. Boettger, 1877 

Type species: Clausilia acuminata Mousson, 1876 (monotypy) 

Mucronaria acuminata (Mousson, 1876) T 

[= Clausilia acrolepta Martens, 1876] 

Clausilia (Mentissa) acuminata Mousson, 1876b: 144-145, pl. 5, fig. 4. 

● Type locality: «Tabizhuri» (vicinities of Tabatzkuri Lake, South Georgia). 


● Distribution: Shav-Nabad Mount, east of Tabatzkuri Lake (South Georgia) (Likharev, Schileyko, MS). 

Mucronaria duboisi (Charpentier, 1852) T 

[= Clausilia subtilis A. Schmidt, 1868; Clausilia subtilis f. minor A. Schmidt, 1868; Clausilia subtilis f. plicata 

O. Boettger, 1880; Clausilia stomatica Westerlund, 1884] 

Clausilia duboisi Charpentier, 1852: 402, pl. 11, fig. 12. 

● Type locality: Crimea. 


● Distribution: forest areas of Transcaucasia (except for Talysh and Lenkoran lowland), North Caucasus

(Vladikavkaz, Pyatigorsk, Zheleznovodsk) (Likharev, Schileyko, MS). 

Mucronaria index (Mousson, 1863) T 

[= Clausilia climax O. Boettger, 1888] 

Clausilia index Mousson, 1863: 401-402. 

● Type locality: «Rédutkaleh, Chesyrkaleh» (now – Kulevi, north of Poti (Georgia). 


● Distribution: West Georgia, mostly along sea coast (Sukhumi to Batumi), less frequently far from the coast 

(Kutaisi, Sairme, Borzhomi) (Likharev, Schileyko, MS). 

Mucronaria pleuroptychia (O. Boettger, 1878) T 

[= Clausilia pleuroptychia var. polygyra Boettger, 1881] 

Clausilia pleuroptychia Boettger, 1878a: 291-292, pl. 10, fig. 1. 

● Type locality: «Syrien». 


● Distribution: central part of West Georgia between Kvirila River in the east and Tzkhenis-Tzkali in the 

west, in the zone of hills and low limestone mountains (Likharev, Schileyko, MS). 

Mucronaria strauchi (O. Boettger, 1878) T 

Clausilia strauchi Boettger, 1878a: 301-303, pl. 10, fig. 6. 

● Type locality: forest near Tianeti, north of Tbilisi (Georgia). 


● Distribution: East Georgia, mostly near Mtzkheta, Tianeti, Manglisi (vicinities of Tbilisi) (Likharev, 


Quadriplicata O. Boettger, 1878 

Type species: Clausilia quadriplicata A. Schmidt, 1868 (monotypy) 

Quadriplicata aggesta aggesta (O. Boettger, 1879) T 

[= Clausilia protracta Westerlund, 1901; Clausilia aggesta var. praegracilis Rosen, 1914] 

Clausilia aggesta Boettger, 1879a: 34-35, pl. 1, fig. 6. 

● Type locality: debris of Kuban River. 


● Distribution: north branches of western part of Great Caucasus, from Novorossiisk to beginnings of Bolshaya 

Laba River (Kuban basin); south branches – from Krasnaya Polyana to Gudauta District of Abkhazia 


Quadriplicata aggesta stauropolitana (Rosen, 1901) T 

Clausilia (Euxina) stauropolitana Rosen, 1901b: 11. 

● Type locality: Stavropol. 

● Lectotype (Likharev, 1962a): Museum of Georgia No. 1578a. 

● Distribution: Stavropol and vicinities (Likharev, Schileyko, MS). 

Quadriplicata dipolauchen (O. Boettger, 1881) T 

Clausilia (Euxina) dipolauchen Boettger, 1881a: 235-236, pl. 9, fig. 18. 

● Type locality: Kveda-Gordi near Tzkheni-tzkali River (Tzulukidze District of Georgia). 

● Lectotype (Likharev, 1962a): Museum of Georgia, No. 1580a. 

● Distribution: foothills (not higher than 500 m above sea level) and some hills of northern border of Colchis 

maritime plain (not lower than 100 m above sea level), between Khobi and Tzkhaltubo-tzkali rivers (Likharev, 


Quadriplicata lederi lederi (O. Boettger, 1879) T 

[=Clausilia lederi var. triadis Boettger, 1881]

Clausilia (Euxina) lederi Boettger, 1879a: 36-37, pl. 1, fig. 5. 

● Type locality: Suram Ridge. 

● Lectotype (Likharev, 1962a): Museum of Georgia No. 1585. 

● Distribution: south branches of Great Caucasus, from Ochamchiri District of Abkhazia to Tbilisi, 

Mtzkheta; branches of Lesser Caucasus near Borzhomi and Abastumani (Likharev, Schileyko, MS). 

Quadriplicata lederi gradata (O. Boettger, 1879) T 

Clausilia (Euxina) gradata Boettger, 1879b: 409, pl. 10, fig. 5. 

● Type locality: “bei Timotissubani, einem alten Kloster in einem Seitenthal der Kura, 18 Werst von Borshom“. 

● Syntype: SMF No. 133580 (“lectotype”). 

● Distribution: western Georgia, Gudauta (Egorov, 2002). 

Quadriplicata lederi martensi Nordsieck, 1983 T 

Nordsieck, 1983: 205-206, pl. 12, fig. 17. 

● Type locality: “Grusien, nahe Bachmaro bei Chochayauri, 1050-2020 m“. 



Quadriplicata pumiliformis (O. Boettger, 1881) T 

[= Clausilia pumiliformis var. rusticana Retowski, 1914; Clausilia pumiliformis var. inarmata Lindholm, 

1913; Clausilia pumiliformis var. circumplicosa Lindholm, 1913] 

Clausilia (Euxina) pumiliformis Boettger, 1881a: 234-235, pl. 9, fig. 17. 

● Type locality: between Sukhumi and Poti. 


● Distribution: mostly forests of maritime areas between Soch and Sukhumi; some populations in vicinities 

of Novorossiisk and Poti; more continentally in chestnut forest zone (Krasnaya Polyana, Tzebelda); western 

branches of Great Caucasus (Belaya River basin up to 1000 m above sea level) (Likharev, Schileyko, MS). 

Quadriplicata quadriplicata (A. Schmidt, 1868) T 

Clausilia quadriplicata Schmidt, 1868: 163. 

● Type locality: Racha Ridge (West Georgia). 


● Distribution: forests of southern parts of Central and East Great Caucasus, northern and northeastern parts 

of Lesser Caucasus; floodland forests of lower Terek; absent in Talysh and Zangezur (Likharev, Schileyko, 

MS). 

Quadriplicata subaggesta (Retowski, 1887) T 

Clausilia (Euxina) subaggesta Retowski, 1887: 37, pl. 1, fig. 4. 

● Type locality: Crimea, sea debris. 

● Lectotype (Likharev, 1962a): IZP. 

● Distribution: Georgia: subtropical maritime area of Adzharia (Likharev, Schileyko, MS). 

Vestia Hesse, 1916 

Type species: Clausilia elata Rossmässler, 1836 (SD Lindholm, 1924) 

Vestia elata (Rossmässler, 1836) T 

Clausilia elata Rossmässler, 1836a: 20, fig. 190. 



● Distribution: Ukraine: Ciscarpathia (Chernovtzy and Ivano-Frankovsk regions), eastward to Vinnitza Region 

(Likharev, Schileyko, MS; Sverlova, 2006).

Vestia gulo (Bielz, 1859) T 

Clausilia turgida var. gulo Bielz, 1859: 222. 



● Distribution: Ukraine: Lesistye Carpathians, Ciscarpathia, eastward to Vinnitza Region (Likharev, 


Vestia turgida turgida (Ziegler in Rossmässler, 1836) T 

Clausilia turgida Ziegler – Rossmässler, 1836a: 20-21, pl. 12, fig. 190. 



● Distribution: Ukraine: Carpathians (Likharev, Schileyko, MS). 

Vestia turgida procera (Bielz, 1853) T 

Clausilia procera Bielz, 1853: 123. 

● Type locality: Tziblesh Mount, North Rumania. 


● Distribution: Ukraine: East Carpathians (Likharev, Schileyko, MS). 

FERUSSACIIDAE Bourguignat, 1883 

Cecilioides Férussac, 1814 

Type species: Buccinum acicula Müller, 1774 (SD Watson, 1828) 

Cecilioides acicula (Müller, 1774) T 

[= Caecilianella acicula var. liesvillei Bourguignat, 1856; Cionella acicula var. nodosaria Boettger, 1879; 

Cionella nautica Westerlund, 1887; Caecilianella acicula var. obtusata Westerlund, 1887; Caecilianella 

praeclara Westerlund, 1898; Caecilianella retteri Rosen, 1901; Cecilioides acicula var. abchasica Retowski, 

1914] 

Buccinum acicula Müller, 1774: 150-151. 

● Type locality: Thangelstedt (Thuringia, Germany). 


● Distribution: mountain Crimea, Caucasus, Central Asia from Kopet Dagh to Pamiro-Alai; occurrence in 

Ukrainian Carpathians and on Russian Plain not proved (Likharev, Schileyko, MS); Ukraine: Ciscarpathia 

and Transcarpathia (Sverlova, 2006). 

Cecilioides raddei (O. Boettger, 1879) T 

Cochlicopa (Hohenwartiana) raddei Boettger, 1879a: 25, pl. 1, fig. 8. 

● Type locality: «Borshom, Mamudly» (100 miles southwestward of Tbilisi, Georgia). 


● Distribution: Crimea, Caucasus (Likharev, Schileyko, MS); Odessa Region of Ukraine (Sverlova, 2006). 

OLEACINIDAE H. et A. Adams, 1855 

Poiretia Fischer, 1887 

Type species: Bulimus algirus Bruguière, 1792 (monotypy) 

● Remark: Likharev and Rammelmeyer (1952) mentioned also the second Recent species in the fauna of 

Caucasus: P. roseni Lindholm, 1924. However, judging from the syntype in ZIN collection, the species was 

described from fossil shells.

Poiretia mingrelica (O. Boettger, 1881) T 

Glandina algira var. mingrelica Boettger, 1881a: 170, pl. 7, fig. 1. 

● Type locality: Kutaisi (Georgia). 

● Lectotype (Subai, 1980): SMF No. 70590. 

● Distribution: Georgia: Abkhazia (Gagry, Novyi Afon, Pitzunda, middle valley of Kodor River to 

Verkhnyaya Tzebelda), Mingrelia (Tzkhenistzkhali River valley near Gordi village, Tzageri, vicinities of Kutaisi) 


PUNCTIDAE Morse, 1864 

Punctum Morse, 1864 

Type species: Helix minutissima Lea, 1841 (OD) 

Punctum boreale Pilsbry et Hirase, 1905 T 

Pilsbry, Hirase, 1905: 717. 

● Type locality: Akkeshi, Kushiro (Hokkaido, Japan). 


● Distribution: Shikotan Island (South Kurile Islands) (Likharev, 1957; Pearce et al., 2002). 

Punctum conspectum (Bland, 1865) T 

[= Helix lepta Westerlund, 1883; Vallonia patens sensu Likharev et Rammelmeyer, 1952, non Reinhardt, 

1883] 

Helix conspectum Bland, 1865: 163, fig. 7. 

● Type locality: San Francisco, California. 


● Distribution: South Kamchatka, Simushir and Kunashir Islands (Kurile Islands), Primorye Territory (Likharev, 

Schileyko, MS; Pearce et al., 2002); Sakhalin (Prozorova, Berezhok, 2004). 

Punctum micropleuros (Paget, 1854) T 

[= Helix pusilla R. Lowe, 1831, non Helix pusilla Vallot, 1801; Patula lederi O. Boettger, 1880] 

Helix micropleuros Paget, 1854: 454. 

● Type locality: "prope Montpellier". 


● Distribution: Caucasus: Talysh and Lenkoran lowland; Hissar Ridge (Izzatullaev, 1975; as Vallonia patens 

non Reinhardt, 1883). 

Punctum pygmaeum (Draparnaud, 1801) T 

Helix pygmaea Draparnaud, 1801: 93. 



● Distribution: mountain and plain areas of temperate latitudes of Eurasia: from Atlantic coast to Kamchatka, 

Sakhalin and Kurile Islands (Likharev, Schileyko, MS). 

● Remark: record of Striatura chishimana (Pilsbry et Hirasei, 1904) from Kunashir Island, Kuriles (by Pearce 

et al., 2002) probably belongs to P. pygmaeum (Prozorova, Bogatov, Kavun, 2006). 

Punctum ussuriense Likharev et Rammelmeyer, 1952 T 

Punctum pygmaeum ussuriense Likharev, Rammelmeyer, 1952: 256, fig. 182. 

● Type locality: Ussuri Natural Reserve (Primorye Territory). 


● Distribution: southern Primorye Territory (Likharev, Schileyko, MS); Sakhalin (Prozorova, Berezhok, 

2004).

HELICODISCIDAE Baker, 1927 

Helicodiscus Morse, 1864 

Type species: Helix lineatus Say, 1817 (non Helix lineata Olivi, 1792, = Planorbisparallelus Say, 1821) 

(monotypy) 

Helicodiscus singleyanus (Pilsbry, 1890) T 

Zonites singleyanus Pilsbry, 1890: 84. 

● Type locality: "New Braunfels" [Colorado]. 

●? Holotype: ANSP No. 60058. 

● Distribution: Ukraine: vicinities of Vinogradov City, Zakarpatskaya Region (Sverlova, 2006) 

(probably introduced). 

DISCIDAE Thiele, 1931 

Discus Fitzinger, 1833 

Type species: Helix ruderata Férussac, 1821 (SD Gray, 1847) 

Discus depressus (A. Adams, 1868) T 

Helix (Patula) depressus A. Adams, 1868: 467. 

● Type locality: " Vladimir Bay" (350 km NW of Vladivostok) 


● Distribution: basin of midle Amur, Primorye Teritory, Sakhalin, Kamchatka, Commander and Kurile Islands, 

northern Khabarovsk Territory (lower Amur, Ayan, Stanovoi Ridge), vicinities of Baikal Lake (Likharev, 


Discus perspectivus (Megerle von Mühlfeld, 1816) T 

[= Helix solaria Menke, 1830] 

Helix perspectivus Megerle von Mühlfeld, 1816: 11. 

● Type locality: Austria, Danube River deposits near Wien. 


● Distribution: Ukraine (Carpathians, Transcarpathia and Ciscarpathia, western Podolsk Hills, area between 

Prut and Dnieper rivers) (Likharev, Schileyko, MS; Sverlova, 2006). 

Discus rotundatus (Müller, 1774) T 

Helix rotundatus Müller, 1774: 29-30. 

● Type locality: Denmark, Norway. 


● Distribution: Baltic countries (Lithuania east of Kaunas, western Latvia, Moozund Archipelago), Kaliningrad 

Region, western regions of Byelorussia and Ukraine (not found in Carpathians), mountain Crimea (one 

finding, probably introduced) (Likharev, Schileyko, MS; Sverlova, 2006). 

Discus ruderatus (Férussac, 1821) T 

[= Helix pauper Gould, 1858; ? Helix cronkhitei Newcomb, 1865; Patula ruderata var. conica Bąkowski et 

Lomnicki, 1892] 

Helix ruderatus Férussac, 1821: 40. 

● Type locality: «Les Hautes-Alpes du Valais, ... près des glaciers du Mt. Pleureur». 


● Distribution: East European Plain, Siberia (including Chukchi Peninsula, Kamchatka, northern Khabarovsk 

Territory), Carpathians, Crimea, Caucasus, Altai, Sayan, vicinities of Baikal Lake (Likharev, Schileyko, MS); 

Kurile Islands (Pearce et al., 2002); Sakhalin (Prozorova, Berezhok, 2004).

ZONITIDAE Mörch, 1864 

PRISZTILOMATINAE Cockerell, 1891 

Hawaiia Gude, 1911 

Type species: Helix kawaiensis L. Pfeiffer, 1855 (= Helix minuscula Binney, 1840) (monotypy) 

Hawaiia minuscula (Binney, 1840) T 

Helix minuscula Binney, 1840: 435, pl. 22, fig. 4. 

● Type locality: Ohio, USA. 

● Lectotype (Pilsbry, 1946): ANSP No. 74416a. 

● Distribution: southern Primorye Territory (Likharev, Schileyko, MS). 

Pristiloma Ancey, 1887 

Type species: Zonites stearnsi Bland, 1875 (SD Baker, 1930) 

Pristiloma japonica Pilsbry et Hirase, 1903 T 

[= Pristiloma arcticum sensu Likharev, 1963 non Lehnert, 1884] 

Pilsbry, Hirase, 1903: 79. 

● Type locality: «Toya, Kuziro, in eastern Yesso» (Hokkaido, Japan). 


● Distribution: southern Kamchatka, northern (Kuroda, Koba, 1933) and southern (Likharev, 1957) Kurile 

Islands, southern Sakhalin (Likharev, Schileyko, MS). 

Striatura Morse, 1864 

Type species: Striatura milium Morse, 1864 (monotypy) 

Striatura aperta (Pilsbry et Hirase, 1904) T 

Punctum apertum Pilsbry, Hirase, 1904b: 5. 

● Type locality: Nemuro (Hokkaido, Japan). 


● Distribution: Iturup and Kunashir Islands (Kurile Islands) (Likharev, Schileyko, MS; Pearce et al., 2002). 

Vitrea Fitzinger, 1833 

Type species: Glischrus (Helix) diaphanus Studer, 1820 (monotypy) 

Vitrea angystropha (O. Böttger, 1880) T 

[= Crystallus lantzi Lindholm, 1913] 

Hyalinia angystropha O. Böttger, 1880b: 380. 


● Lectotype (Pinter, 1972): SMF No. 170983. 

● Distribution: forests of Transcaucasia, except for Talysh and Lenkoran lowland (Likharev, Schileyko, MS). 

Vitrea contortula (Krynicki, 1837) T 

[= Vitrea viridis Westerlund, 1897] 

Helix contortula Krynicki, 1837: 51 (nom. nov. pro Helix contorta Krynicki, 1836). 

● Type locality: «Cauc.» (Pyatigorsk – from labels). 


● Distribution: Caucasus, floodland forests of Krasnodar and Stavropol regions (Likharev, Schileyko, MS).

Vitrea contracta (Westerlund, 1871) T 

[= Hyalinia abchasica Retowski, 1914; Vitrea botterii sensu Likharev et Rammelmeier, 1952 non L. Pfeiffer, 

1853] 

Zonites crystallina var. contracta Westerlund, 1871: 56. 

● Type locality: «Ronneby, Blekinge» [Sweden] 

● Lectotype (Waldén in Riedel, 1966): NMG. 

● Distribution: Baltic countries, Valdai Hills, Carpathians, Ukraine, mountain part of Crimea, sporadically in 

Caucasis (Likharev, Schileyko, MS; Sverlova, 2006). 

Vitrea crystallina (Müller, 1774) T 

[= Vitrea podolica Clessin, 1880] 

Helix crystallina Müller, 1774: 23, 215. 



● Distribution: Carpathian (Sverlova, 2006); western, northwestern and central parts of East-European Plain. 

Records from mountain forests of the Crimea (Jaeckel, 1950; Likharev, Rammelmeier, 1952) are doubtful 


Vitrea nadejdae Lindholm, 1926 T 

Lindholm, 1926: 69. 

● Type locality: mountain Crimea near New Simeiz. 


● Distribution: sporadically found around New Simeiz near Yalta (Crimea) (Likharev, Schileyko, MS). 

Vitrea praetermissa Riedel, 1988 T 

Riedel, 1988a: 73, figs. 1-3. 

● Type locality: Adzharia, Zelenyi Mys near Batumi. 



Vitrea pygmaea (O. Böttger, 1880) T 

[Vitrea etrusca sensu Lindholm, 1926 non Paulucci, 1878] 

Hyalinia pygmaea O. Böttger, 1880a: 118, pl. 4, fig. 8. 

● Type locality: Martkopi near Tbilisi. 

● Lectotype (???): SMF No. 171023/1. 

● Distribution: mountain forests of the Crimea, Central Ciscaucasia (Kalitina, 1953, 1954), Central and Eastern 

Transcaucasia, Western Kopet Dagh. Records from western part of Northern Caucasis (Likharev, Rammelmeier, 

1952) are erroneous (Likharev, Schileyko, MS). 

Vitrea rhododendronis Riedel, 1966 T 

[= Vitrea diaphana erjaveci sensu Riedel, 1959 non Brusina, 1870] 

Riedel, 1966: 45-47, figs. 11-13. 

● Type locality: near Lentekhi, Georgia. 



Vitrea sorella (Mousson, 1863) T 

Zonites sorella Mousson, 1863: 299-300. 

● Type locality: debris or Rioni River. 


● Distribution: type locality (western Transcaucasia, based on the description of the species from the Rioni 

debris) (Likharev, Schileyko, MS).

Vitrea subrimata (Reinhardt, 1871) T 

Hyalina subrimata Reinhardt, 1871: 39. 

● Type locality: «Tschechoslowakei, Mährisches Gesenke, Kessel». 

● Lectotype (Pinter, 1972): SMF No. 3482/1. 

● Distribution: a single finding in Ukraine, Lvov Region, near Boleslav (Bąkowski, Łomnicki, 1892). 

Vitrea transsylvanica (Clessin, 1877) T 

Hyalina transsylvanica Clessin, 1877b: 133, pl. 2, fig. 12. 



● Distribution: Ukraine: forest regions of Carpatian Mountains, Ciscarpathia (Likharev, Schileyko, MS; Sverlopva, 

2006). 

ZONITINAE Mörch, 1864 

Aegopinella Lindholm, 1927 

Type species: Helix pura Alder, 1830 (OD) 

Aegopinella epipedostoma (Bourguignat in Fagot, 1879) T 

[= Aegopinella epipedostoma iuncta Hudec, 1964] 

Zonites epipedostoma Bourguignat – Fagot, 1879: 286. 

● Type locality: «Quartier de Caraman (commune d’Avignolet)» (France). 


● Distribution: Ukrainian Carpathians, Novogrudsk Hills (Baranovichi district) (Likharev, Schileyko, MS). 

Aegopinella minor (Stabile, 1864) T 

[= Hyalina stauropolitana Rosen, 1901a] 

Hyalina nitens var. minor Stabile, 1864: 31. 

● Type locality: not stated (Piedmont, Northern Italy – from title). 


● Distribution: deciduous forests and forest-steppe of Novogrudsk, Middle Russian, Pridneprovye, and Voluno-Podolsk 

hills; foothills of Ukrainian Carpathians; Ukranian Polesye and Dnepropetrovsk Region; mountain 

Crimea; separate heights of North Caucasus. Northermost records are from Svyantzen Hills near Naroch 

Lake (Minsk Region), vicinities of Smolensk, and near Senezhskoe Lake in vicinities of Klin (Moscow Region) 

(Likharev, Schileyko, MS; Sverlova, 2006)). 

● Remark: Examination of respective collection has shown that all records of Ae. nitidula (Draparnaud) and 

Ae. nitens (Michaud) refer to Ae. minor (Elsky, 1862; Retowski, 1888, 1914; Lindholm, 1901; Rosen, 1901b, 

1903, 1911, 1914; Puzanov, 1925, 1927), as well as Krynicki's (1836) indication of the presence of Helix 

nitidosa Férussac (nom. nudum) in vicinities of Kharkov and Volyn. 

Aegopinella nitens (Michaud, 1831) T 

Helix nitens Michaud, 1831: 44. 

● Type locality: "Dep. Ain, Umgebung von Nantua" [France], 

● Lectotype (Forcart, 1959b): Musée d'Histoire Naturelle, Lyon. 

● Distribution: Ukraine: Carpathians and Prut river basin (Sverlova, 2006). 

Aegopinella nitidula (Draparnaud, 1805) T 

Helix nitidula Draparnaud, 1805: 117. 

● Type locality: France, department Pas-de-Calais, Forêt de Guines, 3 km SE of Guines. 

● Neotype (Gittenberger, 1993): NHMV No. 86934. 

● Distribution: certainly known only from a park in Lvov (Ukraine) (Sverlova, 2006).

Aegopinella pura (Alder, 1830) T 

[= Helix lenticula Held, 1837] 

Helix pura Alder, 1830: 37. 

● Type locality: Northumberland, England. 


● Distribution: East-European Plain, Crimea, Stavropol Hills, Caucasus exept for Adzharia (Likharev, 

Schileyko, MS); Carpathians (Sverlova, 2006). Records from Murmansk and Western Siberia (Likharev, 

Rammelmeier, 1952) are erroneous (Likharev, Schileyko, MS). 

GODWINIINAE C.M. Cooke, 1921 

Perpolita Baker, 1928 

Type species: Helix hammonis sensu Baker, 1928, non Strøm, 1765 [= Helix electrina Gould, 1841] (OD) 

Perpolita hammonis (Strøm, 1765) T 

[= Helix viridula Menke, 1830; Helix radiatula Alder, 1830] 

Helix hammonis Strøm, 1765: 435, pl. 6, fig. 16. 

● Type locality: Norway. 


● Distribution: forests of East-European Plain, Caucasus and Siberia (Likharev, Schileyko, MS); Carpathians 

(Sverlova, 2006), Kurile Islands (Pearce et al., 2002); Sakhalin (Prozorova, Berezhok, 2004). 

Perpolita petronella (L. Pfeiffer, 1853) T 

Helix petronella L. Pfeiffer, 1853a: 95. 

● Type locality: Swiss Alps. 

● Lectotype (Forcart, 1960): BMNH. 

● Distribution: forests of East-European Plain, Caucasus and Siberia (Likharev, Schileyko, MS). 

OXYCHILINAE Hesse, 1927 

Cellariopsis A. Wagner, 1915 

Type species: Schistophallus deubeli A. Wagner, 1915 [= Hyalinia cellaria var. orientalis Clessin, 1887] 

(monotypy) 

Cellariopsis orientalis (Clessin, 1887) T 

[= Schistophallus deubeli A. Wagner, 1915] 

Hyalina cellaria var. orientalis Clessin, 1887: 69. 

● Type locality: «Ungarn im Tatragebirge» [Slovak Tatra Mountains]. 


● Distribution: Ukrainian Carpathians, western Podolsk Hills and area between Prutt and Dnieper rivers 

(Likharev, Schileyko, MS; Sverlova, 2006). 

Conulopolita O. Boettger, 1879 

Type species: Hyalinia (Conulopolita) raddei O. Boettger, 1879 (monotypy) 

Conulopolita cavatica (Riedel, 1966) T 

[= Oxychilus cochlospeleus Tzvetkov in Birshtein, 1950, nom. nud.] 

Oxychilus (Conulopolita) cavaticus Riedel, 1966: 188-191, figs. 196-201, pl. 5, fig. 52-54. 

● Type locality: Khosta, cave near Shirokii Pokos. 

● Holotype: ZIN

● Distribution: caves near Khosta, Adler, Gudauta, Sukhumi (Riedel, 1966). 

Conulopolita raddei (O. Boettger, 1879) T 

Hyalinia (Conulopolita) raddei O. Boettger, 1879c: 97-98, pl. 2, fig. 1. 

● Type locality: «in einer Stalaktitenhöhle in Abchasien (Kaukasus)» (a stalactite cave in Abkhazia). 


● Distribution: different caves on western Abkhazia (Gudauta, Sukhumi, Tzebelda) (Likharev, Schileyko, 

MS). 

Conulopolita sieversi (O. Boettger, 1879) T 

[= Hyalinia cellaria var. subapertus O. Boettger, 1879] 

Hyalinia cellaria var. sieversi Boettger, 1879b: 394, pl. 10. fig. 8. 

● Type locality: «Ratscha» (Rachinsky Ridge, western Georgia). 

● Lectotype (Riedel, 1966): SMF No. 3498. 

● Distribution: mountain-forest and subalpine areas of central part of North Caucasus (North Osetia), Daghestan, 

central and eastern Transcaucasia; westward of Suram Ridge: isolated records from Tzebelda 

(Abkhazia), Omalo and Notanebi (Adzharo-Imereti Ridge) (Likharev, Schileyko, MS); Armenia: forest zone 

of northern districts, Zangezur, locally in lower alpine zone, as well as Amasi, Artik, and Sevan districts (Akramowski, 

1976). 

Conulopolita ? stopnevichi (Rosen, 1925) T 

Vitrea stopnevichi Rosen, 1925: 5, fig. 1. 

● Type locality: «Mons Schoana» (Shoan Mountain, Batalpashinsky distict of Kuban Region). 


● Distribution: found on 5 localities of western Great Caucasus, west and northwest of Elbrus (Likharev, 


Discoxychilus Riedel, 1966 

Type species: Discoxychilus lindholmi Riedel, 1966 (OD) 

Discoxychilus lindholmi Riedel, 1966 T 

Riedel, 1966: 227, 284 (nom. nov. pro Patula goctschana Lindholm, 1922, non P. ruderata var. Gorktschaana 

Mousson, 1873, non P. p. Gortschkaana Mousson, 1876, nec P. goctschaana O. Boettger, 1881). 

● Type locality: «Adzarischkali» (Adzhariskali) near Batumi. 

● Lectotype (Riedel, 1966): IZP. 

● Distribution: several places in lower reaches of Chorokh River and vicinities of Batumi (Likharev, 


Eopolita Pollonera, 1916 

Type species: Zonites aequatus Mousson, 1854 [= Eopolita protensa (Férussac, 1886)] (monotypy) 

Eopolita derbentina (O. Boettger, 1886) T 

[= Hyalinia siraphora Westerlund, 1897] 

Hyalinia (Polita) derbentina Boettger, 1886b: 130-131, pl. 3, fig. 3a-d. 

● Type locality: vicinities of Derbent. 


● Distribution: foothills of eastern offshoot of Great Caucasus near Derbent and Makhachkala, northern and 

eastern offshoot of Lesser Caucasus, Zangezur and central part of Armenia (Likharev, Schileyko, MS). 

Morlina A. Wagner, 1915 

Type species: Helix glabra Rossmässler, 1835 (SD Lindholm, 1927)

Morlina glabra striaria (Westerlund, 1881) T 

Zonites glabra var. striaria Westerlund, 1881: 52. 

● Type locality: «Transsilvania (Michelsberg prope Hermannstadt...), Polonia (Ojcow...)». 


● Distribution: Ukrainian Carpathians, Ciscarphatia, western and northwestern Ukraine (Likharev, Schileyko, 

MS; Sverlova, 2006). 

Oxychilus Fitzinger, 1833 

Type species: Helix cellaria Müller, 1774 (SD Herrmannsen, 1847) 

Oxychilus alliarius (Miller, 1822) T 

Helix alliarius Miller, 1822: 379. 

● Type locality: Bristol (England). 


● Distribution: Khiyumaa and Saaremaa islands in the Baltic Sea, two greenhouses in St. Petersburg (Likharev, 


Oxychilus caspius (O. Boettger, 1880) T 

[= Hyalinia schmidti Rosen, 1914] 

Hyalinia (Polita) caspia O. Boettger, 1880b: 379. 

● Type locality: Khamarat, Talysh. 


● Distribution: forests of Lenkoran lowland and Talysh, South-Caspian lowland (Likharev, Schileyko, MS). 

Oxychilus cellarius (Müller, 1774) T 

Helix cellaria Müller, 1774: 28-29. 

● Type locality: Fridrichsdal (vicinities of Copenhagen, Denmark). 


● Distribution: Kalinigrad Region, Lithuania (Kaunas), Latvia (Riga), Estonia (Glinta and Saaremaa Islands), 

Pulkov Hills near St.-Petersburg (Likharev, Schileyko, MS). 

Oxychilus crenimargo (Retowski, 1889) T 

Hyalinia crenimargo Retowski, 1889: 231. 

● Type locality: «Risa» (Rize, northeastern Turkey). 

● Lectotype (Riedel, 1966): IZP. 

● Distribution: vicinities of Batumi (Botanical Garden, Chaisubany) (Likharev, Schileyko, MS). 

Oxychilus decipiens decipiens (O. Boettger, 1886) T 

Hyalina decipiens Boettger, 1886b: 131-132, pl. 3, fig. 1. 

● Type locality: Shakh-Dagh Mount, north-eastern Azerbaijan. 


● Distribution: forests of Great Caucasus and Borzhomi district in Transcaucasia (Likharev, Schileyko, MS). 

Oxychilus decipiens adsharicus Riedel, 1966 T 

Riedel, 1966: 89-91, figs. 68-72. 

● Type locality: botanical garden, Zelenyi Mys near Batumi. 


● Distribution: Transcaucasia: vicinities of Batumi (Likharev, Schileyko, MS). 

Oxychilus deilus deilus (Bourguignat, 1857) T 

[= Hyalinia krynickii Clessin, 1883; Helix filicum Krynicki, 1836 part.] 

Zonites deilus Bourguignat, 1857: 9, pl. 8, figs. 7-9. 

● Type locality: vicinities of Sevastopol.


● Distribution: southern coast of Crimea (Likharev, Schileyko, MS); introduced to Askania-Nova Natural 

Reserve in Kherson Region (Sverlova, 2006). 

Oxychilus difficilis (O. Boettger, 1888) T 

Hyalinia (Retinella) difficilis Boettger, 1888: 150-152. 

● Type locality: Oschten-Fischt Mountains, valley of Kurdzhips River, Guk mountain (western Great Caucasus). 

● Syntypes: SMF Nos. 225580 ("lectotype"), 225581/1, 225582/1, 225583/10 juv. ("paralectotypes"). 

● Distribution: mountain forests of western Great Caucasus; upper reaches of Belaya River (Ciscaucasia); 

from Sochi and Mzymta river basins to Psyrckha River basin (Transcaucasia) (Likharev, Schileyko, MS). 

Oxychilus disciformis Riedel, 1959 T 

Oxychilus caspius disciformis Riedel, 1959: 71, figs. 1, 2, pl. 1, figs. 1-3. 

● Type locality: Northern Iran, Ghilan province. 


● Distribution: forests of Lenkoran lowland and Talysh, South-Caspian lowland (Likharev, Schileyko, MS). 

Oxychilus discrepans (Retowski, 1889) T 

[= Hyalinia difficilis var. batumensis Retowski, 1914] 

Hyalinia difficilis var. discrepans Retowski, 1889: 233. 

● Type locality: «Adsharizkal» (Adzhariskali near Batumi). 

● Syntype: IZP. 

● Distribution: Georgia: Adzharo-Imereti Ridge (to Kobuleti in the north, to Adigeni in Adzhariskali River 

valley in the East, to Chorokh River valley in the south) (Likharev, Schileyko, MS). 

Oxychilus draparnaudi (Beck, 1837) T 

Helicella draparnaudi Beck, 1837: 6 (nom. nov. pro Helix lucida Draparnaud, 1801, non Pultney, 1799). 



● Distribution: sporadically near Moscow, St.-Petersburg, Kharkov, Uzhgorod, and Lvov (Likharev, 


Oxychilus duboisi (Charpentier in Mousson, 1863) T 

Zonites duboisi Charpentier – Mousson, 1863: 295-296. 

● Type locality: Kutaisi. 


● Distribution: Kutaisi (ruins of Bagrat Castle and neighborhood) (Likharev, Schileyko, MS). 

Oxychilus elegans (O. Boettger, 1881) T 

Hyalinia (Mesomphix) elegans O. Boettger, 1881a: 197-198, pl. 8, fig. 12a-c. 

● Type locality: forests of Lenkoran. 


● Distribution: forests of Lenkoran lowland and Talysh mountains (Likharev, Schileyko, MS). 

Oxychilus emmae (Akramowski, 1955) T 

Vitrea emmae Akramowski, 1955: 149, fig. 1. 

● Type locality: Laki ravine near Verin-Agdan village, Idzhevan district of Armenia. 


● Distribution: forests of Lesser Caucasus from Akhaltzikhe to Idzhevan (southern Georgia and northern Armenia) 

(Likharev, Schileyko, MS).

Oxychilus filicum (Krynicki, 1836) T 

Helix filicum Krynicki, 1836: 201. 

Type locality (restr. Riedel, 1966): Lenkoran. 


● Distribution: Talysh and Lenkoran lowland (Likharev, Schileyko, MS). 

Oxychilus horsti (O. Boettger in Reibisch, 1892) T 

[= Hyalinia ampliata Rosen, 1911; Hyalinia duboisi (non Mousson, 1863) et Hyalinia sucinacia (non O. 

Boettger, 1883) auct.] 

Hyalinia horsti O. Boettger – Reibisch, 1892: 50. 

● Type locality: valley of Belaya River. 


● Distribution: western part of Great Caucasus (Likharev, Schileyko, MS). 

Oxychilus imperator Riedel, 1966 T 

[= Schistophallus duboisi auct., non Mousson, 1863] 

Riedel, 1966: 161, figs. 165-167, pl. 3, figs. 31-36. 

● Type locality: valley of Psyrtzkha river, Novyi Afon, Abkhasia. 


● Distribution: Abkhasia (Likharev, Schileyko, MS). 

Oxychilus inopinatus (Ulicný, 1887) T 

[Hyalina opinata Ulicný in Clessin, 1887 (nom. err.); Hyalinia plutonia Kimakowicz, 1890] 

Hyalina inopinata Ulicný, 1887: 111, fig. 36/1. 

● Type locality: «Pavlovske vrchy», southern Moravia, Czechia. 


● Distribution: Volyn-Podolsk Hills, Ukraine (Likharev, Schileyko, MS). 

Oxychilus iphigenia (Lindholm, 1926) T 

Hyalina iphigenia Lindholm, 1926: 167. 

● Type locality: Sredniye Lemeny village near Novyi Simeiz, western part of South coast of Crimea. 



Oxychilus kobelti (Lindholm, 1910) T 

Retinella kobelti Lindholm, 1910: 223. 

● Type locality: ravine of Savluk-Su River, Alma River basin (north-western offshoot of Babugan-Yaila 

Ridge, Crimea). 

● Lectotype (Riedel, 1972): ZIN. 

● Distribution: Mountain Crimea: forest regions of Babugan-Yaila and foothills of Chatyr-Dagh. One finding 

on Southern coast near Aidanila village (near Nikitsky Botanical Garden) (Likharev, Schileyko, MS). 

Oxychilus koutaisanus koutaisanus (Mousson, 1863) T 

[= Hyalinia kutaisana auct.; Hyalinia kutaisiana auct.; ? Zonites lucidus var. selectus Mousson, 1863; Hyalinia 

kutaisiana var. transitans O. Boettger, 1883 part.] 

Zonites cypricus var. koutaisanus Mousson, 1863: 295. 



● Distribution: central and southern Georgia (Likharev, Schileyko, MS). 

Oxychilus koutaisanus mingrelicus (Mousson, 1863) T 

[Hyalina mingrelica var. intermissa Kobelt, 1879; Hyalinia pontica O. Boettger, 1881Hyalinia kutaisiana 

var. transitans O. Boettger, 1883 part.; Hyalinia reticulata O. Boettger, 1883]

Zonites mingrelicus Mousson, 1863: 294. 

● Type locality: «atterrissements du Rhéon, ... Koutais» (Rioni River, Kutaisi). 


● Distribution: West Caucasus, from basin of Psekups River to basin of Laba River in Ciscaucasia, from Sochi 

to Abkhasia and Mingrelia in Transcausasia. Absent in southern Georgia (Likharev, Schileyko, MS). 

Oxychilus oschtenicus (O. Boettger, 1888) T 

Hyalinia (Polita) oschtenica Boettger, 1888: 150. 

● Type locality: forests of Oschten-Fischt Mountains on West Great Caucasus. 

● Syntype: SMF No. 160942. 

● Distribution: western part of Great Caucasus; from Kurdjips River to Dakhovsky ravine (Belaya River) in 

Ciscaucasia; from Sochi (Khosta) to valley of Kodori River and off Tkvarcheli (Abkhasia) in Transcaucasia 


Oxychilus suaneticus suaneticus (O. Boettger, 1883) T 

Hyalinia suanetica O. Boettger, 1883: 148, pl. 5, fig. 3. 

● Type locality: Svanetia. 

● Syntypes: SMF Nos. 225695 ("lectotype"), 225696/1+3 juv., 225697/1 ("paralectotypes". 

● Distribution: western part of Great Caucasus (Likharev, Schileyko, MS). 

Oxychilus suaneticus likharevi (Riedel, 1966) T 

Schistophallus suaneticus likharevi Riedel, 1966: 166, figs. 168-171, pl. 2, figs. 25-27. 

● Type locality: Khosta, Yew-Boxwood copse. 



Oxychilus subeffusus (O. Boettger, 1879) T 

[= Hyalinia effusa O. Boettger, 1879 non Helix effusa L. Pfeiffer, 1866; Hyalinia subeffusa var. depressa O. 

Boettger, 1881; Hyalinia subeffusa var. daghestanica O. Boettger, 1886] 

Hyalinia (Vitrea) subeffusa Boettger, 1879b: 395-396 (nom. nov. pro Hyalina effusa O. Boettger, 1879). 

● Type locality: «Mamudly», southern Georgia. 


● Distribution: Great Caucasus northward to Zheleznovodsk, central and south Georgia, Armenia, northeastern 

Lesser Caucasus to Talysh (Likharev, Schileyko, MS). 

Oxychilus sucinacius sucinacius (O. Boettger, 1883) T 

[= Hyalinia denselineata P. Hesse, 1914; Hyalinia sericata Lindholm, 1922] 

Hyalinia sucinaceus O. Boettger, 1883: 150, pl. 5, fig. 2. 

● Type locality: Nakeral mountains (southern offshoots of Rachinsky Ridge, western Georgia). 


● Distribution: forests on Rachinsky, Lechkhum, Adzharo-Imereti and Trialeti Ridges (western Georgia) 


Oxychilus sucinacius zakatalicus (Likharev et Riedel, 1962) T 

Schistophallus sucinacius zakatalicus Likharev, Riedel, 1962: 14, figs. 1, 2. 

● Type locality: Zakatal Reserve,Verketel Ridge near Kamplov cave. 


● Distribution: mountain forests of southern offshoots of central part of Great Caucasus at border between 

Georgia and Azerbaijan (Likharev, Schileyko, MS). 

Oxychilus translucidus (Mortillet, 1854) T 

[= Hyalinia komarowi O. Boettger, 1881a] 

Helix translucidus Mortillet, 1854: 9, pl. 1, fig. 4.

● Type locality: Trabzon, Turkey. 

● Syntypes: unknown. 

● Distribution: in parks and gardens of some vilages along Black sea coast of Caucasus, from Sochi to Batumi; 

parks of Moscow, Kutaisi and botanical garden in Ashkhabad; greenhouses in Moscow, Rostov-on- 

Don, Alma-Ata, Dushanbe and Tashkent (Likharev, Schileyko, MS). 

Oxychilus (?) diaphanellus (Krynicki, 1837) T 

[= Helix diaphana Krynicki, 1833 nom. praeocc. non Studer, 1820; Hyalina taurica Clessin, 1881b; Hyalina 

planaria Clessin, 1883; Hyalinia zonulata Westerlund, 1886] 

Helicella diaphanella Krynicki, 1837: 53. 

● Type locality: «Taur.» (Crimea between Skel and Merdven – in Krynicki, 1833 and on label). 

● Syntypes: ZIN, MNHN. 

● Distribution: mountain forests of Main Ridge, presumably Yajla and Chatyr-Dagh and Southern Coast, 

Crimea (Puzanov, 1927); Kharkov, Zaporozhje and Kiev (Sverlova, 2006). 

«Oxychilus» andronakii (Lindholm, 1914) T 

Hyalinia andronakii Lindholm, 1914a: 33. 

● Type locality: Kwarzchana», 8 km north from Artvin (northeastern Turkey). 

● Lectotype ZIN (?) 

● Distribution: Chorokh River basin (Likharev, Schileyko, MS). 

«Oxychilus» birsteini Tzvetkov, 1940 T 

Tzvetkov, 1940b: 57, fig. 1. 

● Type locality: «cave in Andreevka», Abkhasia. 

● Syntypes: ZMMU Nos. Lc-14469, Lc-14470. 

● Distribution: Nizhnyaya Andreevskaya cave near Akhalsheny (27 km north from Sukhumi) (Birstein, 

1940). 

«Oxychilus» lederi (O. Boettger, 1880) T 

Hyalinia (Polita) lederi O. Boettger, 1880a: 117-118, pl. 4, fig. 2. 

● Type locality: Tbatani, northeastern Georgia. 


● Distribution: a single record from upper reaches of Alazani River, 2100 m above see level (Likharev, 


«Oxychilus» retowskii (Lindholm, 1914) T 

Crystallus retowskii Lindholm, 1914a: 35. 

● Type locality: Abastumani (Southern Georgia). 

● Holotype: Geological Institute of Azerbaijan Academy of Science (Baku). 

● Distribution: Lesser Caucasus from Abastumani (Adzharo-Trialeti Ridge) to Dilizhan (Bazum Ridge) (Likharev, 


Riedeliconcha Schileyko, 2003 

Type species: Hyalina depressa Sterki, 1880 (OD) 

Riedeliconcha depressa (Sterki, 1880) T 

Hyalina depressa Sterki, 1880: 104`-105. 

● Type locality: "am Randen (Jura) und im Wuhtachthal (Südost-Schwarzwald, Triasboden), also auf 

schweizerischen sowohl wie aud deutschen (Baden) Gebiete". 


● Distribution: Ukrainian Carpathians (Likharev, Schileyko, MS).

Vitrinoxychilus Riedel, 1963 

Type species: Hyalinia suturalis O. Boettger, 1881 (OD) 

Vitrinoxychilus subsuturalis (O. Boettger, 1888) T 

Hyalinia subsuturalis Boettger, 1888: 149-150. 

● Type locality: forests on Kurdjips River valley (tributary of Belaya River). 

● Syntypes: SMF No. 165895, MNHN. 

● Distribution: sporadically in western Great Caucasus: near Maikop, Kurdjips River valley, Yew-Boxwood 

copse (Khosta), Inguri River valley near Khaishi village (Likharev, Schileyko, MS). 

Vitrinoxychilus suturalis O. Boettger, 1881 T 

Hyalinia (Polita) suturalis Boettger, 1881a: 190-191, pl. 8, fig. 9a-d. 



● Distribution: Adzharo-Imereti, Suram, Rachinsky, Svanetsky and Kodor ridges of Caucasus (Likharev, 


DAUDEBARDIIDAE Kobelt, 1906 

Bilania Schileyko, 1986 

Type species: Daudebardia boettgeri Clessin, 1883 (OD) 

Bilania boettgeri (Clessin, 1883) T 

Daudebardia boettgeri Clessin, 1883: 38, pl. 2, fig. 9, 10. 

● Type locality: Strateiz near Yalta, Crimea. 


● Distribution: montain forests of Crimea (Likharev, Schileyko, MS). 

Carpathica A. Wagner, 1895 

Type species: Daudebardia (Carpathica) kimakowiczi A. Wagner, 1895 [= Daudebardia calophana Westerlund, 

1881] (SD Forcart, 1950) 

Carpathica calophana (Westerlund, 1881) T 

[= Daudebardia (Carpathica) kimakowiczi A. Wagner, 1895] 

Daudebardia calophana Westerlund, 1881: 51. 

● Type locality: Przemysl, Poland. 

● Syntypes: NMG No. 13. 

● Distribution: Carpathians and near-Carpathian areas of Ukraine (Likharev, Schileyko, MS). 

Daudebardia Hartmann, 1821 

Type species: Helix rufa Draparnaud, 1805 (SD Forcart, 1950) 

Daudebardia brevipes (Draparnaud, 1805) T 

Draparnaud, 1805: 119. 

● Type locality: not designated. 


● Distribution: Ukraine – vicinities of Lvov (Sverlova, 2006). 

Daudebardia nivea Schileyko, 1988 T 

Schileyko 1988b: 1730. fig. 1 (I-IV). 

● Type locality: Georgia, Oficho Cave near Kumistavi village, 6 km from Tzkhaltubo, Caucasus. 

● Holotype: ZMMU No. Lc-14439.


Daudebardia rufa (Draparnaud, 1805) T 

[= Daudebardia haliciensis Westerlund, 1881] 

Helix rufa Draparnaud, 1805: 118-119, pl. 8, figs. 26-29. 



● Distribution: the species was recorded from Poland near Przemysl (as D. haliciensis Westerlund, 1881), i.e. 

in immediate vicinity of Ukraine (Likharev, Schileyko, MS); in Ukraine sporadically occurs in Lvov and 

Chernovtzy regions (Sverlova, 2006). 

Inguria Schileyko, 1986 

Type species: Daudebardia wagneri Rosen, 1911 (OD) 

Inguria wagneri (Rosen, 1911) T 

[= Daudebardia heydeni Riedel, 1978, part.] 

Daudebardia wagneri Rosen 1911: 94, pl. 2, fig. 5. 

● Type locality: Maikop and immediate vicinity. 


● Distribution: western offshoots of Great Caucasus, from Maikop in the north to the Tekhuri River valley in 

the south, isolated findings in western Lesser Caucasus on Godersinsky pass of Arsiyansky Ridge (Likharev, 


Sieversia Kobelt, 1880 

Type species: Daudebardia heydeni O. Boettger, 1879 (monotypy) 

Sieversia heydeni (O. Boettger, 1879) T 

[= Daudebardia sieversi O. Boettger, 1880a; Daudebardia pawlenkoi O. Boettger, 1880a] 

Daudebardia heydeni Boettger, 1879a: 3, pl. 1, fig. 1. 

● Type locality: Kazbek (erroneous locality, probably Suram Ridge – Boettger, 1880a: 112). 


● Distribution: maritime regions of West Caucasus (Likharev, Schileyko, MS). 

Sieversia lederi (O. Boettger, 1881) T 

[= Daudebardia pontica Simroth, 1912; Daudebardia caucasica Simroth, 1912; Daudebardia rufa sensu 

Reibisch, 1892 non Draparnaud, 1805] 

Daudebardia lederi Boettger, 1881a: 172-173, pl. 7, fig. 2a-b. 



● Distribution: West Ciscaucasia from Maikop to the Black Sea and to mouth of the Chorokh River in the 

south (Likharev, Schileyko, MS). 

Szuchumiella H. Wagner, 1945 

Type species: Daudebardia (Libania) jetschini A. Wagner, 1895 (SD Forcart, 1950) 

Szuchumiella jetschini (A. Wagner, 1895) T 

[Daudebardia kalischewskii Simroth, 1910 restr. Riedel, 1978; Daudebardia simrothi H. Wagner, 1945 restr. 

Riedel, 1978] 

Daudebardia (Libania) jetschini A. Wagner, 1895: 618-619, pl. 5, fig. 30a-b. 

● Type locality: Psyrtskha near Novy Afon, Abkhazia. 


● Distribution: maritime areas of West Caucasus from Gudauta to Tkvarcheli and Kodori River basin (Likharev, 

Schileyko, MS).

VITRINIDAE Fitzinger, 1833 

VITRININAE Fitzinger, 1833 

Vitrina Draparnaud, 1801 

Type species: Helix pellucida Müller, 1774 (monotypy) 

Vitrina exilis Morelet, 1858 T 

Morelet, 1858: 8. 

● Type locality: not stated (Kamchatka – from title). 


● Distribution: Iturup, Paramushir, Shumshu, Atlasova Islands (Kurile Islands) (Pearce et al., 2002). 

Vitrina pellucida pellucida (Müller, 1774) T 

[= Vitrina beryllina C. Pfeiffer, 1821 «Bellevnegarten von Cassel» restr. Forcart, 1955; Vitrina globosa O. 


Helix pellucida Müller, 1774: 15. 

● Type locality: Fridrihsdal near Copenhagen. 

Neotype (Forcart, 1955): ZMUC. 

● Distribution: Palearctic (Likharev, Schileyko, MS). 

Vitrina pellucida alaskana Dall, 1905 T 

[= Vitrina pfeifferi Newcomb, 1861 nom. praeocc., non Deshayes, 1851). 

Dall, 1905: 37 (nom. nov. pro Vitrina pfeifferi Newcomb, 1861). 

● Type locality: Karson River valley Nevada State, USA. 


● Distribution: Commander Islands (Likharev, Schileyko, MS). 

Vitrina rugulosa Martens, 1874 T 

[= Vitrina alexandri Westerlund, 1896] 

Martens, 1874: 7. 

● Type locality: Karakasuk area near Kokand. 


● Distribution: mountain regions of Central Asia (including Kopet-Dagh) and Altai (Likharev, Schileyko, 

MS). 

PHENACOLIMACINAE Schileyko, 1986 

Phenocolimax Stabile, 1859 

Type species: Helicolimax major Férussac, 1807 (SD Fischer in Paulucci, 1878) 

Phenacolimax annularis (Studer, 1820) T 

[= ? Vitrina conoidea E. Martens, 1874; Helicarion sieversi Mousson, 1876; Vitrina subconica O. Boetger, 

1878; Vitrina komarowi O. Boettger, 1879; Vitrina annularis f. caucasica Westerlund, 1896; Vitrina raddei 

O. Boettger, 1889] 

Glischrus annularis Studer, 1820: 86. 

● Type locality: «Kt. Wallis, Tourbillon bei Sitten» restr. Forcart, 1957 (Switzerland, Kt. Wallis). 

● Lectotype (Forcart, 1957): NMBE.

● Distribution: Carpathians, Crimea, central and eastern Caucasus, Talysh and mountains of Central Asia 

(Likharev, Schileyko, MS); Odessa (Sverlova, 2006). 

Trochovitrina O. Boettger, 1880 

Type species: Lampadia lederi O. Boettger (monotypy) 

Trochovitrina lederi (O. Boettger, 1879) T 

[= Vitrina lederi var. subcarinata O. Boettger, 1880] 

Lampadia lederi Boettger, 1879a: 7, pl. 1, fig. 2. 

● Type locality: Kazbek Mount. 


● Distribution: Suram Ridge, Lenkoran lowland and mountain-forest zone of Talysh (Likharev, Schileyko, 

MS). 

SEMILIMACINAE Schileyko, 1986 

Eucobresia Baker, 1929 

Type species: Vitrina diaphana Draparnaud, 1805 (OD) 

Eucobresia nivalis (Dumont et Mortillet, 1852) T 

[= Phenacolimax alpina Koch in Hesse, 1923 part; Vitrina kochi auct. (non Andreae, 1884)] 

Vitrina nivalis Dumont, Mortillet, 1852: 209-211. 

● Type locality: «Bassin de Genève. Vallée inférieure du Haut-Rhфne. Azindaz, au-dessus de Bex. Bassin de 

Bonneville. Vallée de l’Arve. Col du Bonhomme, а 2,300 m. Bassin de Moutiers. Vallée de la Haute-Isère. 

Petit-St-Bernard, а 2,000 m. Vallée du Doron. Col de la Valnoise, а plus de 3,000 m.» (Swiss Alps). 

Syntypes (Zilch, 1979): SMF No. 170130/3. 

● Distribution: Ukrainian Carpathians (Likharev, Schileyko, MS). 

Semilimax Agassiz, 1845 

Type species: Helix semilimax Férussac, 1802 (SD Hesse, 1923) 

Semilimax kotulae (Westerlund, 1883) T 

Vitrina kotulae Westerlund, 1883a: 54-55. 

● Type locality: «Galicia in M. Tatra, 900-2200’ s. m.» (Poland). 


● Distribution: Carpathians in Ukraine (Likharev, Schileyko, MS). 

Semilimax semilimax (Férussac, 1802) T 

[= Vitrina elongata Draparnaud, 1805] 

Helix semilimax Férussac, 1802: 236, pl. 1, figs. A-D. 

● Type locality: Switzerland, Billafingen near Baden. 


● Distribution: Carpathians in Ukraine (Likharev, Schileyko, MS). 

GASTRODONTIDAE Tryon, 1868 

Zonitoides Lehmann, 1862 

Type species: Helix nitida Müller, 1774 (monotypy)

Zonitoides arboreus (Say, 1816) T 

Helix arboreus Say, 1816: sp. no. 2, pl. 4 fig. 4. 

● Type locality: North America. 


● Distribution: introduced to parks and gardens of Eastern Hemisphere (Likharev, Schileyko, MS). 

Zonitoides nitidus (Müller, 1774) T 

Helix nitida Müller, 1774: 32. 

● Type locality: Fridrichsdal (vicinities of Copenhagen, Denmark). 



EUCONULIDAE H. Baker, 1928 

Discoconulus Reinhardt, 1883 

Type species: Hyalina sinapidium Reinhardt, 1877 (SD Pilsbry, 1928) 

Discoconulus sinapidium (Reinhardt, 1877) T 

Hyalina sinapidium Reinhardt, 1877c: 318, pl. 10, fig. 5. 

● Type locality: «Yedo, Uweno, Nikkobirge» (Japan). 


● Distribution: southern Primorye Territory (Likharev, Schileyko, MS). 

Euconulus Reinhardt, 1883 

Type species: Helix fulva Müller, 1774 (SD Pilsbry in Pilsbry et Ferriss, 1910) 

Euconulus fulva Müller, 1774 T 

[= Helix trochiformis Montagu, 1803; Helix fulva var. alderi Gray in Turton, 1840] 

Helix fulva Müller, 1774: 56-57. 




ARIOPHANTIDAE Godwin-Austen, 1888 

Macrochlamys Benson, 1832 

Type species: Helix (?) petrosa Hutton, 1834 (SD Blanford, Godwin-Austen, 1908) 

Macrochlamys clessini Westerlund, 1902 T 

[= Macrochlanys schmidti Clessin, 1894 nom. ppraeocc. non Brancsik, 1891; Macrochlamys kasachstani 

Tzvetkov, 1940] 

Westerlund, 1902: 19, 35 (nom. nov. pro Macrochlanys schmidti Clessin, 1894). 

● Type locality: Ketmen Ridge, gorge near Podgorny vyselok. 


● Distribution: Northern Tien-Shan (Likharev, Schileyko, MS). 

Macrochlamys kasnakowi (Westerlund, 1898) T 

[= Hyalinia maureri Rosen, 1901; Macrochlamys lindbergi Likharev et Starobogatov, 1967] 

Hyalinia kasnakowi Westerlund, 1898: 155. 

● Type locality: «Turkestan, Sary-Pu».


● Distribution: Southern Tien-Shan, Alai, Pamir, Hindu Kush (Likharev, Schileyko, MS). 

Macrochlamys sogdiana (Martens, 1871) T 

[= Macrochlamys sogdiana var. coeligena Gude, 1902] 

Helicarion sogdianus Martens, 1871: 65, pl. 1, figs. 1-3. 

● Type locality: «Turkestan, Fedchenko expedition». 

● Lectotype (Muratov, 1991): ZMMU No. Lc-1076. 

● Distribution: Tien-Shan, Pamir, Ferghana valley, valleys of Zaravshan, Syr-Daria and Amu-Daria rivers 


Macrochlamys turanica Martens, 1874 T 

[Macrochlamys schmidti Brancsik, 1891; Macrochlamys korshinskii Westerlund, 1896; Hyalinia retteri 

Rosen, 1901; Parvatella issikensis Cockerell, 1928] 

Martens, 1874: 7, pl. 1, fig. 3. 

● Type locality: «Aksay-Tau, Fedchenko expedition». 

● Lectotype (Muratov, 1991): ZMMU No. Lc-3785. 

● Distribution: Tien-Shan and northern foothills of Pamir (Likharev, Schileyko, MS). 

AGRIOLIMACIDAE H. Wagner, 1935 

Deroceras Rafinesque, 1820 

Type species: Limax gracilis Rafinesque, 1820 (= Limax laevis Müller, 1774) (monotypy) 

Deroceras agreste (Linnaeus, 1758) T 

[= Agriolimax fedschenkoi Koch et Heynemann, 1874; Agriolimax transcaucasicus coeciger Simroth, 1901] 

Limax agrestis Linnaeus, 1758: 652. 



● Distribution: almost everywhere (Likharev, Schileyko, MS). 

Deroceras altaicum (Simroth, 1886) T 

[= Agriolimax buchar var. coeciger Simroth, 1910; Agriolimax altaicus var. transitorius Waldén, 1957] 

Agriolimax altaiucus Simroth, 1886: 28, pl. 1, fig. 14. 

● Type locality: Altai. 


● Distribution: Western Tien-Shan, (Kirgiz and Zailijskij Alatau), Dzungar Ridge, Altai, Sayan, Cisbaikalia 

and Transbaikalia, Amur Region, Primorye Territory, Kamchatka, Sakhalin, Kurile Islands (Likharev, 

Schileyko, MS); 

Deroceras bakurianum (Simroth, 1912) T 

[= Agriolimax schemachensis Simroth, 1912] 

Agriolimax bakurianus Simroth, 1912: 40, pl. 3, fig. 34, pl. 7, fig. 23. 

● Type locality: Bakuriani (Georgia). 

● Lectotype (Likharev, Wiktor, 1980): ZIN. 

● Distribution: forest areas of Transcaucasia, Daghestan near Tlyarata (Likharev, Schileyko, MS); Western 

Caucasus (Sochi), Crimea (Wiktor, Jurkowska, 2007). 

Deroceras caucasicum (Simroth, 1901) T 

[= Agriolimax dymczewiczi sensu Simroth, 1901 non Kaleniczenko, 1851; Agriolimax caspius Simroth, 1901; 

Lytopelte grusina Simroth, 1901; Lytopelte caucasica armenia Akramowski, 1948; Deroceras hamatus 

Skljar, 1975] 

Lytopelte caucasicus Simroth, 1901: 171, pl. 17, figs. 5-14.

● Type locality: Lagodekhi, valley of left tributary of Alazani River. 


● Distribution: central and eastern Caucasus (Likharev, Schileyko, MS); introduced to Central Asia (Likharev, 

Wiktor, 1980) and Far East (vicinities of Vladivostok, eastward to Nakhodka, northward to Ussuriisk) 

(Chernyshev, 1999, 2006); Crimea, Donetzk City, Kiev, Lugansk, and Donetsk regions (Sverlova, 2006; 

Sverlova et al., 2009). 

Deroceras ilium (Simroth, 1901) T 

Agriolimax ilium Simroth, 1901: 159, pl. 15, figs. 29-32. 

● Type locality: Il mountain (current name: Sapitskaya Budka) near Vladikavkaz. 


● Distribution: upper reaches of the Terek, Podkumok, Kodori rivers and upper reaches of eastern tributaries 

of the Kuban River (Likharev, Schileyko, MS). 

Deroceras laeve (Müller, 1774) T 

[= Limax gracilis Rafinesque, 1820; Limax hyperboreus Westerlund, 1876; Agriolimax psudodioicus 

Velitchkovsky, 1910; Deroceras schulzi Tzvetkov et Matyokin, 1946] 

Limax laevis Müller, 1774: 1-2. 




Deroceras moldavicum (Grossu et Lupu, 1961) T 

Lytopelte moldavicum Grossu, Lupu, 1961: 28, figs. 1-2. 

● Type locality: Sukevitsa (Suchava district, northern Romania). 

● Holotype: Natural History Museum of Bucuresti No. 13166. 

● Distribution: Ukrainian Carpathians (Zakarpatskaya and Ivano-Frankovsk regions) (Likharev, Schileyko, 

MS); Lvov near-Dniester areas of Ternopol and Chernovtzy regions (Sverlova, 2006). 

Deroceras occidentalis (Grossu et Lupu, 1966) T 

Lytopelte occidentalis Grossu, Lupu, 1966: 25, figs. 1-3. 

● Type locality: Styna-de-Vale, Bikhor mountain (western Romania). 


● Distribution: south-western part of Ukrainian Carpathians (Zakarpatskaya and Ivano-Frankovsk regions) 

and Chernovtsy District (Baidashnikov, 1989). 

Deroceras osseticum (Simroth, 1901) T 

[= Agriolimax subagrestis var. minutus Simroth, 1901 part.] 

Agriolimax osseticum Simroth, 1901: 161, pl. 14, figs. 22-26. 

● Type locality: Tkibuli (Western Georgia). 


● Distribution: western part of North Caucasus and western Transcaucasia (Likharev, Schileyko, MS); Adzharia: 

Batume (Wiktor, Jurkowska, 2007). 

Deroceras praecox Wiktor, 1966 T 

Wiktor, 1966: 449, figs. 1-10, 15, 16. 

● Type locality: Pokshyvna, eastern Sudetes, Poland. 

● Holotype: Natural History Museum of Wroclav. 

● Distribution: probably western Ukraine (Likharev, Schileyko, MS). 

Deroceras reticulatum (Müller, 1774) T 

[= Agriolimax agrestis auct., non Linnaeus, 1758] 

Limax reticulatus Müller, 1774: 10.

● Type locality: Rosenburg, Fridrichsdal (Denmark). 


● Distribution: Arkhangelsk, Leningrad, Pskov, Novgorod, Yaroslavl, Moscow, Tula, Penza, Orel, Kursk, 

Saratov and Chelyabinsk regions; Tatarstan; Estonia (Tallinn); Lithuania (Shaulyai district); Ukraine (Kiev, 

Zhitomir, Odessa, Yalta, Sevastopol, Ivano-Frankovsk, Yaremcha); North Caucasus (Maikop, Vladikavkaz); 

Transcaucasia (Erevan); Kazakhstan (Alma-Ata); Central Asia (Khamzabad, Ferghana district) (Likharev, 


Deroceras rodnae Grossu et Lupu, 1965 T 

Grossu, Lupu, 1965: 28, fig. 2. 

● Type locality: Inau mountain, Rodna block, Eastern Carpathians, Romania. 


● Distribution: Ukraine: Carphathians including adjacent areas of Ciscarpathia and Transcarpathia (Sverlova, 

2006). 

Deroceras sturanyi (Simroth, 1894) T 

[= Agriolimax laevis aust., non Müller, 1774] 

Agriolimax sturanyi Simroth, 1894: 392, pl. 19, figs. 5, 6. 

● Type locality: vicinities of Ohrid Lake (Yugoslavia). 

● Lectotype (Likharev, Wiktor, 1980): NHMV No. 19493b. 

● Distribution: NW Ukraine, Carpathians, Odessa Region, Zaporozhje Region, Donetsk and Lugansk regions 

(Sverlova, 2006; Sverlova et al., 2009); sporadically in Moscow and Voronezh regions; gardens, greenhouses 

and botanical gardens of Alma-Ata (Uvalieva, 1979), St.-Petersburg, Chelyabinsk, Rostov-on-Don, Khodzhent 

and Khorog area (Likharev, Schileyko, MS). 

Deroceras subagreste (Simroth, 1892) T 

[= Agriolimax agresticulus Simroth, 1894; Agriolimax ananowi Simroth, 1901] 

Agriolimax subagrestus Simroth, 1892: 43. 

● Type locality: Maikop. 


● Distribution: North Caucasus; introduced to greenhouses of Pushkin (Leningrad Region), Rostov-on-Don 

and gardens of Tashkent (Likharev, Schileyko, MS); south-eastern Ukraine (Sverlova et al., 2009); Abkhasia 

(Wiktor, Jurkowska, 2007). 

Deroceras tauricum (Simroth, 1901) T 

[= Agriolimax dymczewiczi sensu Simroth, 1885 non Kaleniczenko, 1851; Agriolimax crimense Simroth, 

1901; Deroceras ramosus Skljar, 1975] 

Agriolimax tauricum Simroth, 1901: 153, fig. 7. 

● Type locality: montain Crimea. 


● Distribution: forest and forest-steppe areas of Crimea, mountain Crimea (Likharev, Schileyko, MS; Sverlova 

et al., 2009). 

Deroceras turcicum (Simroth, 1894) T 

Agriolimax turcicum Simroth, 1894: 392, pi. 19, figs. 2-5. 

● Type locality:"Ochrida-See". 

● Lectotype (Likharev, Viktor, 1980): NHMV No. 19491. 

● Distribution: Ukraine: Podolsk Hills, area between Prut and Dniester rivers (Sverlova, 2006); Moldavia (Baidashnikov, 

1993, 1996). 

Krynickillus Kaleniczenko, 1851 

Type species: Krynickillus melanocephalus Kaleniczenko, 1851 (SD Pilsbry, 1922)

Krynickillus melanocephalus Kaleniczenko, 1851 T 

Kaleniczenko, 1851: 221, pl. 5, fig. 2. 

● Type locality: vicinities of Stavropol. 


● Distribution: almost all Caucasus, forests of Stavropol Territory, mountain part of Crimea (Likharev, 

Schileyko, MS); Ukraine: Kiev, Lvov, vicinities of Yasinovataya town in Donetzk Region (Sverlova, 2006); 

Latvia (Wiktor, Jurkowska, 2007). 

Lytopelte O. Boettger, 1886 

Type species: Amalia longicollis O. Boettger, 1886 (= Amalia maculata Koch et Heynemann in Martens, 

1874) (monotypy) 

Lytopelte maculata (Koch et Heynemann in Martens, 1874) T 

[= Amalia longicollis O. Boettger, 1886a; Lytopelte transcaspia Rosen, 1894; Deroceras kandaharensis Altena, 

1970] 

Amalia maculata Koch et Heynemann – Martens, 1874: 2, pl. 1, fig. 4, pl. 3, fig. 35. 

● Type locality: vicinities of Samarkand. 


● Distribution: Caucasus (Lenkoran lowland and foothills of Talysh); two findings on south slope of Great 

Caucasus (Akhsu, Padar); Central Asia (fruit-forests of Ferghana Ridge, valleyes of Zeravshan, Vakhsh and 

Pyandzh rivers); Western Kopet-Dagh (Likharev, Schileyko, MS). 

Megalopelte Lindholm, 1914 

Type species: Megalopelte simrothi Lindholm, 1914 (monotypy) 

Megalopelte simrothi Lindholm, 1914 T 

Lindholm, 1914b: 167. 

● Type locality: Dzansul (Murgut-Su river basin, Chorokh vilayet, Northeastern Turkey). 


● Distribution: Batumi Botanical Garden (Adzharia); off Akarmara village near Tkvarcheli (Abkhazia) (Likharev, 


BOETTGERILLIDAE Goethem, 1972 

Boettgerilla Simroth, 1910 

Type species: Boettgerilla compressa Simroth, 1910 (monotypy) 

Boettgerilla compressa Simroth, 1910 T 

Simroth, 1910: 531, pl. 7, figs. 29-33. 

● Type locality: Verkhnyaya Tzebelda (Abkhazia). 


● Distribution: type locality and near Kutaisi on ruins of Bagrat Castle (Likharev, Schileyko, MS). 

Boettgerilla pallens Simroth, 1912 T 

[= Boettgerilla vermiformis Wiktor, 1959] 

Simroth, 1912: 55, pl. 3, fig. 50, pl. 8, fig. 32. 

● Type locality: Gudauta (Abkhasia) 


● Distribution: in nature: Abkhazia, western Georgia near Lentekhi and Upper Svane-tia; Ciscarpathia (Baidashnikov, 

1988); parks and gardens of Moscow, Vyborg, St-Petersburg, Lvov, Ivano-Frankovsk, Chernovtzy, 

Kiev, Sochi, Vladikavkas, Erevan, Chelyabinsk (Likharev, Schileyko, MS; Sverlova, 2006); Varzobskaya 

botanical station on Hissar Ridge (Tajikistan) (Likharev, Izzatulaev, 1972).

LIMACIDAE Rafinesque, 1815 

LIMACINAE Rafinesque, 1815 

Caspilimax P. Hesse, 1926 

Type species: Limax keyserlingi Martens, 1880 (OD) 

Caspilimax keyserlingi (Martens, 1880) T 

[= Limax tigris O. Boettger, 1886; Limax talyschanus O. Boettger, 1886; Limax baeri Simroth, 1898; Gigantomilax 

robustus Simroth, 1901; Limax persicus Simroth, 1910] 

Limax keyserlingi Martens, 1880: 396. 

● Type locality: Astrabad (now Gorgan, Northern Iran). 


● Distribution: forests of Talysh and Lenkoran lowland, sporadically in Khudat and Izmail districts (Likharev, 


Caucasolimax Likharev et Wiktor, 1980 

Type species: Limax caucasicus Simroth, 1898 (OD) 

Caucasolimax caucasicus (Simroth, 1898) T 

[= Limax ananowi Simroth, 1898; Limax amaloides Simroth, 1898; Limax ananowi var. alticola Simroth, 

1912; Limax svaneticus Simroth, 1912] 

Limax caucasicus Simroth 1898: 55. 

● Type locality: Mayorsha montain, Kazbek. 


● Distribution: central part of Great Caucasus (Likharev, Schileyko, MS). 

Gigantomilax O. Boettger, 1883 

Type species: Amalia (Gigantomilax) lederi O. Boettger, 1883 (monotypy) 

Gigantomilax brunneus (Simroth, 1901) T 

Monochroma brunneus Simroth, 1901: 96, pl. 6, figs. 1-18. 

● Type locality: Nagornyi Karabakh, Karagjol Lake. 


● Distribution: Zangezur mountain system (Likharev, Schileyko, MS). 

Gigantomilax daghestanus (Simroth, 1898) T 

[= Limax florenskii Simroth, 1912] 

Limax daghestanus Simroth, 1898: 54. 

● Type locality: Gunib (Central Daghestan). 


● Distribution: Central Daghestan (Gunib, Mekhelta, Agvali), southern Armenia (Leninkan, Vedinsky and 

Daralages zoogeoraphical area) (Akramowski, 1971; Likharev, Schileyko, MS). 

Gigantomilax koenigi (Simroth, 1912) T 

[= Limax oltinus Simroth, 1912; Limax jailanus Simroth, 1912] 

Limax koenigi Simroth, 1912: 10, pl. 1, fig. 4, pl. 5, fig. 4. 

● Type locality: Ker-Ogly (vilayet Erzerum, Turkey). 


● Distribution: Adzharia (valley of Machakhelis-tzkali River), Armenia (Idzhevan, Noemberyan and Vedinsky 

districts), Talysh (Likharev, Schileyko, MS).

Gigantomilax lederi (O. Boettger, 1883) T 

Amalia lederi O. Boettger, 1883: 143, pl. 4, fig. 1. 

● Type locality: Svanetia (Georgia). 

● Lectotype (Likharev, Wiktor, 1980): SMF No. 155289. 

● Distribution: southern slopes and offshoots of Great Caucasus from Mzymta River basin to Suram pass and 

Borzhomi area, Adzharo-Imereti and Shavshetky ridges, Teberda Reserve (Likharev, Schileyko, MS); Azerbaijan 

(Wiktor, Jurkowska, 2007). 

Gigantomilax lenkoranus Simroth, 1912 T 

[= Gigantomilax talyschanus Simroth, 1912] 

Simroth, 1912: 48, pl. 3, fig. 44, pl. 7, fig. 27. 

● Type locality: Kyz-Kalasi mountain, Zuvand (Talysh, Azerbaijan). 


● Distribution: Talysh Ridge (southeastern Azerbaijan), near tops of Marayurt, Kyz-Jrdy, Kizkalasi and 

Shindan-Kalasi mountains (Likharev, Schileyko, MS). 

Gigantomilax monticola monticola (O. Boettger, 1881) T 

[= Gigantomilax brunneus Simroth, 1912; Gigantomilax borshomensis Simroth, 1912] 

Limax monticola O. Boettger, 1881a: 180-182, pl. 7, fig. 6a-c. 

● Type locality: «8000’ Höhe auf dem Taparowan in Hocharmenien» (vicinities of Paravani lake, South 

Georgia). 

● Holotype: SMF No. 155348/1 

● Distribution: between Dzhavakhet, Samsar and Trialet ridges (South Georgia) (Likharev, Schileyko, MS). 

Gigantomilax monticola armeniacus (Simroth, 1886) T 

[= Limax Dymczewiczi sensu Martens, 1880 non Kaleniczenko, 1851; Limax primitivus Simroth, 1894; Limax 

colchicus Simroth, 1898; Limax valentini Simroth, 1898; Limax ordubadensis Simroth, 1901; Limax incipiens 

Simroth, 1912; Limax kaznakovi Simroth, 1912; Limax simplex Simroth, 1898; Limax schmidti Simroth, 

1912; Limax schelkovnikovi Simroth, 1912; Limax voronovi Simroth, 1912] 

Limax armeniacus Simroth, 1886: 27, pl. 1, figs. 4-7. 

● Type locality: Sevanga peninsula, Sevan Lake (Armenia). 


● Distribution: most part of mountain systems of Lesser Caucasus (Likharev, Schileyko, MS). 

Lehmannia Heynemann, 1862 

Type species: Limax marginatus Müller, 1774 (monotypy) 

Lehmannia jaroslaviae Grossu, 1967 T 

[= Lehmannia vrancensis Lupu, 1973] 

Grossu, 1967: 120-124, figs. 1-6. 

● Type locality: "Jaroslave, vallée du Topolog, région Arges" (Romania). 

● Holotype: Grossu collection No. 1131 (Natural History Museum of Bucuresti). 

● Distribution: a single finding near Kipriani village (Moldavia) (Likharev, Schileyko, MS). 

Lehmannia macroflagellata Grossu et Lupu, 1962 T 

Grossu, Lupu, 1962: 198, figs. 9-11. 

● Type locality: Pitrozul (Romania). 

● Holotype: National History Museum Bucuresti No. 13133. 

● Distribution: Ukrainian Carpathian (Chernogora and Chivchin Mountains) (Likharev, Schileyko, MS; Baidashnikov, 

1988).

Lehmannia marginata (Müller, 1774) T 

[= Limax arborum Bouchard-Chantereaux, 1838; Limax livonicus Schrenk, 1848] 

Limax marginatus Müller, 1774: 10. 



● Distribution: Baltic countries, Karelian isthmus, Valaam Island (Ladoga Lake), Byelorussian and Ukrainian 

Polesye, Lvov, Lvov, Ivano-Frankovsk, Cherkassy, Ternopol, Chernovtzy and Zakarpatskaya regions (Likharev, 


Lehmannia valentiana (Férussac, 1821) T 

[= Limax poirieri Mabille, 1883] 

Limax valentianus Férussac, 1821: 21. 

● Type locality: Valencia (Spain). 


● Distribution: gardenfarms of St.-Petersburg, Rostov-on-Don and Tashkent (Likharev, Schileyko, MS). 

Limax Linnaeus, 1758 

Type species: Limax maximus Linnaeus, 1758 (SD Férussac, 1819) 

Limax bielzi Seibert, 1873 T 

Seibert, 1873: 195. 

● Type locality: "Mistek" [Severomoravsky Kraj, Czech Republic]. 


● Distribution: vicinities of Zhornava village, Zakarpatskaya Region, Ukraine (Sverlova, 2006). 

Limax cinereoniger Wolf, 1803 T 

[= Limax antiquorum Férussac, 1819 part.] 

Wolf, 1803: 7. 

● Type locality: Germany. 


● Distribution: Baltic countries, southern Karelia, western and central regions of European part of Russia, 

Byelorussia, northern and western Ukraine (Likharev, Schileyko, MS). 

Limax flavus Linnaeus, 1758 T 

[= Limax variegatus Draparnaud, 1801] 

Linnaeus, 1758: 652. 



● Distribution: cellars, basements and greenhouses of Moscow, Kaliningrad, Tartu, Saratov and Rostov-on- 

Don, Yalta (Likharev, Schileyko, MS); Odessa, Zmeinyi Island in the Black Sea (Sverlova, 2006). 

Limax maculatus (Kaleniczenko, 1851) T 

[= Limax variegatus sensu Moquin-Tandon, 1855 non Draparnaud, 1801; Limax ecarinatus O. Boettger, 

1881; Limax grossui Lupu, 1970] 

Krynickillus maculatus Kaleniczenko, 1851: 226, pl. 6, fig. 2. 

● Type locality: Kutchuk-Koi (now Kiparisnoe, southern coast of Crimea). 


● Distribution: mountain Crimea and Caucasus; cellars and greenhouses of Pushkin, St.-Petersburg, Sevastopol, 

Vladikavkaz, Tbilisi and Astrakhan (Likharev, Schileyko, MS); urbanized biotopes in Ukraine: in Nikolaev, 

Odessa, Zaporozhje, Berdyansk, Donetzk, Novoazovsk, Belozersk District of Kherson Region, 

northward to Vasilkov City in Kiev Region (Sverlova, 2006).

Limax maximus Linnaeus, 1758 T 

[= Limax cinereus Müller, 1774; Limax antiquorum Férussac, 1819 part.] 

Linnaeus, 1758: 652. 



● Distribution: a common species in parks and gardens in European part of Russia (Likharev, Schileyko, 

MS); Ukraine: Lvov, Chernovtzy, Zhitomir, Vinnitza, set-dements of Volyn, Ivano-Frankovsk, Chernigov, 

Kiev regions, forest biotopes of Slovechansko-Ovruch Range (Zhitomir Region) (Sverlova, 2006). 

Malacolimax Malm, 1868 

Type species: Limax tenellus Müller, 1774 (monotypy) 

Malacolimax tenellus (Müller, 1774) T 

[= Limax serotinus Schrenk, 1848] 

Limax tenellus Müller, 1774: 11. 



● Distribution: northwestern, western and central areas of European part of Russia (Likharev, Schileyko, 

MS); western, northwestern and northern Ukraine (Sverlova, 2006). 

Turcomilax Simroth, 1901 

Type species: Gigantomilax (Turcomilax) nanus Simroth, 1901 (monotypy) 

Turcomilax ferganus (Simroth, 1910) T 

Gigantomilax ferganus Simroth, 1910: 528, pl. 7, figs. 22-25. 

● Type locality: eastern slope of Chatkal Ridge. 


● Distribution: Chatkal Ridge (Likharev, Schileyko, MS). 

Turcomilax nanus (Simroth, 1901) T 

[= Gigantomilax abramowi Simroth, 1910; Gigantomilax pischpekensis Simroth, 1910] 

Gigantomilax nanus Simroth, 1901: 177, pl. 19, figs. 15-20. 

● Type locality: Arslanbob, Baubash-Ata Mount (Ferghana Ridge). 


● Distribution: Ferghana, Chatkal and Kirgiz ridges (Likharev, Schileyko, MS). 

Turcomilax natalianus (Michaelis, 1892) T 

[= Lehmania stummeri Simroth, 1910; Gigantomilax iliensis Altena, 1954] 

Limax natalianus Michaelis, 1892: 31. fig. 3. 

● Type locality: mountains of eastern Kazakhstan. 


● Distribution: Kirgiz, Kungei, Dzungar and Tarbagatai ridges (Central Asia) (Likharev, Schileyko, MS). 

Turcomilax turkestanus (Simroth, 1898) T 

[= Limax dengis Simroth, 1910] 

Limax turkestanus Simroth, 1898: 54. 

● Type locality: «Turkestan.» 


Distribution:vicinities of Przhevalsk (Terskei-Alatau); Zailijskij Alatau; Zhetyzhol Ridge (Likharev, 


Turcomilax tzvetkovi Likharev et Wiktor, 1980 T 

Likharev, Wiktor, 1980: 258, figs. 309, 310.

● Type locality: 12 km from Turgen village, Alma-Ata Region, Zailijskij Alatau. 


● Distribution: Turgen ravine (eastern part of Zailijskij Alatau) (Likharev, Schileyko, MS). 

EUMILACINAE Likharev et Wiktor, 1980 

Eumilax O. Boettger, 1881 

Type species: Limax (Milax) Brandti Martens, 1880 (OD) 

Eumilax brandti Martens, 1880 T 

[= Paralimax multirugatus O. Boettger, 1888; Paralimax marmoratus Simroth, 1901; Paralimax niger Simroth, 

1901; Paralimax salamandroides Simroth, 1901; Paralimax gyratus Simroth, 1901; Paralimax gracillis 

Simroth, 1901; Paralimax ochraceus Simroth, 1901; Paralimax albocarinatus Simroth, 1901; Eumilax 

brandti var. ciscaucasica Rosen, 1901; Eumilax kalischewskii Simroth, 1910] 

Limax brandti Martens, 1880: 143. 



● Distribution: mountain forests and meadows on south slopes of Great Caucasus, Suram, Trialet, Adzharo- 

Imereti and Shavshet chains of Lesser Caucasus (Likharev, Schileyko, MS). 

Eumilax intermittens (O. Boettger, 1883) T 

[= Paralimax raddei Simroth, 1901; Paralimax minutus Simroth, 1901; Paralimax orientalis Simroth, 1912; 

Paralimax griseus Simroth, 1912] 

Limax intermittens O. Boettger, 1883: 145, pl. 4, fig.7. 



● Distribution: mountain forests and meadows on south and north offshoots of Great Caucasus (Likharev, 


Metalimax Simroth, 1896 

Type species: Metalimax elegans Simroth, 1901 (monotypy) 

Metalimax elegans Simroth, 1901 T 


● Type locality: Tkibuli (Western Georgia). 


● Distribution: southern offshoots of Great Caucasus (Rachinsky, Zgriisky and Kodor ridges) in western 

Georgia and Abkhazia (Tkibuli, Tzageri, Tkvarcheli) (Likharev, Schileyko, MS). 

Metalimax varius (O. Boettger, 1884) T 

Paralimax varius O. Boettger, 1884: 147. 

[= Paralimax reibischi Simroth, 1892] 

● Type locality: Psyrtzkha (near Novyi Afon, Abkhazia). 


● Distribution: mountain and foothill forests of northwestern Great Caucasus; greenhouses of Sochi and 

Rostov-on-Don (Likharev, Schileyko, MS). 

LIMACOPSIDAE Gerhardt, 1936 

Bielzia Clessin, 1887

Type species: Limax coerulans M. Bielz, 1851 (monotypy) 

Bielzia coerulans (M. Bielz, 1851) T 

[= Limax schwabi Frauenfeld, 1864] 

Limax coerulans Bielz, 1851: 14. 

● Type locality: South Carpathians (Romania). 


● Distribution: Ukrainian Carpathians (Likharev, Schileyko, MS); sporadically on plains of Ukraine: area 

between Prut and Dniester rivers, western Podolsk Hills, vicinities of Vinnitza and Kiev (Sverlova, 2006). 

TRIGONOCHLAMYDIDAE Hesse, 1882 

TRIGONOCHLAMYDINAE Hesse, 1882 

Boreolestes Schileyko et Kijashko, 1999 

Type species: Boreolestes likharevi Schileyko et Kijashko, 1999 (OD) 

Boreolestes likharevi Schileyko et Kijashko, 1999 T 

Schileyko, Kijashko, 1999: 40-41, figs. 1, 2. 

● Type locality: North-Western Caucasus, Oshten-Fisht Mountains, west-facing slope of Mount Oshten. 


● Distribution: North-Western Caucasus, Oshten-Fisht Mountains (Schileyko, Kijashko, 1999). 

Boreolestes sylvestris Kijashko in Schileyko et Kijashko, 1999 T 

Schileyko, Kijashko, 1999: 41-43, figs. 3, 4. 

● Type locality: North-Western Caucasus, Molchepa riverside (right tributary of Belaya River), 6 km from 

Ghooseriple. 


● Distribution: North-Western Caucasus, upper part of Belaya River basin (Schileyko, Kijashko, 1999). 

Drilolestes Lindholm, 1925 

Type species: Pseudomilax retowskii O. Boettger, 1884 (OD) 

Drilolestes retowskii (O. Boettger, 1884) T 

[= Pseudomilax reibischi Simroth, 1901; Pseudomilax ananowi Simroth, 1901; Pseudomilax orientalis Simroth, 

1912] 

Pseudomilax retowskii O. Boettger, 1884: 146. 

● Type locality: Psyrtzkha near Novyi Afon (Abkhazia). 


● Distribution: forest and alpine zones of Great Caucasus and northern offshoots of Lesser Caucasus eastward 

of Dzhevakheti ridge (Likharev, Schileyko, MS). 

Hyrcanolestes Simroth, 1901. 

Type species: Parmacella velitaris Martens, 1880 (OD) 

Hyrcanolestes velitaris (Martens, 1880) T 

[= Pseudomilax bicolor O. Boettger, 1881; Hyrcanolestes valentini Simroth, 1901; Hyrcanolestes armeniacus 

Simroth, 1910; Hyrcanolestes kaznakovi Simroth, 1912; Hyrcanolestes varius Simroth, 1912; Hyrcanolestes 

obscurus Simroth, 1912; Hyrcanolestes fursovi Simroth, 1912; Hyrcanolestes declivis Simroth, 1912; 

Chrysalidomilax sphingiformis Simroth, 1912] 

Parmacella velitaris Martens, 1880: 154.

● Type locality: Astrabad (now Gorgan), northern Iran. 


● Distribution: forests of Lenkoran lowland and Talysh, northern and eastern offshoots of Lesser Caucasus, 

southern offshoots of eastern Great Caucasus (Likharev, Schileyko, MS). 

Khostalestes Suvorov, 2003 

Type species: Khostalestes kochetkovi Suvorov, 2003 

Khostalestes kochetkovi Suvorov, 2003 T 

Suvorov, 2003: 150-152, figs. 1-3. 

● Type locality: Northwestern Caucasus, Krasnodar Territory, Kavkazsky State Natural Biosphere Reserve, 

Khosta yew-box tree wood. 



Lesticulus Schileyko, 1988 

Type species: Lesticulus nocturnus Schileyko, 1988 (OD) 

Lesticulus nocturnus Schileyko, 1988 T 

Schileyko, 1988b: 1733, fig. 2. 

● Type locality: Oficho Cave near Kumistavi village (near Tzkhaltubo, western Georgia). 



Selenochlamys O. Boettger, 1883 

Type species: Selenochlamys pallida O. Boettger, 1883 (monotypy) 

Selenochlamys pallida O. Boettger, 1883 T 

[= Selenochlamys plumbea Simroth, 1912] 

Boettger, 1883: 142, pl. 5, fig. 1. 



● Distribution: near Kutaisi, Gelati, Tkibuli, Chikhareshi, Lentekhi (western Georgia), Tkvarcheli and 

Amtkel (Abkhazia) and south of Maikop (Likharev, Schileyko, MS). 

Trigonochlamys O. Boettger, 1881 

Type species: Trigonochlamys imitatrix O. Boettger, 1881 (monotypy) 

Trigonochlamys imitatrix O. Boettger, 1881 T 

[= Pseudomilax lederi O. Boettger, 1881; Trigonochlamys semiplumbeus O. Boettger, 1886; Trigonochlamys 

boettgeri Retowski, 1889; Trigonochlamys minor Simroth, 1901; Phrixolestes adshricus Simroth, 1901; 

Phrixolestes ponticus Simroth, 1901; Trigonochlamys pontica Simroth, 1912; Trigonochlamys armeniaca 

Simroth, 1912; Trigonochlamys distans Simroth, 1912] 

Boettger, 1881a: 177-178, pl. 7, fig. 5a-c. 



● Distribution: Georgia: forest zone of Adzharo-Imereti and Shavshetsky ridges, southern slopes of Rachinsky 

ridge (near Kutaisi and Tkibuli); Armenia: subtropical microsites in Megrin and Kafan areas (Akramowski, 

1976); Azerbaijan: outer slopes of Lesser Caucasus (Khanlar and Shamkhor areas), near Kusari 


Troglolestes Ljovushkin et Matiokin, 1965 

Type species: Troglolestes sokolovi Ljovushkin et Matiokin, 1965 (monotypy)

Troglolestes sokolovi Liovushkin et Matiokin, 1965 T 

Liovushkin, Matiokin, 1965: 35. 

● Type locality: Vorontzovskaya Cave near Sochi. 



PARMACELLIDAE Gray, 1860 

Candaharia Godwin-Austen, 1888 

Type species: Parmacella rutellum Hutton, 1849 (monotypy) 

Candaharia aethiops (Westerlund, 1896) T 

Parmacella anthiops Westerlund, 1896: 183 (nom. err. – Lindholm, 1922a: 305). 

● Type locality: along Uzun-kur-su River (Ferghana Ridge). 

● Holotype: lost. 

● Distribution: Kirgiz, Ferghana, Hissar, Darvaz, Peter the Great ridges and Khorog (Pamir) (Likharev, 


Candaharia izzatullaevi Likharev et Wiktor, 1980 T 

Likharev, Wiktor, 1980: 352, color plate, fig 2, figs. 475-479. 

● Type locality: off Pendzhikent (Tajikistan). 


● Distribution: type locality, near irrigation system of Botanical Garden of Tajik Academy of Sciences 

(Dushanbe), vicinities of Dzhizak station (Uzbekistan) (Likharev, Schileyko, MS). 

Candaharia levanderi (Simroth, 1901) T 

[= Parmacella kaznakovi Simroth, 1912; Parmacella roseni Simroth, 1912] 

Parmacella levanderi Simroth, 1901: 203, pl. 20. fig. 4, 10, 21, 22, pl. 21, figs. 1-5. 

● Type locality: Samarkand. 

● Lectotype (Likharev, Wiktor, 1980): Zoological Museum of Helsinki University. 

● Distribution: Alai mountain system from Zeravshan Ridge in the west to Alai Ridge and Peter the Great 

Ridge in the east, Chatkal Ridge (Central Asia) (Likharev, Schileyko, MS). 

Candaharia rutellum (Hutton, 1849) T 

[= Parmacella kainarensis Simroth, 1912; Vitrina baccate Hutton, 1849] 

Parmacella rutellum Hutton, 1849: 649. 

● Type locality: Kandahar (Afghanistan). 


● Distribution: mountains and offshoots of Tien-Shan, Alai and Pamir-Darvaz mountain systems (Likharev, 

Schileyko, MS); environs of St.-Petersburg (Wiktor, Jurkowska, 2007). 

Parmacella Cuvier, 1804 

Type species: Parmacella olivieri Cuvier, 1804 (monotypy) 

Parmacella ibera Eichwald, 1841 T 

[= Parmacella olivieri sensu Simroth, 1883 non Cuvier, 1804; Clathropodium vitrinaeformis Westerlund, 

1897; Parmacella simrothi Germain, 1912] 

Parmacella olivieri var. ibera Eichwald, 1841: 250-251, pl. 38, figs. 1-3. 

● Type locality: "in Iberia" (Georgia). 


● Distribution: western Kopet Dagh, eastern Transcaucasia, southern offshoots of Great Caucasus, northern

and norteastern offshoots of Lesser Caucasus, coast of Caspian Sea from Makhachkala to Yalama, Lenkoran 

lowland and Talysh mountains. Introduced to Caucasian coast of Black Sea (Likharev, Schileyko, MS). 

MILACIDAE Ellis, 1926 

Milax Gray, 1855 

Type species: Limax gagates Draparnaud, 1801 (SD Hesse, 1926) 

Milax caucasicus (Simroth, 1912) T 

Amalia caucasica Simroth, 1912: 52, pl. 3, fig. 48, pl. 8, fig. 31. 

● Type locality: Borzhomi (Georgia). 


● Distribution: type locality, near Lenkoran, and offshoots of Talysh mountains (Likharev, Schileyko, MS). 

Tandonia Lesson et Pollonera, 1882 

Type species: Limax marginatus Draparnaud, 1801 (non Müller, 1774) (= Limax rusticus Millet, 1843) (SD 

Hesse, 1926) 

Tandonia cristata (Kaleniczenko, 1851) T 

[= Milax cristatus var. nanus Grossu et Lupu, 1961] 

Krynickillus cristatus Kaleniczenko, 1851: 225, pl. 6, fig. 1. 

● Type locality: Otuz (now Shchebetovka) Southern Crimea. 


● Distribution: Sevastopol, Yalta and Simferopol (Crimea), Odessa and Uzhgorod (Likharev, Schileyko, 

MS). 

Tandonia kaleniczenkoi (Clessin, 1883) T 

[= Milax dobrogicus Grossu, 1882; Milax samsunensis sensu Forcart, 1953 non Forcart, 1942] 

Amalia kalenzkoi Clessin, 1883: 39, pl. 2, fig. 11 (nom. err.). 

● Type locality: Strateis (district of Yalta). 


● Distribution: Sevastopol, Yalta, Simferopol (Crimea) (Likharev, Schileyko, MS). 

Tandonia kusceri (H. Wagner, 1931) T 

[= Milax rusticus longipenis Grossu et Lupu, 1961; Milax rusticus f. balcanicus Grossu et Lupu, 1961; Milax 

bojanensis Hudec, 1964] 

Milax kusceri H. Wagner, 1931: 200. 

● Type locality: Svetka Petka near Nisha (Yugoslavia). 

● Holotype: NHMV. 

● Distribution: a park and Botanical Garden in Odessa, Nikolaev and several localities of Odessa and Nikolaev 

regions (Likharev, Schileyko, MS; Sverlova, 2006); NW of Simferopol in Crimea (Leonov, 2007). 

PHILOMYCIDAE Gray, 1847 

Meghimatium Hasselt, 1823 

Type species: Meghimatium striatus Hasselt, 1823 (monotypy) 

Meghimatium bilineatum (Benson, 1842) T 

Incilaria bilineatum Benson, 1842: 486 

● Type locality: Chzhou-Shan (China).


● Distribution: Primorye Territory and central part of Amur River basin (Likharev, Schileyko, MS). 

ARIONIDAE Gray, 1840 

ARIONINAE Gray, 1840 

Arion Férussac, 1819 

Type species: Arion empiricorum Férussac, 1819 [part. syn. Limax ater or Limax rufus Linnaeus, 1758] (SD 

Hesse, 1926) 

Arion ater (Linnaeus, 1758) T 

[= Arion empiricorum Férussac, 1819 pars.] 

Limax ater Linnaeus, 1758: 652. 



● Distribution: probably can be found in Karelia (Likharev, Schileyko, MS). 

Arion circumscriptus Johnston, 1828 T 

[= Arion bourguignati Mabille, 1868] 

Johnston, 1828: 76. 



● Distribution: found near St.-Petersburg and Zheleznovodsk (North Caucasus) (Likharev, Schileyko, MS); 

Lvov and Lvov region, park in Vinnitza (Sverlova, 2006). 

Arion distinctus Mabille, 1868 T 

Mabille, 1868: 137. 

● Type locality: "Sèvres, Pare de St. Cloud" [France]. 

● Neotype (de Winter, 1984): Rijksmuseum van Natuurlijke Historie, Leiden, No. alc. 9120. 

● Distribution: Ukraine: Lvov, Ivano-Frankovsk, Zhitomir, vicinities of Pustomyty town (Lvov Region), 

probably Uzhgorod and Kiev (Sverlova, 2006). 

Arion fasciatus (Nilsson, 1823) T 

[= Arion nilssoni Pollonera, 1887] 

Limax fasciatus Nilsson, 1823 (non 1822): 3-5. 

● Type locality: Lund (Sweden). 


● Distribution: found in Moscow, Tver, Lipetzk, Orel, Kursk, Pskov, Kaluga; Zakarpatskaya, Ivano- 

Frankovsk, Chernovtzy, Lvov, Ternopol, Kiev regions, near St.-Petersburg, Arkhangelsk and Kiev (Likharev, 


Arion hortensis Férussac, 1819 T 

Férussac, 1819: 65-66, pl. 2, figs. 4-5. 

● Type locality: Paris and its environs (France). 


● Distribution: a single finding near Uzhgorod (Zakarpatskaya Region) (Likharev, Schileyko, MS). 

Arion rufus Linnaeus, 1758 T 

[= Arion empiricorum Férussac, 1819 pars] 

Limax rufus Linnaeus, 1758: 652.



● Distribution: near Pushkino and Kuskovo (Moscow Region) and southern Karelia near Sandal Lake (Likharev, 


Arion sibiricus Simroth, 1901 T 


● Type locality: Irkutsk. 


● Distribution: Central and Eastern Siberia, middle and southern parts of Amur River basin, Primorye Territory, 

Sakhalin and Kunashir Island (Likharev, Schileyko, MS). 

Arion silvaticus Lohmander, 1937 T 

Arion circumscriptus var. silvaticus Lohmander, 1937: 94, figs. 10-11. 

● Type locality: Southern Sweden. 


● Distribution: off Shaulyai and Pagegjaya (Lithuania), Pushkin and Mozhaiskoje (near St.-Petersburg), Carpathians, 

western Podolsk Hills, area between Prut and Dniester rivers near Yaremcha (Ivano-Frankovsk Region,) 

(Likharev, Schileyko, MS; Sverlova, 2006). 

Arion subfuscus (Draparnaud, 1805) T 

[= Arion krynickii Kaleniczenko, 1851; Arion brunneus Lehmann, 1862; Arion esthonicus Poska-Teiss, 1927] 

Limax subfuscus Draparnaud, 1805: 125-126, pl. 9, fig. 8. 

● Type locality: «le Sorézois et la montagne Noire» (southeastern France). 


● Distribution: widely distributed in European part of Russia, common in Carpathians, western and eastern 

slopes of Central and South Ural, Ukraine, North Caucasus near Vladikavkaz. Absent in Crimea (Likharev, 


Arion vulgaris Moquin-Tandon, 1855 T 

[= Arion lusitanicus auctt., non Mabille, 1868] 

Arion rufus var. vulgaris Moquin-Tandon, 1855: 10, pl. 1, fig. 1. 



● Distribution: a single finding in Kaluga Region (near Fersikovo village, 39 km east of Kaluga) (Likharev, 

Schileyko, MS, as Arion lusitanicus). 

HELICIDAE Rafinesque, 1815 

ARIANTINAE Mörch, 1864 

Arianta Turton, 1831 

Type species: Helix arbustorum Linnaeus, 1758 (OD) 

Arianta arbustorum (Linnaeus, 1758) T 

Helix arbustorum Linnaeus, 1758: 771. 

● Type locality: «in Europae arbustis». 

● Types: probably in BMNH. 

● Distribution: Ukrainian Carpathians, western Podolsk Hills (Sverlova, 2006); Byelorussia, Baltic countries, 

introduced to vicinities of St.-Petersburg and Moscow (Likharev, Schileyko, MS); northern Moldavia (Coadã, 

2007).

Arianta petrii (Kimakowicz, 1890) T 

Helix aethiops var. petrii Kimakowicz, 1890: 204. 

● Type locality: Carpathians. 


● Distribution: Ukraine – Carpathians, Chernovtzy Region (Likharev, Schileyko, MS; Sverlova, 2006). 

Campylaea Beck, 1837 

Type species: Helix faustina Ziegler in Rossmässler, 1835 (OD) 

Campylaea faustina (Ziegler in Rossmässler, 1835) T 

[= Helix faustina var. associata Rossmässler, 1835; Helix faustina f. citrinula Rossmässler, 1835] 

Helix faustina Ziegler – Rossmässler, 1835b: 4-5, pl. 6, fig. 93. 

● Type locality: «Galicien». 


● Distribution: Carpathians and western Ukraine (Likharev, Schileyko, MS; Sverlova, 2006). 

Drobacia Brusina, 1904 

Type species: Helix banatica Rassmaessler, 1838 (OD) 

Drobacia banatica (Partsch in Rossmässler, 1838) T 

Helix banatica Partsch – Rossmässler, 1838: 14, pl. 7, fig. 457. 

● Type locality: «im Banat, bei Mehadia» (Romania). 


● Distribution: Ukrainian Carphatians (Likharev, Schileyko, MS). 

Helicigona Férussac, 1821 

Type species: Helix lapicida Linnaeus, 1758 (SD Pilsbry, 1895) 

Helicigona lapicida (Linnaeus, 1758) T 

Helix lapicida Linnaeus, 1758: 768. 

● Type locality: «in Europae rupibus terrestribus». 


● Distribution: Kalinigrad Region, maritime Latvia (Likharev, Schileyko, MS). 

Isognomostoma Fitzinger, 1833 

Type species: Helix personata Lamarck, 1792 (OD) 

Isognomostoma isognomostomum (Gmelin, 1791) T 

[= Helix personata Lamarck, 1792] 

Helix isognomostomos Gmelin, 1791: 3621. 

● Type locality: «Virginia, Thuringia, Alsatia». 


● Distribution: Carpathians, Ciscarpathia and Transcarpathia, western Podolsk Hills, area between Prut and 

Dnieper rivers, Khmelnistky Region (Likharev, Schileyko, MS; Sverlova, 2006). 

HELICINAE Rafinesque, 1815 

Caucasotachea C. Boettger, 1911 

Type species: Helix atrolabiata Krynicki, 1833 (OD)

Caucasotachea atrolabiata (Krynicki, 1833) T 

[= Helix stauropolitanus A. Schmidt, 1854; Caucasotachea argonautarum Roschkovski, 1919 nom. nov. pro 

Helix atrolabiata auct. non Krynicki, 1833; Helix atrolabiata f. tricolor O. Boettger, 1881; Helix atrolabiata 

f. maculato-fasciata O. Boettger, 1881; Helix atrolabiata f. decussata O. Boettger, 1881; repanda Mousson, 

1879; laeta Westerlund, 1898; albispira Lindholm, 1913; voronoviae Lindholm, 1913; alternans Retowski, 

1914; transiens Retowski, 1914] 

Helix atrolabiata Krynicki, 1833: 425-428, pl. 10 a-c. 

● Type locality: Mashuk and Besh-Tau mountains near Pyatigorsk. 


● Distribution: horseshoe-like area of western wing of Great Caucasus from north, west and south. Sporadically 

in western Colchis lowland and off Kutaisi (Likharev, Schileyko, MS). 

Caucasotachea calligera (Dubois de Montpéreux, 1840) T 

[= Helix calligera var. pallasii Mousson, 1863; Helix atrolabiata var. intercedens Retowski, 1889; Helix 

atrolabiata var. nemoraloides Kobelt, 1903] 

Helix calligera Dubois de Montpéreux, 1840: 87. 

● Type locality: Caucasus. 


● Distribution: southern slopes of Great Caucasus, Colchis lowland, western Lesser Caucasus (Likharev, 


Caucasothachea lencoranea (Mousson, 1863) T 

Helix atrolabiata var. lencoranea Mousson, 1863: 203. 

● Type locality: Lenkoran. 


● Distribution: Lenkoran lowland and forests of Talysh (Caucasus) (Likharev, Schileyko, MS). 

Cepaea Held, 1837 

Type species: Helix nemoralis Linnaeus, 1758 (SD Herrmannsen, 1846) 

Cepaea hortensis (Müller, 1774) T 

Helix hortensis Müller, 1774: 52-55. 



● Distribution: Kaliningrad Region, Latvia, Estonia and sporadically in Vitebsk Region (Likharev, Schileyko, 

MS); introduced in western Ukraine (Sverlova, 2006). 

Cepaea nemoralis (Linnaeus, 1758) T 

Helix nemoralis Linnaeus, 1758: 773. 

● Type locality: «in Europae arboribus». 

Prabable syntypes: UUZM No. 757. 

● Distribution: locally along coast of Baltic Sea (Kaliningrad Region, Latvia, Estonia) (Likharev, Schileyko, 

MS); Lvov (Sverlova, 2006). 

Cepaea vindobonensis (C. Pfeiffer, 1828) T 

[= Helix austriaca Menke, 1828; Helix ervensis Krynicki, 1837] 

Helix vindobonensis Pfeiffer, 1828: 15-16, pl. 4, figs. 6-7. 

● Type locality: "im Erzherzogthum Oesterreich und Illyrien". 


● Distribution: forest-steppe and steppe zones of East-European Plain, steppes of Crimea and foothills of 

North Caucasus (Likharev, Schileyko, MS). 

Cryptomphalus Agassiz, 1837

Type species: Helix aspersa Müller, 1774 (SD Pilsbry, 1889) 

Cryptomphalus aspersa (Müller, 1774) T 

Helix aspersa Müller, 1774: 59. 

● Type locality: «Italia». 


● Distribution: a single finding in Crimea near Feodosia (Likharev, Schileyko, MS). 

Eobania Hesse, 1915 

Type species: Helix vermiculata Müller, 1774 (OD) 

Eobania vermiculata (Müller, 1774) T 

Helix vermiculata Müller, 1774 

● Type locality: Italy. 


● Distribution: Crimea (Likharev, Schileyko, MS); introduced to Odessa Region (Sverlova, 2006). 

Helix Linnaeus, 17758 

Type species: Helix pomatia Linnaeus, 1758 (SD Montfort, 1810) 

Helix albescens Rossmässler, 1839 T 

[= Helix obtusata Rossmässler, 1837 nom. praeocc., non Serres, 1824; Helix vulgaris Rossmässler, 1839 

nom. praeocc., non Da Costa, 1778; Helix obtusalis Bourguignat, 1860 nom. nov. pro H. obtusata Rossmässler, 

1837; Helix obtusalis var. bicincta Kobelt, 1877; Helix obtusata var. balionis Retowski, 1889; Helix 

intermissa Westerlund, 1897; Helix vulgaris kubanensis Kobelt, 1906; Helix vulgaris roseni Kobelt, 1906] 

Rossmässler, 1839: 10, pl. 44, figs. 585-586. 

● Type locality: Northern Italy. 


● Distribution: Odessa to DOnetsk regions, mountain and steppe Crimea (Ukraine) (Sverlova, 2006); western 

and northern parts of North Caucasus; sporadically in Daghestan, Georgia, Armenia (Likharev, Schileyko, 

MS). 

Helix buchi Dubois de Montpéreux in Pfeiffer, 1853 T 

[= Helix buchi f. minor O. Boettger, 1881; Helix buchi sieversi Kobelt, 1903; Helix pomatia duschekensis 

Kobelt, 1906; Helix buchi var. adsharica Kobelt, 1906; Helix gogerdziana Mumbadze, Tarkhnishviliet Pokryscko, 

2008] 

Helix buchi Dubois de Montpéreux – Pfeiffer, 1853a: 181. 

● Type locality: «Grusia». 


● Distribution: Georgia near Abkhazia, Verin Agdan village (Idzhevan district, Armenia) (Akramowski, 

1976; Likharev, Schileyko, MS). 

Helix christophi O. Boettger, 1881 T 

[= Helix christophi var.? atrolabioides Kobelt, 1914] 

Boettger, 1881a: 217-218. 

● Type locality: Adzharia. 


● Distribution: southwestern Transcaucasia (Likharev, Schileyko, MS). 

Helix lucorum Linnaeus, 1758 T 

[= Helix taurica Krynicki, 1833; Helix radiosa Rossmässler, 1838; Helix taurica mut. martensi O. Boettger, 

1883; Helix ancyrensis haussknechti Kobelt, 1906] 

Linnaeus, 1758: 773.

● Type locality: «in Europae arboribus». 

● Neotype (ICZN Opinion 1996, 2002): Zoological Museum of the University of Copenhagen. 

● Distribution: mountain Crimea, Black Sea coast of Caucasus (north of Sochi), Colchis lowland and bordering 

mountain ridges, Armenia, Shemakha and Talysh (Likharev, Schileyko, MS). 

Helix lutescens Ziegler in Rossmässler, 1837 T 

Helix lutescens Ziegler – Rossmässler, 1837: 4, pl. 21, fig. 292. 

● Type locality: «Lemberg in Galicien» (Lvov, Ukraine). 

Neotype (Zilch, 1952b): SMF No. 9850. 

● Distribution: Carpathians in western Ukraine and Moldavia, Volyno-Podolsk Hills, Zhitomir, Odessa and 

Nikolaev regions, western Ukranian Polesye and adjacent Belorussia (Likharev, Schileyko, MS; Sverlova, 

2006). 

Helix nordmanni Parreyss in Mousson, 1854 T 

Mousson, 1854: 366-367. 

● Type locality: «le Somketh, l’Imereth et l’Armenie». 


● Distribution: southern Georgia (Likharev, Schileyko, MS). 

Helix pomatia Linnaeus, 1758 T 

Linnaeus, 1758: 771. 

[synonyms see in Zilch, 1952] 

● Type locality: «in Angliae, Galliae nemoribus». 

● Probable syntypes: UUZM No. 866. 

● Distribution: Baltic countries, Ukraine, western Byelorussia, introduced to vicinities of Moscow, Kursk, 

Kiev, Poltava and Prioksko-Terrasny Reserve (Likharev, Schileyko, MS; Sverlova, 2006). 

Levantina Kobelt, 1871 

Type species: Helix spiriplana Olivier, 1821 (SD Pilsbry, 1894) 

Levantina djulfensis (Dubois de Montpéreux, 1840) T 

[= Helix urmiensis Kobelt, 1889; Helix placida Westerlund, 1897; Helix casta Westerlund, 1898] 

Helix djulfensis Dubois de Montpéreux, 1840: 24. 

● Type locality: Djulfa. 


● Distribution: Araks River valley (Megrin district) vicinities of Bzhni (Razdan district) (Armenia); Djulfa, 

Ordubad, Zangilan (Likharev, Schileyko, MS). 

Levantina escheriana (Bourguignat, 1864) T 

Helix escheriana Bourguignat, 1864b: 105, pl. 12, figs. 8-11. 

● Type locality: Mesopotamia (now Iraque). 


● Distribution Armenia (Norashen district, Kyarki village; vicinities of Areni and Ekhegnadzor); Azerbaijan 

(Nakhichevan region) (Likharev, Schileyko, MS; Schütt, Subai, 1996). 

Levantina longinqua (Schütt et Subai, 1996) T 

Assyriella longinqua Schütt, Subai, 1996: 128-129, pi. 7, fig. 29 (nom. nov. pro Helix (Levantina) kurdistana 

var. dohrni Kobelt, 1902, non Helix dohrni Paulucci, 1882). 

● Type locality: "Samarkand" [Uzbekistan]. 


● Distribution: south Uzbekistan (Schiitt, Subai, 1996).

BRADYBAENIDAE Pilsbry, 1939 

Acusta Martens, 1860 

Type species: Helix ravida Benson, 1842 (OD) 

Acusta ravida (Benson, 1842) T 

[= Helix selskii Schrenck, 1867; Helix cincto-inflata Mousson, 1887; Helix weyrichii sensu Cockerell, 1924 

non Schrenck, 1867] 

Helix ravida Benson, 1842: 486. 

● Type locality: «Chusan». 


● Distribution: lower and middle parts of Amur River basin, the entire Primorye Territory (Egorov, Ivanov, 

1997). 

Fruticicola Held, 1837 

Type species: Helix fruticum Müller, 1774 (SD Herrmannsen, 1847) 

Fruticicola alaica (Kuznetsov, 1998) T 

Bradybaena alaica Kuznetsov, 1998: 87-88, fig. 1. 

● Type locality: Kirgizia, Osh Region, Naukat District, Kichik-Alai Ridge, Mazar-Sai River (tributary of 

Kirgiz-Alai) valley upstream from Eski-Naukat settlement, Naukat Experimental Forestry, Experimental 

Point Karagoi, eastern slope, 2500-2700 m. 


● Distribution: Osh Region of Kirgizia (Kuznetsov, 1998). 

Fruticicola almaatini (Skvortzov, 1940) T 

[= Fruticola alma-atina var. lindholmi Skvortzov, 1940] 

Fruticola almaatini Skvortzov, 1940: 372-373. 

● Type locality: Maloe Almaatinskoe ravine, Zailijskij Alatau. 

● Syntypes: ZMMU Nos. Lc-14476, Lc-14409. 

● Distribution: Kazakhstan, northern Tien-Shan, Zailijskij Alatau Ridge, Maloe Almaatinskoe ravine, Kirgiz 

Ala Too and Ferghana ridges, Susamyrskaya valley (Egorov, Ivanov, 1997). 

● Remark: In the original description of the species, Skvortzov (1940) did not designate any type specimens 

but mentioned 2 specimens and gave their dimensions. Therefore, the shell illustrated in the paper by Egorov 

and Ivanov (1997) (No. Lc-14476), was erroneously designated as holotype. 

Fruticicola bilaticincta (Martens, 1882) T 

[= Fruticola saturata Londholm, 1926] 

Helix paricincta var. bilaticincta Martens, 1882a: 5-6, pl. 1, figs. 12-13. 

● Type locality: «valem fluvii Ili ejusque affluentium ... ad fl. Kungess circa 1300 met. altit.». 


● Distribution: Ferghana and Chatkal ridges (western Tien-Shan) (Egorov, Ivanov, 1997). 

Fruticicola cavimargo cavimargo (Martens, 1879) T 

[= Helix ? aspasta Westerlund, 1896; Cathaica iacosta Möllendorff, 1899; Cathaica sügoschuriana Weber, 

1913] 

Helix cavimargo Martens, 1879: 126. 


● Lectotype (Zilch, 1968): SMF 9048. 

● Distribution: central and eastern Tien-Shan, northern Pamiro-Alai (Egorov, Ivanov, 1997). 

● Remark: Zilch (1968) cited «Gipfel des Kandagar-Gebirges, Ordos» as the locality of the lectotype.

Fruticicola cavimargo tarbagataica (Matiokin, 1966) T 

Bradybaena cavimargo tarbagataica Matiokin, 1966: 120. 

● Type locality: collective farm May-Bulak, near Karakol River, Tarbagatai Ridge. 

● Holotype: ZMMU No Lc-14426. 

● Distribution: Semipalatinsk Region, Tarbagatai Ridge, Manrak and Saur ridges, southern Altai, northern 

slope of Kurchum and southern foothills of Naryn ridges (Egorov, Ivanov, 1997). 

Fruticicola dichrozona (Martens, 1885) T 

[= Bradybaena pavlovskii sensu Matiokin, 1966 part.] 

Helix dichrozona Martens, 1885: 17. 

● Type locality: Namangan. 


● Distribution: Chatkal Ridge and southern foothills (Likharev, Schileyko, MS). 

Fruticicola fedtschenkoi (Martens, 1874) T 

[= Helix sturanyana Kobelt, 1893] 

Helix fedtschenkoi Martens, 1874: 16, pl. 1, fig. 9. 

● Type locality: off Kulikalan Lake in Zeravshan Valley. 

● Lectotype (Egorov, Ivanov, 1997): ZMMU No. Lc-645. 

● Distribution: Karatau, Zeravshan, Alai and, possibly, Chatkal ridges, on Digmay plateau, at foothills of 

Kuram and Turkestan ridges (Egorov, Ivanov, 1997). 

Fruticicola fruticum (Müller, 1774) T 

[= Helix fruticum var. europaea W. Dybowski, 1901] 

Heilx fruticum Müller, 1774: 71. 



● Distribution: East-European Plain, Crimea, Ukaranian Carpathians, North Caucasus (Likharev, Schileyko, 

MS; Sverlova, 2006). 

Fruticicola helvola helvola (Frivaldszky in L. Pfeiffer, 1853) T 

[= Bradybaena likharevi Uvalieva, 1967] 

Helix helvola Frivaldszky – Pfeiffer, 1853b: 188-189. 

● Type locality: «Sibiria». 


● Distribution: southwestern and southern Altai (Eastern Kazakhstan Region) (Egorov, Ivanov, 1997). 

Fruticicola helvola anachoretica (Matiokin, 1966) T 

Bradybaena anachoretica Matiokin, 1966: 127, pl. 4, fig. 5. 

● Type locality: Tarbagatai. 

● Types: unknown (S&L) ● Holotype: ZIN (E&!, 1997). 

● Distribution: Semipalatinsk Region, Tarbagatai Ridge, Eastern Kazakhstan, southwestern Altai, Ulbinsky 

Ridge (Likharev, Schileyko, MS; Egorov, Ivanov, 1997). 

Fruticicola lantzi (Lindholm, 1927) T 

[= Fruticola lantzi dextrorsa Tzvetkov, 1938; Fruticola lantzi sinistrorsa Tzvetkov, 1938; Fruticola lantzi f. 

cingulicarens Lindholm, 1927; Fruticola lantzi var. brauni Skvortzov, 1940; Fruticola lantzi f. albina Tzvetkov, 

1941; Fruticola lantzi f. steppensis Tzvetkov, 1941; Fruticola lantzi f. xerophila Tzvetkov, 1941; Fruticola 

lantzi f. silvestris Tzvetkov, 1941; Fruticola lantzi f. montana Tzvetkov, 1941] 

Fruticola lantzi Lindholm, 1927d: 261. 

● Type locality: Varukh village near Kokand. 

● Syntypes: ZMMU No. Lc-14486.

● Distribution: most ridges of northern and western Tien-Shan, Pamiro-Alai and Alai ridges (Likharev, 


Fruticicola necopinata (Tzvetkov, 1941) T 

Pseudiberus necopinatus Tzvetkov, 1941: 285. 

● Type locality: Tisken-Su side ravine of Turgen ravine (Zailijskij Alatau Ridge). 


● Distribution: Zailijskij Alatau Ridge, ravines of Turgen and Tisken-Su rivers, Kirgiz Ala Too ridge, Alamedin 

valley (northern Tien-Shan) (Egorov, Ivanov, 1997). 

Fruticicola perlucens (Rosen, 1901) T 

[= Helix indigena Westerlund, 1898 nom. praeocc., non Mabille, 1895] 

Helix perlucens Rosen, 1901a: 9 

● Type locality: Garm on Surkhob River. 


● Distribution: foothills of Chatkal ridge (western Tien-Shan), Alai, Peter the Great, Hissar and Zeravshan 

ridges (Pamiro-Alai) (Egorov, Ivanov, 1997). 

Fruticicola phaeozona phaeozona (Martens, 1874) T 

[= Helix orithyia Martens, 1874] 

Helix phaeozona Martens, 1874: 13, pl. 1, fig. 8, pl. 3, fig. 39. 

● Type locality: «Kokand Khanate». 


● Distribution: Ferghana and Zailijskij Alatau ridges (Tien-Shan), Alai Ridge (Egorov, Ivanov, 1997). 

Fruticicola phaeozona pseudarianta (Lindholm, 1927) T 

Fruticola phaeozona pseudarianta Lindholm, 1927c: 195. 

● Type locality: «Ferghana-Gebiet, Tal. des Fl. Tshithkan (linker Nebenfluss des Uzun-Achmat), Tal. des Fl. 

Turduk (linker Tributar des Karasu»). 

● Lectotype (Egorov, Ivanov, 1997): ZIN. 

● Distribution: Ferghana Ridge near Arslanbob village, ravines of Chichkhan and Turduk rivers (Egorov, 

Ivanov, 1997). 

Fruticicola plectotropis plectotropis (Martens, 1864) T 

[= Helix palmeni Westerlund, 1898; Helix palmeni f. accincta Westerlund, 1898; Cathaica sügoschuriana 

Weber, 1913] 

Helix plectotropis Martens, 1864a: 114, pl. 3, figs. 3-5. 

● Type locality: Tien-Shan («O-Turkestan: Karagalik» - Egorov, Ivanov, 1997). 


● Distribution: Kirgiz Ala Too, Zailijskij Alatau, Ketmen, Kungei Ala Too ridges (northern Tien-Shan) 

(Egorov, Ivanov, 1997). 

Fruticicola plectotropis mesophila (Tzvetkov, 1941) T 

Cathaica plectotropis m. mesophila Tzvetkov, 1941: 293. 

● Type locality: Issyk ravine. 


● Distribution: northern Tien-Shan, Zailijskij Alatau ridge, Issyk ravine, Turgen ravine (Likharev, Schileyko, 

MS). 

Fruticicola plectotropis saxatilis (Tzvetkov, 1941) T 

Cathaica saxatilis Tzvetkov, 1941: 298-299. 

● Type locality: Issyk ravine, 50 km eastward from Alma-Ata, 1200 to 2000 m above sea level. 

● Lectotype (Egorov, Ivanov, 1997): ZMMU No. Lc-14433.

● Distribution: Zailijskij Alatau Ridge, Issyk ravine, Kirgiz Ala Too Ridge, Alamedin valley and Chon-Tash 

ravine (Egorov, Ivanov, 1997). 

Fruticicola plectotropis scalaris (Tzvetkov, 1941) T 

[= Cathaica scalaris m. alticola Tzvetkov, 1941] 

Cathaica scalaris Tzvetkov, 1941: 297-298. 

● Type locality: Talgar ravine, 1200-1300 m above sea level. 


● Distribution: Zailijsky Alatau Ridge, Talgar ravine (Egorov, Ivanov, 1997). 

Fruticicola plectotropis scopulosus (Tzvetkov, 1941) T 

Cathaica plectotropis m. scopilosus Tzvetkov, 1941: 293-294. 

● Type locality: Maloe Almaatinskoe ravine. 

● Lectotype (Egorov, Ivanov, 1997): ZIN. 

● Distribution: Zailijskij Alatau ridge, Maloe Almaatinskoe ravine (northern Tien-Shan), Kirgiz Ala Too 

Ridge, Alamedin valley, Kungei Ala Too Ridge, northern shore of Issyk-Kul Lake, near Bosteri town 

(Egorov, Ivanov, 1997). 

Fruticicola plectotropis stschukini (Lindholm, 1927) T 

Pseudiberus stschukini Lindholm, 1927c: 200. 

● Type locality: upper reaches of Turduk River, Chatkal Ridge. 


● Distribution: Chatkal Ridge, valley of Turduk River (western Tien-Shan), Kirgiz Ala Too Ridge, Karabalty 

valley, Kara-Haty ravine, northern slope of Talas-Alatau Ridge, Besh-Tash valley (Egorov, Ivanov, 1997); 

Zailijskij Alatau Ridge, Talgar valley (Likharev, Rammelmeyer, 1952). 

Fruticicola plectotropis transiliensis (Tzvetkov, 1940) T 

Cathaica transiliensis Tzvetkov, 1940a: 391. 

● Type locality: Alayak ravine near Uzun-Aghach village, Zailijskij Alatau Ridge. 



Fruticicola schrenckii (Middendorff, 1851) T 

[= Helix sibirica L. Pfeiffer, 1853; Helix annexa Westerlund, 1897] 

Helix schrenckii Middendorff, 1851: 302, pl. 30, figs. 20-26. 

● Type locality: Siberia. 


● Distribution: northeastern Europe, Siberia, Altai and Sayan mountains, Kamchatka (Shileyko, Likharev, 

MS; Egorov, Ivanov, 1997) 

Fruticicola scythica (Westerlund, 1898) T 

[= Fruticicola roxane Lindholm, 1927] 

Helix scythica Westerlund, 1898: 168. 

● Type locality: Ferghana Ridge («Pass zwischen Dgehularik und Habin»). 

● Lectotype (Schileyko, 1978b): ZIN. 

● Distribution: Ferghana, Naryn-Too and Baubash-Ata ridges (Tien-Shan), Peter the Great Ridge on Pamiro- 

Alai (Egorov, Ivanov, 1997). 

Fruticicola skwortzowi Tzvetkov, 1940 T 

Tzvetkov, 1940a: 389 

● Type locality: Kara-Kastek river canyon, May-Tyube Pasture. 

● Syntypes: ZMMU No. Lc-14428.

● Distribution: Zailijskij Alatau Ridge (Likharev, Schileyko, MS). 

Fruticicola squamulosa (Izzatullaev et Schileyko, 1980) T 

Bradybaena squamulosa Izzatullaev, Schileyko, 1980: 1220-222, fig. 1A, B. 

● Type locality: Kyrgyzstan, northern shore of Issyk-Kul Lake, Cholpon-Ata, Progress settlement. 


● Distribution: type locality; Kazakhstan, estuary of the Koktal River (Egorov, Ivanov, 1997). 

Fruticicola stoliczkana stoliczkana (Nevill, 1878) T 

[= Helix opposita Westerlund, 1898] 

Helix stoliczkana Nevill, 1878: 3, pl. 1, figs. 4-6. 

● Type locality: Sasak Taka and Pasrobat, west of Yarkland. 

● Probable syntypes: SMF No. 22834. 

● Distribution: northern and central Tien-Shan (Egorov, Ivanov, 1997). 

Fruticicola stoliczkana chuika (Egorov in Egorov et Ivanov, 1997) T 

Bradybaena stoliczkana chuika Egorov – Egorov, Ivanov, 1997: 39: fig. 32. 

● Type locality: Kyrgyzstan, valley of Chu river, 50 km from Rybachje village. 


● Distribution: northern Tien-Shan, Boam ravine (part of Chu river valley between Kungei Ala Too and Kirgiz 

Ala Too ridges) (Likharev, Schileyko, MS). 

Fruticicola stoliczkana pavlovskii (Likharev, 1955) T 

[= Eulota pavlovskii var. depressa Likharev, 1955] 

Eulota pavlovskii Likharev, 1955: 183-185, fig. 2. 

● Type locality: Kum Tar River, northern Tien-Shan. 


● Distribution: Kyrgyzstan, northern Tien-Shan, Kum Tar River, near Rybachje village on right side of the 

Chu valley (Egorov, Ivanov, 1997). 

Fruticicola tomyris Lindholm, 1927 T 

Lindholm, 1927c: 196. 

● Type locality: Chichkan River valley, Ferghana Ridge. 



Fruticicola transbaicalia transbaicalia (Schileyko, 1978) T 

Bradybaena transbaicalia transbaicalia Schileyko, 1978b: 128-130, textfigs. 56, 57, pl. 1, fig. 10. 

● Type locality: Transbaikalia, Khamar-Daban Ridge, right bank of Mamai River. 


● Distribution: Khamar-Daban Ridge, Krasnoyarsk (Schileyko, 1978b; Egorov, Ivanov, 1997). 

Fruticicola transbaicalia sayanica (Kuznetsov in Egorov et Ivanov, 1997) T 

Bradybaena transbaicalia sayanica Kuznetsov – Egorov, Ivanov, 1997: 9, fig. 4. 

● Type locality: Russia, East Siberia, Khakassia, Biisk district, West Sayan mountain system, Alan Ridge, 

left side of the Yenisei valley, 3-4 km north-north-eastward of Cheriomushki village, foot of eastern slope of 

«1699 m» mount 20-30 m over railway, along foot of steep rocks. 


● Distribution: Sayan mountains (Egorov, Ivanov, 1997). 

Fruticicola tzwetkovi (Uvalieva et Soboleva, 1973) T 

[= Cathaica brodskii Tzvetkov et Tzvetkova, 1950 nom. nud.] 

Bradybaena tzwetkovi Uvalieva, Soboleva, 1973: 1717, figs. 1, 2.

● Type locality: Alma-Atinsky State Reserve, right bank of Levyi Talgar River. 

● Holotype: Institute of Zoology of Kazakhstan Academy of Sciences No. 5184. 

● Distribution: Zailijskij Alatau Ridge and right side of Talgar ravine (Likharev, Schileyko, MS). 

Fruticicola zhecseni Rymzhanov, 1979 T 

Bradybaena plectotropis smsp. zheceni Rymzhanov, 1979: 55, figs. 4-5. 

● Type locality: Urochishche Assy, Zailijskij Ridge. 


● Distribution: Zailijskij Alatau and Kungei ridges, Turgen ravine (northern Tien-Shan) (Likharev, 

Schileyko, MS; Egorov, Ivanov, 1997). 

Karaftohelix Pilsbry, 1927 

Type species: Eulota fiscina Fulton, 1905 (OD) 

Karaftohelix arcasiana (Crosse et Debeaux, 1863) T 

[= Helix arcasiana f. normalis Schrenck, 1867; Helix arcasiana f. depressior Schrenck, 1867; Eulota 

murenensis Cockerell, 1926; Ganesella virgo Pilsbry, 1926; Helix fruticum Gerstfeldt, 1859 nom. praeocc., 

non Müller, 1774] 

Helix arcasiana Crosse, Debeaux, 1863: 386. 

● Type locality: «Shang-hai». 


● Distribution: basin of Amur River (near Khabarovsk, Blagoveshchesk-on-Amur), Razdolno-Khankayskaya 

lowland, Pogranichny Ridge, vicinity of Nakhodka, eastern coast of Ussuri Bay, Furugelm Island (Egorov, 

Ivanov, 1997). 

Karaftohelix bocageana bocageana (Crosse, 1864) T 

[= Helix miranda A. Adams, 1868; Eulota weyrichii sensu Likharev, Rammelmeyer, 1952 part.] 

Helix bocageana Crosse, 1864: 284. 

● Type locality: ? China. 


● Distribution: southeastern Sakhalin, Shikotan Island (Egorov, Ivanov, 1997). 

Karaftohelix bocageana chishimana (Pilsbry et Hirase, 1904) T 

[= Eulota weyrichii sensu Likharev, Rammelmeyer, 1952 part.] 

Eulota chishimana Pilsbry, Hirase, 1904a: 116-117. 

● Type locality: Kunashir Island. 


Distribution:Kunashir Island (Likharev, Schileyko, MS). 

Karaftohelix bocageana urupensis (Pilsbry, 1927) T 

Eulota urupensis Pilsbry, 1927a: 17, pl. 1, figs. 1-3. 

● Type locality: Urup Island (Kurile Islands). 

● Holotype: ANSP No. 99950. 


Karaftohelix bocageana weyrichii (Schrenck, 1867) T 

[= Helix sirotina A. Adams, 1868; Eulota fiscina Fulton, 1905] 

Helix weyrichii Schrenck, 1867: 669, pl. 26, figs. 11-13. 

● Type locality: Sakhalin Island («Bai d’Estaing an der Westküste und bei Manuë an der Ostküste»). 


● Distribution: southern Sakhalin Island (Egorov, Ivanov, 1997).

Karaftohelix capillata (Schileyko et Bratchik in Schileyko, 1978) T 

Bradybaena capillata Schileyko et Bratchik – Schileyko, 1978b: 347-349, textfigs. 468-471. 

● Type locality: Malyi Island (Peter the Great Bay). 


● Distribution: Malyi, Naumov, Pakhtusov, Kozlov, Matveev, De-Livron, Durnovo, Pelis, Stenin, Klykov 

and Rimskij-Korsakov Islands (Egorov, Ivanov, 1997). 

Karaftohelix dieckmanni (Mousson, 1887) T 

[= Helix eutheta Westerlund, 1897] 

Helix dieckmanni Mousson, 1887: 13, pl. 1, fig. 2. 

● Type locality: Amur River basin. 


● Distribution: from basin of Lower and Middle Amur River to Peter the Great Bay, southern Primorye and 

adjacent islands (Likharev, Schileyko, MS). 

Karaftohelix diversita (Schileyko et Bratchik in Schileyko, 1978) T 

Bradybaena diversita Schileyko et Bratchik – Schileyko, 1978b: 140-141, textfigs. 82-84. 

● Type locality: Dva Brata Islands (Peter the Great Bay). 


● Distribution: Dva Brata, Matveev, De-Livron, Durnovo and Rimskij-Korsakov Islands (Egorov, Ivanov, 

1997). 

Karaftohelix duiensis (Westerlund, 1897) T 

Helix duiensis Westerlund, 1897b: 121. 

● Type locality: northern Sakhalin. 


● Distribution: east coast of Sakhalin Island along Due River; Kholmsk district, Racuma village; southeastern 

Sakhalin Island, Nevelsk district (Egorov, Ivanov, 1997); Kunashir Island (Pearce et al., 2002). 

Karaftohelix fragilis (Pilsbry, 1926) T 

Eulota fragilis Pilsbry, 1926: 455, pl. 33, fig. 2. 

● Type locality: «Kom-san near Senchon» (northern part of Korean Peninsula). 

● Holotype: ANSP No. 99957. 

● Distribution: southern Primorye Territory (Egorov, Ivanov, 1997). 

Karaftohelix incognita (Schileyko, 1988) T 

Bradybaena incognita Schileyko, 1988a: 1305, fig. 1, II. 

● Type locality: Sakhalin Island. 



Karaftohelix intermedia (Rymzhanov, 1983) T 

Bradybaena intermedia Rymzhanov, 1983: 30-32, figs. 1, 2. 

● Type locality: Karkara ravine, Kungei Ala Too ridge, North Tien-Shan. 


● Distribution: Kungei Ala Too and Ketmen ridges, Khan-Tengry massif, Dzungar Alatau, Saur and Sarym- 

Sakty ridges (Egorov, Ivanov, 1997). 

? Karaftohelix kudiensis (Cockerell, 1924) T 

Hygromia kudiensis Cockerell, 1924: 581. 

● Type locality: basin of Kudiya river, Primorye Territory. 


● Distribution: type locality and near Rudnaya Bay (Egorov, Ivanov, 1997).

Karaftohelix kurodana (Pilsbry, 1926) T 

Eulota kurodana Pilsbry, 1926: 455, pl. 33, figs. 3, 4. 

● Type locality: Pukhan-san north of Seoul, Korea. 

● Holotype: ANSP No. 99960a. 

● Distribution: Primorye Territory, Pogranichnyi district, Pogranichnyi Ridge (Egorov, Ivanov, 1997). 

Karaftohelix maacki (Gerstfeldt, 1859) T 

[= Helix maacki f. elatior Schrenck, 1867; Helix maacki var. depressior Schrenck, 1867; Eulota maacki f. 

optima Cockerell, 1924; Helix middendorffi Mousson, 1887 nom. praeocc., non Gerstfeldt, 1859] 

Helix maacki Gerstfeldt, 1859: 518, fig. 27. 

● Type locality: «Am mittleren Amur, vorzüglich zwischen den Mündungen des Songari und Ussuri». 


● Distribution: basin of lower and middle Amur River in Primorye Territory (Likharev, Schileyko, MS). 

Karaftohelix middendorffi (Gerstfeldt, 1859) T 

[= Helix graeseri Mousson, 1887; Eulota eulemnisca Cockerell, 1924; Hygromia amatoris Cockerell, 1924] 

Helix middendorffi Gerstfeldt, 1859: 521-522, fig. 29. 

● Type locality: Amur River between mouths of Sungari and Ussuri rivers. 


● Distribution: Primorye Territory and middle Amur River basin (Egorov, Ivanov, 1997). 

Karaftohelix plana (Schileyko et Bakurov in Schileyko, 1988) T 

Bradybaena plana Schileyko et Bakurov – Schileyko, 1988: 1305, fig. 2. 

● Type locality: Primorye Territory, Anuchinsk district, Chernyshovka station. 


● Distribution: type locality, Rimskij-Korsakov and Durnovo islands in Peter the Great Bay (Egorov, Ivanov, 

1997). 

Karaftohelix strelkovi (Likharev et Rammelmeyer, 1952) T 

Eulota strelkovi Likharev, Rammelmeyer, 1952: 408, fig. 340. 

● Type locality: southern Sakhalin. 


● Distribution: southern Sakhalin, Iturup, Shikotan, Kunashir Islands (Egorov, Ivanov, 1997; Pearce et al., 

2002). 

Karaftohelix ussuriensis (Westerlund, 1897) T 

Helix ussuriensis Westerlund, 1897b: 122. 

● Type locality: Nakhodka. 

● Lectotype (Schileyko, 1978b): NMG No. 738. 

● Distribution: coastal part of southern Primorye and islands of Peter the Great Bay (Likharev, Schileyko, 

MS). 

Karaftohelix vulcanica (Schileyko, 1978) T 

Bradybaena vulcanica Schileiko, 1978b: 142-144, textfigs. 87, 88, pl. 2, fig. 20. 

● Type locality: foot of Golovnin volcano (southern Kunashir Island). 


● Distribution: Kunashir Island (Likharev, Schileyko, MS). 

Ponsadenia Schileyko, 1978 

Type species: Helix semenovi Martens, 1864 (OD)

Ponsadenia dentata (Rymzhanov, 1983) T 

Bradybaena dentata Rymzhanov, 1983: 32, figs. 3, 4. 

● Type locality: Kungei Ridge, near Kul-say Lake. 


● Distribution: type locality and Issyk ravine (Zailijskij Alatau Ridge) (Likharev, Schileyko, MS). 

Ponsadenia duplocincta (Martens, 1879) T 

[= Helix paricincta Martens, 1879; Helix paricincta var. bisbicincta Martens, 1882; Helix stenroosi Westerlund, 

1898; Helix stenroosi f. tetrica Westerlund, 1898] 

Helix duplocincta Martens, 1879: 125. 



● Distribution: Central Asia from Ili River to eastern Kirgiz Ala Too Ridge, Ketmen Ridge, except for northern 

slopes of Zailijskij Alatau and Kirgiz Ala Too Ridges, northern shore of Issyk-Kul Lake (Likharev, 


Ponsadenia hirsuta (Matiokin, 1966) T 

Bradybaena hirsuta Matiokin, 1966: 124, pl., 4, fig. 7. 

● Type locality: Dzungar Ridge and Tarbagatai. 

● Syntypes: ZMMU Nos. Lc-9244, 9253-9257. 

● Distribution: Tarbagatai, Dzungar and Terskei ridges (Shil.) Zailijskij Alatau (Egorov, Ivanov, 1997). 

Ponsadenia pseudoferghanica (Schileyko, 1978) T 

Bradybaena pseudoferghanica Schileyko, 1978b: 158-160, textfigs. 114, 115, pl. 4, fig. 37. 

● Type locality: Kirgiz Ridge, near Chardovan village. 


● Distribution: Tarbagatai, Dzungar, Zailijskij and Kirgiz ridges (Likharev, Schileyko, MS). 

Ponsadenia semenovi (Martens, 1864) T 

[= Cathaica hermanni Gude, 1904; Cathaica keiseri Cockerell, 1928] 

Helix semenovi Martens, 1864a: 115, pl. 3, figs. 6-8. 

● Type locality: Tien-Shan. 


Distributuon: Tien-Shan, except for Pskem and Ugam ridges, Dzungar Ridge, Tarbagatai, southern Altai, 

Saur Ridge (Likharev, Schileyko, MS; Egorov, Ivanov, 1997). 

HELICODONTIDAE Hesse, 1907 

LINDHOLMIOLINAE Schileyko, 1978 

Lindholmiola Hesse, 1931 

Type species: Helix lens Férussac, 1821 (OD) 

Lindholmiola corcyrensis (Férussac, 1819) T 

[= Helix contorta Rossmässler, 1838] 

Helix corcyrensis Férussac, 1819: 21, pl. 69 E, figs. 1-5. 

● Type locality: not stated (Corfu Island – from the name derivation). 


● Distribution: a single finding in Durleshtsky forest near Kishinev (Schileyko, 1978b).

HYGROMIIDAE Tryon, 1866 

TRICHIINAE Ložek, 1956 

Caucasigena Lindholm, 1927 

Type species: Helix eichwaldi L. Pfeiffer, 1846 (OD) 

Caucasigena abchasica (Lindholm, 1927) T 

[= Helix pontica sensu O. Boettger, 1883 non Fischer, 186] 

Helix abchasica Lindholm, 1927b: 133. 

● Type locality: «Tagereise nördl. Suchum». 


● Distribution: northern and southern slopes of western Great Caucasus (Likharev, Schileyko, MS). 

Caucasigena armeniaca (L. Pfeiffer, 1846) T 

[= Helix alpina Menetries, 1832 non Férussac, 1822; Helix nivalis Krynicki, 1837 non Orbigny, 1835] 

Helix armeniaca Pfeiffer, 1846 (1841-1846): 77. 

● Type locality: «Armenia». 


● Distribution: southern and eastern slopes of Great Caucasus from Kazbek to Shakhdag, Bazum Ridge (Armenia), 

Kyapaz Mount (Mrovdagh Ridge, Azerbaijan) (Likharev, Schileyko, MS). 

Caucasigena eichwaldi (L. Pfeiffer, 1846) T 

[= Helix eichwaldi var. nivicola Lindholm, 1913] 

Helix eichwaldi Pfeiffer, 1846 (1841-1846): 77. 



● Distribution: eastern and northeastern slopes of Great Caucasus. Near Krstovsky pass it can be found on 

southern slopes (Likharev, Schileyko, MS). 

Caucasigena rengarteni (Lindholm, 1913) T 

[= Helix gerassimovi Lindholm, 1913] 

Helix rengarteni Lindholm, 1913: 142. 

● Type locality: central Caucasus, Baksan River valley. 


● Distribution: canyons in the north of central part of Great Caucasus, from Malka and Podkumok river valleys 

in the west to Ardon River valley in the east (Likharev, Schileyko, MS). 

Caucasigena schaposchnikovi (Rosen, 1911) T 

[= Helix antoni Rosen, 1914 non L. Pfeiffer, 1876; Helix antonwagneri Lindholm, 1927; Caucasigena 

schaposchnikovi var. balkariensis Lindholm, 1929] 

Helix schaposchnikovi Rosen, 1911: 113, pl. 3, figs. 2a-c. 

● Type locality: Oshten and Fisht mountains in western Great Caucasus. 


● Distribution: sporadically on northern slopes of Great Caucasus (Likharev, Schileyko, MS). 

Caucasigena schileykoi Tavasiev et Tavasieva, 1980 T 

Tavasiev, Tavasieva, 1980: 144, figs. 1-11. 

● Type locality: North Osetiya, Alagir district, Tamisk village, 40 km west of Vladikavkaz. 

● Holotype: Museum of Severo-Osetinsky Reserve No. M-104. 


Caucasigena thalestris (Lindholm, 1927) T 

[= Helix flaveola Mousson, 1863 non Kaleniczenko, 1853] 

Trochulus thalestris Lindholm, 1927b: 123, 130. 

● Type locality: between Sukhumi and Poti. 


● Distribution: West Caucasus from Stavropol to Poti (Likharev, Schileyko, MS). 

Caucasigena tschetschenica (Retowski, 1914) T 

Helix tschetschenica Retowski, 1914: 292. 

● Type locality: Bonos-Mta Mount (Chechnya). 


● Distribution: type locality and middle reaches of Fiagdon River (Likharev, Schileyko, MS). 

Diodontella Lindholm, 1929 

Type species: Diodontella stschukini Lindholm, 1929 (OD) 

Diodontella nubigena Lindholm, 1929 T 

Lindholm, 1929a: 210, pl. 11, fig. 2. 

● Type locality: upper reaches of Chegem River. 



Diodontella stschukini Lindholm, 1929 T 

Lindholm, 1929a: 206, pl. 11, fig. 1. 

● Type locality: lime rocks near top of Cherakh-Kend (Chegem Canyon). 


● Distribution: Chegem Canyon, 1500 m above sea level (Likharev, Schileyko, MS). 

Edentiella Polinski, 1929 

Type species: Helix edentula Draparnaud, 1805 (OD) 

Edentiella bakowskii (Polinski, 1924) T 

Fruticola bielzi var. bakowskii Polinski, 1924: 196-198, figs. 72-74. 

● Type locality: «Dora und Tatarуw in oberen Prut-Tal sowie Osthäuge der Czarnohora in den Ostkarpaten». 


● Distribution: Carpathians (Sverlova, 2006). 

Helicella Férussac, 1821 

Type species: Helix itala Linnaeus, 1758 (SD ICZN Opinion 431, 1956) 

Helicella candicans (L. Pfeiffer, 1841) T 

Helix candicans Pfeiffer, 1841: 220. 

● Type locality: Hungary, south of Tapoltza, near Balaton Lake. Monogr. Helic. viv.: Hungaria (prope 

Szigleget ad lacum Balaton...) et prope Como; var β et γ in Tauria. 


● Distribution: western part of Ukraine and Moldavia, Zhitomir and Kiev (Likharev, Schileyko, MS; Sverlova, 

2006). 

Helicopsis Fitzinger, 1833 

Type species: Helix striata Müller, 1774 (monotypy) 

Helicopsis dejecta (Cristofori et Jan, 1832) T 

[= Helix arenosa Krynicki, 1836; Helix substriata Clessin, 1881; Xerophila arenosa bistriata Puzanov, 1926;

Xerophila substriata planorum Puzanov, 1927] 

Helix dejecta Cristofori, Jan, 1832 – Rossmässler, 1838: 34, pl. 38, fig. 520. 

● Type locality: Crimea. 


● Distributon: mountain and steppe Crimea, vicinities of Melitopol and Novorossiisk, shores of Yalpug Lake 

(Odessa Region), southern Donetsk Region southern (Likharev, Schileyko, MS; Sverlova, 2006). 

● Remark: the authorship of the name is usually cited as "Cristofori et Jan, 1832" (e.g. Schileyko, 1978). However, 

there in no such name in the work of Cristofori and Jan. The earliest mentioning of the name we could find 

is that in Rossmässler (1838). Major monographs of the 19th century (e.g., Pfeiffer, 1848) also cite only Rossmässler 

as primary souce of data on the species. 

Helicopsis filimargo (Krynicki, 1833) T 

[= Helix taurica Rossmässler, 1837; Helicella lantzi Lindholm, 1926; Xerophila elata Puzanov, 1927; Xerophila 

milaschevitschi vulcanica Puzanov, 1927] 

Helix filimargo Krynicki, 1833: 435. 

● Type locality: "Habitat in collibus calcareis Tauriae (Sevastopol, ... Khersones, Kachi, Shulya)". 

● Syntypes: ZIN, MNHN. 

● Distribution: mountain Crimea, vicinities of Odessa, Lugansk Region (Likharev, Schileyko, MS; Sverlova, 

2006). 

Helicopsis instabilis (Rossmässler, 1838) T 

[= Helicella instabilis var. bakowskyana Clessin, 1879; Helix jachnoi Clessin, 1887] 

Helix instabilis Rossmässler, 1838: 33, pl. 38, fig. 518. 

● Type locality: «bei Lemberg in Galicien». 

● Types: probably in SMF. 

● Distribution: Dnestr basin (Likharev, Schileyko, MS). 

Helicopsis likharevi Schileyko, 1978 T 

Schileyko, 1978b: 212-213, textfigs. 232, 233, pl. 10, fig. 96. 

● Type locality: Kopet-Dagh, Shekhin-Dere Canyon. 


● Distribution: Kopet-Dagh (Likharev, Schileyko, MS). 

Helicopsis paulhessei (Lindholm, 1936) T 

[= Helicella gasprensis Hesse, 1934] 

Helicella paulhessei Lindholm, 1936: 439, fig. 1. 

● Type locality: «Southern coast of Crimea, Gaspra manor, Koreiz». 



Helicopsis retowskii (Clessin, 1883) T 

[= Helicella gireiorum Lindholm, 1926] 

Helicella retowskii Clessin, 1883: 47, pl. 3, fig. 1. 

● Type locality: Yalta (Crimea). 


● Distribution: Crimea (Likharev, Schileyko, MS). 

Helicopsis striata (Müller, 1774) T 

[= Helicogena lunulata Krynicki, 1833; Martha cereoflava podolica Polinski, 1922; Helicella eudakowi 

Skvortzov, 1938 nom. nud.] 

Helix striata Müller, 1774: 38. 

● Type locality: «in Saxonia». 

● Types: unknown.

● Distribution: Moldavia, western Ukraine, NW Black Sea Maritime Territory, Dnepropetrovsk and Sumy 

regions of Ukraine, Kursk and Voronezh regions of Russia (Likharev, Schileyko, MS; Sverlova, 2006). 

Hygrohelicopsis Schileyko, 1978 

Type species: Hygrohelicopsis darevskii Schileyko, 1978 (OD) 

Hygrohelicopsis darevskii Schileyko, 1978 T 

Schileyko, 1978a: 14-15, textfigs. 42-46, pl. 3, fig. 9. 

● Type locality: left bank of middle part of Chegem River. 


● Distribution: Chegem River Canyon, 2000-2500 m above sea level (Likharev, Schileyko, MS). 

Kokotschashvilia Hudec et Lezhawa, 1969 

Type species: Helix holotricha O. Boettger, 1884 (OD) 

Kokotschashvilia caucasicola (Lindholm, 1913) T 

[= Caucasigena ossetica Tavasieva, 1982] 

Helix caucasicola Lindholm, 1913: 138. 

● Type locality: central Caucasus, southern side of Krestovsky pass, 2370 m above sea level. 


● Distribution: central part of Great Caucasus (Likharev, Schileyko, MS). 

Kokotschashvilia eberhardi Schileyko, 1978 T 


● Type locality: central part of Great Caucasus between Kazbegi and Sioni villages. 


● Distribution: highland of western Great Caucasus (Likharev, Schileyko, MS). 

Kokotschashvilia holotricha (O. Boettger, 1884) T 

Helix holotricha Boettger, 1884: 151. 

● Type locality: Abkhazia, Psyrtzkha near Novyi Afon. 


● Distribution: Black Sea shore of Caucasus and eastward along northern (Maikop) and southern (Tekhuri 

River basin) slopes of Great Caucasus (Likharev, Schileyko, MS). 

Kokotschashvilia makvalae (Hudec et Lezhawa, 1969) T 

Trichia makvalae Hudec, Lezhawa, 1969: 41, figs. 1, 5. 

● Type locality: Balda village, Gegechkori district of Georgia. 


● Distribution: type locality and Kvaisa (Kvansi) village, Dzhavi district of Georgia (Likharev, Schileyko, 

MS). 

Kokotschashvilia phaeolaema (O. Boettger, 1886) T 

[= Helix phaeolama var tenuitesta Lindholm, 1913] 

Helix (Fruticocampylaea) phaeolaema O. Boettger, 1886b: 137-138, pl. 3, fig. 4 a-e. 

● Type locality: Shakh-Dagh Mount, East Caucasus. 


● Distribution: central and eastern parts of northern slopes of Great Caucasus (Likharev, Schileyko, MS). 

Kokotschashvilia tanta Schileyko, 1978 T 


● Type locality: Lebarde village, Gegechkori district of Georgia. 

● Holotype: ZIN.


Leucarchaica Schileyko et Pazylov, 1990 

Type species: Leucarchaica rudimentifera Schileyko et Pazylov, 1990 (OD) 

Leucarchaica rudimentifera Schileyko et Pazylov, 1990 T 

Schileyko, Pazylov, 1990: 871, fig. 1. 

● Type locality: Ferghana Ridge, Kara-Bulak Canyon near Arslanbob. 



Leucozonella Lindholm, 1927 

Type species: Helix rubens Martens, 1874 (OD) 

Leucozonella angulata (Westerlund, 1896) T 

[= Fruticicola boewiana Tzvetkova, 1950] 

Helix rubens var. angulata Westerlund, 1896: 186-187. 

● Type locality: «Turkesian [sic]. Artaschaty; Prov. Samarkand, Kreis Pendschekent, nahe Steinkohlgruben 

gegenüber Pisan-Pojane». 


● Distribution: from southwestern Tien-Shan to Tarbagatai and steppe regions of East Kazakhstan (Likharev, 


Leucozonella caria Schileyko, 1978 T 


● Type locality: Hissar Ridge near Khodzha-Obigarm. 


● Distribution: Hissar Ridge (Likharev, Schileyko, MS). 

Leucozonella caryodes (Westerlund, 1896) T 

[= Fruticola conospira Lindholm, 1927] 

Helix rubens var. caryodes Westerlund, 1896: 186. 

● Type locality: «Turkestan und Dshungarei. Länds dem Fl. Kugart, auf Abhängen 6000-6500’. Aslam-bob; 

Stadt Lepsinsk; Kreis Sergiopol am Fl. Baskan». 


● Distribution: central and northern Tien-Shan (Likharev, Schileyko, MS). 

Leucozonella crassicosta Schileyko, 1978 T 

Schileyko, 1978b: 180-181, textfigs. 161-164, pl. 6, fig. 55. 

● Type locality: Chatkal Ridge, left bank of Kasansai River upstream from Ala-Buka village. 


● Distribution: southwestern slopes of Chatkal Ridge along left bank of Kasansai river and northern part of 

eastern slopes of Kuramin Ridge (Likharev, Schileyko, MS). 

Leucozonella ferghanica (Lindholm, 1927) T 

Cathaica ferghanica Lindholm, 1927d: 264. 

● Type locality: near Sary-Chelek Lake. 


● Distribution: northern part of Chatkal Ridge and vicinities of Namangan (Likharev, Schileyko, MS). 

Leucozonella globuliformis (Lindholm, 1931) T 

Fruticicola globuliformis Lindholm, 1931: 54, fig. 1. 

● Type locality: near Gulcha, Alai Ridge.


● Distribution: Alai Ridge near Gulcha and near Papan village (Pazylov, Schileyko, 1992). 

Leucozonella hypophaea (Lindholm, 1927) T 

[= Fruticicola balgutensis Lindholm, 1927; Fruticicola balgutensis f. elatior Lindholm, 1927] 

Fruticicola hypophaea Lindholm, 1927c: 198. 

● Type locality: Chatkal Ridge, Dzhuvan-Suget River valley. 


● Distribution: Ferghana and Alai ridges (Likharev, Schileyko, MS). 

Leucozonella intermedia Uvalieva, 1995 T 

Uvalieva, 1995: 9-10, textfig. 3, pl. 1, fig. 3. 

● Type locality: Pskem Ridge, Kalasan-Sai ravine. 



Leucozonella mesoleuca (Martens, 1882) T 

[= Helix rubens var. limitata Westerlund, 1896] 

Helix mesoleuca Martens, 1882b: 105. 

● Type locality: Arshaty, Alai Ridge. 


● Distribution: Kirgiz, Talas, Chatkal, Ferghana, Alai, Zaalai, Turkestan and Kuramin ridges and foothills 


Leucozonella mica (Tzvetkova, 1950) T 

Fruticola mica Tzvetkova, 1950: 331. 

● Type locality: Mynzhilke Canyon near Lugovaya station, Kirgiz Ridge. 

● Lectotype (Schileyko, 1988a): ZMMU No. Lc-14407. 


Leucozonella planulata Uvalieva, 1995 T 


● Type locality: Zailijskij Ridge, Syugatinskie Mountains. 


● Distribution: Zailijskij and Kungei ridges (Uvalieva, 1995). 

Leucozonella retteri (Rosen, 1897) T 

[= Helix retteri f. minor Rosen, 1901] 

Helix retteri Rosen, 1897: 170. 

● Type locality: Amankutan near Samarkand. 


● Distribution: Hissar, Peter the Great, Turkestan and Zeravshan ridges (Likharev, Schileyko, MS). 

Leucozonella rubens (Martens, 1874) T 

[= Helix rubens var. concolor Martens, 1882; Helix rubens var. finschiana Martens, 1882; Helix rubens var. 

zeiliana Martens, 1882; Helix rubens var. subangulata Rosen, 1901] 

Helix rubens Martens, 1874: 12, pl. 1, fig. 6. 

● Type locality: basin of upper Zeravshan River. 


● Distribution: sporadically along all the mountains of Central Asia except for Kopet-Dagh (Likharev, 

Schileyko, MS).

Leucozonella rufispira (Martens, 1874) T 

Helix rufispira Martens, 1874: 9, pl. 1, fig. 38. 

[= Helix rufispira var. albidorsalis Martens, 1881; Helix rufispira var. maracandensis Rosen, 1901; Helix 

rufispira var. farabensis Rosen, 1901; Helix rufispira var. exserta Rosen, 1901; Helix rufispira var. fuscior 

Rosen, 1903; Cathaica rufispira hispida Jaeckel, 1956] 

● Type locality: vicinities of Samarkand, Pyandzhikent, Magian and Kulikalan Lake. 


● Distribution: Tien-Shan and foothills of Pamir (Likharev, Schileyko, MS). 

Leucozonella schileykoi Pazylov et Kuchbaev in Pazylov, Kuchbaev et Daminova, 2001 T 


● Type locality: Turkestan Ridge, Chuyanchi village on Aksur River. 

● Holotype: Gulistan State University. 


Leucozonella translucens Uvalieva, 1995 T 


● Type locality: Kirgiz Ridge, Tuiyk-Zhar. 


● Distribution: Kirgiz Ridge (Uvalieva, 1995). 

Nanaja Schileyko, 1978 

Type species: Nanaja cumulata Schileyko, 1978 (OD) 

Nanaja chatkalica Kuznetsov, 1996 T 

Kuznetsov, 1996: 161, fig. 1 A, B. 

● Type locality: Uzbekistan, Tashkent Region, Bostanlyk District, 6 km southward of Burchmulla settlement, 

Keksui Ridge, right bank of Chatkal River, lower third of south-western slope of Paltau Mountain. 


● Distribution: Chatkal River valley (Kuznetsov, 1996). 

Nanaja cumulata Schileyko, 1978 T 


● Type locality: Pskem Ridge, upper part of Pskem River, upstream from Nanai village. 


● Distribution: Pskem River valley (Likharev, Schileyko, MS). 

Nanaja illuminata Schileyko, 1978 T 


● Type locality: western Tien-Shan, narrow side ravine in Pskem River valley, upstream from Nanai village. 


● Distribution: Pskem River valley (Likharev, Schileyko, MS). 

Odontotrema Lindholm, 1927 

Type species: Odontotrema diplodon Lindholm, 1927 (OD) 

Odontotrema diplodon Lindholm, 1927 T 

Lindholm, 1927d: 268. 

● Type locality: near Sary-Chelek Lake. 


● Distribution: northeastern part of Chatkal Ridge (Likharev, Schileyko, MS).

Odontotrema monodom Schileyko, 1988 T 

Schileyko, 1988a: 1307, fig. 3. 

● Type locality: Ashmailya Canyon near Lugovaya station, Kirgiz Ridge. 

● Holotype: ZMMU No. Lc- 14442. 

● Distribution: western part of Kirgiz Ridge (Likharev, Schileyko, MS). 

Plicuteria Schileyko, 1978 

Type species: Helix lubomirskii Slуsarski, 1881 (OD) 

Plicuteria lubomirskii (Slуsarski, 1881) T 

[= Helix clessini Ulićny, 1885] 

Helix lubomirskii Slуsarski, 1881: 29. 

● Type locality: "na S-tym Krzyzu (Lysej Gorze) w Lipcu ... nadto ... na dziedzincu star-ego zamku w 

Ogrodziencu w Lipcu" (Upper Silesia, Poland). 


● Distribution: Carpathiansб Ciscarpathia and western Podolsk Hills (Likharev, Schileyko, MS; Sverlova, 

2006). 

Teberdinia Schileyko, 1978 

Type species: Helix zolatarevi Lindholm, 1913 (OD) 

Teberdinia flavolimbata (O. Boettger, 1883) T 

[= Helix brjanskii Rosen, 1914; Helix zolatarevi Lindholm, 1913; Caucasigena reminiscenta Schileyko, 

1978] 

Helix flavolimbatus O. Boettger, 1883: 162, pl. 5, fig. 6 a-d. 

● Type locality: Svanetiya, Esheri village. 


● Distribution: Alpine and subalpine zones of western Great Caucasus (Likharev, Schileyko, MS). 

Trochulus Chemnitz, 1786 

Type species: Helix hispida Linnaeus, 1758 (SD Herrmannsen, 1849) [=Trichia Hartmann, 1840] 

Trochulus bielzi (A. Schmidt in Bielz, 1860) T 

Helix bielzi A. Schmidt – Bielz, 1860: 199. 

● Type locality: northern Transylvania, Romania. 


● Distribution: Carpathians (Likharev, Schileyko, MS). 

Trochulus concinna (Jeffreys, 1830) T 

[= Helix nothra Westerlund, 1897] 

Helix concinna Jeffreys, 1830: 336. 

● Type locality: Great Britain, Welsh, Glamorgan near Swonsea. 


● Distribution: probably western part of Ukraine and Byelorussia (Likharev, Schileyko, MS). 

Trochulus hispidus (Linnaeus, 1758) T 

Helix hispida Linnaeus, 1758: 771. 

● Type locality: «in Svecia». 


● Distribution: nearly entire European Russia (Likharev, Schileyko, MS). 

Trochulus villosulus (Ziegler in Rossmässler, 1838) T 

[= Helix pietruskyana L. Pfeiffer, 1846]

Helix villosa var. villosula Ziegler – Rossmässler, 1838: 1. 

● Type locality: Hungary. 

● Types: probably SMF. 

● Distribution: Carpathians and Ciscarpathia (Likharev, Schileyko, MS). 

Xeropicta Monterosato, 1893 

Type species: Helix krynickii Krynicki, 1833 (monotypy) 

Xeropicta candaharica (L. Pfeiffer, 1846) T 

[= Helix millepunctata O. Boettger, 1889] 

Helix candaharica Pfeiffer, 1846: 37. 

● Type locality: «Candahar, East Indies» (Afghanistan). 


● Distribution: Central Asia from eastern Kopet-Dagh to Ferghana Ridge, Ferghana valley, Chatkal and 

western Kirgiz ridges and foothills (Likharev, Schileyko, MS). 

Xeropicta derbentina (Krynicki, 1836) T 

[= Helix derbentina var. isomera Mousson, 1863; Helix derbentina var. suprazonata Mousson, 1863] 

Helix derbentina Krynicki, 1836: 192. 

● Type locality: Pyatigorsk. 


● Distribution: Crimea; Odessa, Nikolaev, Kherson, Zaporozhye regions of Ukraine; Sakharna Reserve and 

Kishinev (Chisinau) parks in Moldova; Caucasus (Likharev, Schileyko, MS; Sverlova, 2006). 

Xeropicta krynickii (Andrzejowski in Krynicki, 1833) T 

[= Helix theodosia Clessin, 1881; Xerophila krynickii var. odessana Lindholm, 1908; Xerophila orianda Puzanov, 

1925] 

Helix krynickii Andrzejowski – Krynicki, 1833: 434. 

● Type locality: "Habitat inter montes calcareos Tauriae (Sevastopol, Inkerman, Shulya)". 


● Distribution: Crimea, Odessa, Nikolaev, Kherson regions of Ukraine; vicinities of Anapa, Novorossiisk, 

Astrakhan, Lenkoran lowland to Talysh by river valleys (Likharev, Schileyko, MS; Sverlova, 2006). 

Xeropicta parableta (O. Boettger, 1881) T 

Helix (Xerophila) parableta Boettger, 1881a: 212-213, pl. 8, fig. 15 a-c. 

● Type locality: debris of Araks River near Nakhichevan. 


● Distribution: highland-steppe zone of Talysh (Likharev, Schileyko, MS). 

HYGROMIINAE Tryon, 1866 

Cernuella Schlüter, 1838 

Type species: Helix variabilis Draparnaud, 1801 (= Cochlea virgata Da Costa, 1778) (SD Gude, Woodward, 

1921) 

Cernuella virgata (Da Costa, 1778) T 

[= Helix variabilis Draparnaud, 1801; Xerophila euxina Clessin, 1883] 

Cochlea virgata Da Costa, 1778: 79, pl. 4, fig. 8. 

● Type locality: Oxfordshire, Hempshire, Cornwall, Cambridge (England). 


● Distribution: southern coast of Crimea and vicinities of Novorossiisk (Likharev, Schileyko, MS).

Chilanodon Westerlund, 1897 T 

Type species: Helix sibirica Westerlund, 1897 (= Helix gerstfeldti Dybowski, 1901) (monotypy) 

Chilanodon bicallosa (L. Pfeiffer, 1853) T 

Helix bicallosa Pfeiffer, 1853a: 638. 

● Type locality: «Siberia». 


● Distribution: Altai, southern Tomsk and Novosibirsk regions (Likharev, Schileyko, MS; Udaloi, Novikov, 

2005b). 

Chilanodon gerstfeldti (W. Dybowski, 1901) T 

[= Helix sibirica Westerlund, 1897 non L. Pfeiffer, 1853; Chilanodon gerstfeldti var. depressior Lindholm, 

1912] 

Helix gerstfeldti Dybowski W., 1901b: 136-138, textfig. 

● Type locality: "Mundung des Ussuri in den Amour". 


● Distribution: Cisbaikalia and Transbaikalia, along Amur River basin to Khabarovsk (Likharev, Schileyko, 

MS). 

Circassina Hesse, 1921 

Type species: Helix circassica Mousson, 1863 (OD) 

Circassina christophori (Rosen, 1911) T 

[= Helix uruschtenka Lindholm, 1927, nom. nov. pro Helix christophori Rosen, 1911] 

Helix christophori Rosen, 1911: 110. 

● Type locality, "am Ufer des Uruschten, c. 30 Werst sudlich Psebai" (southern Krasnodar Territory, Russia). 


● Distribution: upper drainage area of the Malaya Laba River in the northwestern Caucasus (Hausdorf, 2001). 

Circassina frutis frutis (L. Pfeiffer, 1859) T 

[= Helix fiaveola Kaleniczenko, 1853 (partim); ?Helix colchica Mousson, 1863; Helix circassica var. rudis 

Retowski, 1889] 

Helix frutis Pfeiffer, 1859: 252. 

● Type locality: Reduktaleh" (= Redut-Kale = Qulevi, Georgia). 

● Lectotype (Hausdorf, 2001): ZMZ No. 505748a. 

● Distribution: western Georgia from Adzharia in the southeast to Kutaisi and surround 

ings of Sukhumi in the northeast; Stavropol (Hausdorf, 2001). 

Circassina frutis akramowskii Schileyko, 1972 T 

Schileyko, 1972: 1135. 

● Type locality: off Parz-Lich Lake in northern Armenia. 

● Holotype: Institute of Zoology of Academy of Sciences of Armenia. 

● Distribution: northern Armenia (Likharev, Schileyko, MS). 

Circassina frutis circassica Charpentier in Mousson, 1863 T 

[= Helix Nymphaea Mousson, 1863 (partim?); Circassina bojenae Hudec et Lezhawa, 1969] 

Helix circassica Charpentier – Mousson, 1863: 370-371. 

● Type locality: « Imereth, Nakolakevi» (Nakalakevi, Imeretia, Georgia). 

● Leclotype (Hausdorf, 2001): ZMZ No. 50575a. 

● Distribution: southern slopes of Caucasus in northern Georgia and adjacent Russia, Trialeti Ridge in southern 

Georgia (Hausdorf, 2001).

Circassina frutis veselyi (Frankenberger, 1919) T 

[= Helix aurea Rosen, 1911, non Dillwyn, 1817; Circassina circassica simpla Schileyko, 1972; Metafruticicola 

(Metafruticicola) schileykoi Hudec, 1972, nom. nov. pro Helix aurea Rosen, 1911] 

Fruticicola (Monacha) veselyi Frankenberger, 1919: 72, fig. 6. 

● Type locality: "Ananur" (Ananuri, Georgia). 

● Lectotype (Kroupa, 1988): Národní Muzeum, Praha, Czechia. 

● Distribution: north slope of Caucasus from the Belaya valley in the west to the Assa valley in the east, also 

eastern Georgia and northernmost Armenia; probable isolated occurrences on southern slopes of Caucasus 

(Abkhazia and Imereti Ridge in southwestern Georgia) (Hausdorf, 2001). 

Circassina pachnodes (O. Boettger, 1884) T 

Helix pachnodes Boettger, 1884: 150. 

● Type locality: "Uetsch-Dere" (Georgia). 

● Lectotype (Hausdorf, 2001): SMF No. 6232. 

● Distribution: southwest slope of Caucasus from the Shakhe valley in the northwest to the Bzyb valley in 

the southeast (Hausdorf, 2001). 

Circassina pergranulata Hausdorf, 2001 T 

[= Hela stephaniae Hudec et Lezhawa, 1970 (partim)] 

Hausdorf, 2001: 442-443, figs. 9, 17, 26, 30. 

● Type locality: Mamdzykha Mount above Gagra, Abkhazia, Georgia. 

● Holotype: ZMH No. 2864. 

● Distribution: surroundings of Gagra, Abkhazia, Georgia (Hausdorf, 2001). 

Circassina septentrionalis Hausdorf, 2001 T 

Hausdorf, 2001: 443-444, figs. 10,18, 27, 30. 

● Type locality: Russia: Maykop. 

● Holotype: SMF No. 138802a. 

● Distribution: north slope of Caucasus (Maikop, Dakhovskaya) and south slope as far southwest as Lazarevskoye 

(Hausdorf, 2001). 

Circassina stephaniae (Hudec et Lezhawa, 1970) T 

Hela stephaniae Hudec, Lezhawa, 1970: 16, fig. 1, 10. 

● Type locality: Novyi Afon, Abkhazia. 

● Holotype: Národní Muzeum, Praha, Czechia, No. 7066, 7067. 

● Distribution: type locality (Hausdorf, 2001). 

Fruticocampylaea Kobelt, 1871 

Type species: Helix narzanensis Krynicki, 1836 (SD Lindholm, 1928) 

Fruticocampylaea kobiensis (O. Boettger, 1883) T 

[= Helix kobiensis var. fluxa Rosen, 1914; ? Helix schelkovnikovi Rosen, 1914] 

Helix kobiensis Boettger, 1883: 166. 

● Type locality: Pasanauri near Aragvi River, Georgia. 

● Probable syntype: SMF No. 6199. 

● Distribution: southern slopes of central Great Caucasus (Likharev, Schileyko, MS). 

Fruticocampylaea narzanensis (Krynicki, 1836) T 

[= Helix narzanensis var. suanetica O. Boettger, 1883; Helix narzanensis var. macromphala O. Boettger, 

1883; Helix narzanensis var. cyclothira O. Boettger, 1883; Fruticocampylaea schamyli Lindholm, 1922] 

Helix narzanensis Krynicki, 1836: 172. 

● Type locality: «In Caucaso». 


● Distribution: almost entire Caucasus ecxept for Colchis lowland, Adzharia, Kuro-Araks lowland and Ta-

lysh (Likharev, Schileyko, MS). 

Kalitinaia Hudec et Lezhawa, 1967 

Type species: Helix schelkovnikovi Bogatschev, 1936 (= Kalitinaia perspectiva Hausdorf, 1993) (OD) 

Kalitinaia arcadiana (Schileyko, 1967) T 

Helicella arcadiana Schileyko, 1967: 28, fig. 1, 3A. 

● Type locality: Azerbaijan, Talysh, Zuvand Plateau, 2 km eastward of Kosmalyan village. 

● Holotype: ZMMU No. Lc-14479, 4844. 

● Distribution: Zuvand, Talysh (Likharev, Schileyko, MS). 

Kalitinaia crenimargo (L. Pfeiffer, 1848) T 

[= Helix crenimargo var. obtusior Mousson, 1876; ? Helix acutistria Boettger, 1880] 

Helix crenimargo Pfeiffer, 1848: 174. 



● Distribution: central and eastern Ciscaucasia, Daghestan, eastern Transcaucasia (Likharev, Schileyko, MS). 

Kalitinaia gvozdevi (Soboleva, Rymzhanov et Plachov, 1995) T 

Xerosecta gvozdevi Soboleva et al., 1995: 141-143, figs. 1-2. 

● Type locality: Aktau-Buzachinsky Reserve, 30 km west of Tauchik settlement. 

● Holotype: Institute of Zoology of National Academy of Sciences of Kazakhstan Republic. 


Kalitinaia orientalis (Uvalieva et Sacharnova in Uvalieva, 1995) T 

Xerosecta orientalis Uvalieva et Sacharnova – Uvalieva, 1995: 12-13, textfig. 5, pl. 1, fig. 5. 

● Type locality: West Kazakhstan, Obshchie Syrty. 



Kalitinaia perspectiva Hausdorf, 1993 T 

Hausdorf, 1993: 44 (nom. nov. pro Helix schelkovnikovi Bogatschev, 1936, non Helix schelkovnikovi Rosen, 

1914). 

● Type locality: Apsheron deposits (Upper Pliocene) of Kura depression near Mingechauri. 


● Distribution: near Kura River, vicinities of Kaspi (eastern Georgia) and Samukha (western Azerbaijan) 


Kalitinaia tiflisiana (Lindholm, 1913) T 

Xerophila tiflisiana Lindholm, 1913: 20. 

● Type locality: Mta-Tzminda mountain slope, Tbilisi. 


● Distribution: Tbilisi with vicinities and Darjyal Canyon (Likharev, Schileyko, MS). 

Lindholmomneme Haas, 1936 

Type species: Helix rhysota Westerlund, 1896 (= Helix nordenskioldi Westerlund, 1876) (OD) 

Lindholmomneme altaica (Westerlund, 1896) T 

Helix (Trichia) rhysota var. altaica Westerlund, 1896 

● Type locality: «Am See Jabagan» (Russia, Mountain Altai, Ust-Kan District, Anui Ridge, near Yabogan 

settlement on the shore of Yabogan Lake). 


● Distribution: West Altai, Abakan Ridge (Kuznetsov, Schileyko, 1999).

Lindholmomneme nordenskioldi (Westerlund, 1876) T 

[= Helix rhysota Westerlund, 1896; Helix rhysota var. altaica Westerlund, 1896] 

Helix (Eulota) nordenskioldi Westerlund, 1876a: 97-98. 

● Type locality: «ad Jenissei adversus Chantojskoj (lat. 68º5’), prope Podk. Tunguska (lat. 61º) et inter Krasnojarsk 

(lat. 56º) et Tomsk». 

● Syntypes: SMNH, possibly ZMMU (Kuznetsov, Schileyko, 1999). 

● Distribution: Altai, Salair and Abakan ridges (Kuznetsov, Schileyko, 1999). 

Lindholmomneme notophila (Cockerell, 1924) T 

Hygromia notophila Cockerell, 1924: 581. 

● Type locality: Primorye Territory, vicinities of Kongauz. 


● Distribution: Primorye Territory, vicinities of Kongauz, Chernigovka and coast of Peschanka Bay near 

Vladivostok (Likharev, Schileyko, MS); Sakhalin (Prozorova, Berezhok, 2004). 

Lindholmomneme rhysota (Westerlund, 1896) T 

Helix (Trichia) rhysota Westerlund, 1896: 13. 

● Type locality: «Am Fusse der Berge Kyrlygan und Alatau (im oberen Laufe von Nenj nahe Tomj)» (Russia, 

Altai Territory, Solton District, Salair Ridge, slope of Kyrlygan Mount and in mountains in upper reaches 

of the Nenya River). 

● Lectotype (Kuznetsov, Schileyko, 1999): Zoologiska Institutionen, Göteborg No. 504. 

● Distribution: Salair, Abakan, West Sayan ridges (Kuznetsov, Schileyko, 1999). 

Lindholmomneme turbinatum Uvalieva, 1995 T 

Uvalieva, 1995: 10-12, textfig. 4, pl. 1, fig 4. 

● Type locality: Semipalatinsk Region, Zhana-Semei District, near pioneer camp «Orbita» (Kazakhstan, right 

side of Irtysh River valley). 


● Distribution: Kazakhstan, Irtysh River basin (Kuznetsov, Schileyko, 1999). 

Lindholmomneme westerlundi Kuznetsov in Kuznetsov et Schileyko, 1999 T 

Kuznetsov, Schileyko, 1999: 34-36, fig. 5. 

● Type locality: Russia, Krasnoyarsk Territory, Divnogorsk District, East Sayan, "Stolby" Reserve, floodlands 

of right bank of Laletina River (right tributary of Yenisei River), 2.5-3 km southward of "Krasnoyarsk- 

Divnogorsk" road. 


● Distribution: Yenisei River valley on the territory of West and East Sayan (from the dam of Sayano-Shusha 

hydroelectric power plant to «Stolby» Reserve) (Kuznetsov, Schileyko, 1999). 

? Lindholmomneme boevi (Uvalieva, 1967) T 

Bradybaena boevi Uvalieva, 1967: 216, figs. 4, 5. 

● Type locality: north of Altai village, southern Altai. 



● Remark: Schileyko (1978b) believes that this species most probably does not belong to Bradybaenidae but 

is similar to representatives of Hygromiidae. The original description and figures do not allow to decide on its 

taxonomic position without an additional study. 

Monachoides Gude et Wodward, 1921. 

Type species: Helix incarnata Müller, 1774 (OD)

Monachoides incarnata (Müller, 1774) T 

Helix incarnata Müller, 1774: 63. 

● Type locality: not designated. 



Monachoides vicina (Rossmässler, 1842) T 

Helix vicina Rossmässler, 1842: 3, pl. 51, fig. 689. 

● Type locality: Carpathians. 


● Distribution: Ukraine: Carpathians, Transcarpathia and Ciscarpathia, Podolsk Hills, area between Prut and 

Dniester rivers, eastward to Vinnitza Region, northward to Rovno Region (Sverlova, 2006). 

Noneulota Schileyko et Horsak, 2007 

Type species: Noneulota surprisa Schileyko et Horsak, 2007 (OD) 

Noneulota khakassica Schileyko, Horsak et Cheremnov in Schileyko et Horsak, 2007 T 

Schileyko, Horsak, 2007: 71, fig. 3. 

● Type locality: Altai, Khakassia, Tashtyp setdement 



Noneulota surprisa Schileyko et Horsak, 2007 T 

Schileyko, Horsak, 2007: 70-71, figs. 1-2. 

Type locality: Russia, Altai Republic, Altai Mountains, Turochak district, Ust'-Lyzha setdement, a slope 1.5 

km NE of the setdement, 51°51'19.1"N, 87°06'57.5"E, 435 m above sea level. 



Perforatella Schlüter, 1838 

Type species: Trochus bidens Chemnitz, 1786 (= Helix bidentata Gmelin, 1791) (monotypy) 

Perforatella bidentata (Gmelin, 1791) T 

Helix bidentata Gmelin, 1791: 3642. 



● Distribution: Central European Plain (Likharev, Schileyko, MS). 

Perforatella dibothrion (Kimakowicz, 1883) T 

Fruticicola dibothrion Kimakowicz, 1883: 47. 

● Type locality: vicinities of Mukatchevo. 


● Distribution: Carpathians, Ciscarpathia, western Ukraine (Sverlova, 2006). 

Pseudotrichia Likharev, 1949 

Type species: Tricheulota shadini Likharev, 1949 (= Helix rubiginosa A. Schmidt, 1853) (monotypy) 

Pseudotrichia rubiginosa (A. Schmidt, 1853) T 

[= Fruticicola czarnohorica Polinski, 1924; Tricheulota shadini Likharev, 1949] 

Helix rubiginosa A. Schmidt, 1853: 3. 

● Type locality: vicinities of Wrocław, Poland. 


● Distribution: almost entire Palearctic (Likharev, Schileyko, MS).

Shileykoia Hudec, 1969 

Type species: Helix daghestana Kobelt, 1877 (OD) 

Shileykoia daghestana (Kobelt, 1877) T 

[= Helix (?) aliostoma Westerlund, 1898; Helix eichwaldi var. albocostata Retowski, 1914; Helix eichwaldi 

var. königi Retowski, 1914; Shileykoia lithophila Schileyko, 1972] 

Helix daghestana Kobelt, 1877: 28, pl. 126, fig. 1217. 

● Type locality: Daghestan. 

● Probable syntype: SMF No. 103244 (coll. C.R. Boettger 1906, labelled by Zilch as “Neotype”, most probably 

taken by Boettger from Kobelt’s material) (R. Janssen, pers. comm.). 

● Distribution: mountain Daghestan and slopes of eastern Great Caucasus (Likharev, Schileyko, MS). 

Stygius Schileyko, 1970 

Type species: Zenobiella aculeata Uvalieva, 1964 (OD) 

Stygius aculeata (Uvalieva, 1964) T 

Zenobiella aculeata Uvalieva, 1964: 201, figs. 1-3. 

● Type locality: Altai, vicinities of Verkhnekatunsk. 


● Distribution: Altai, southern Tomsk and Novosibirsk regions (Udaloi, Novikov, 2005b). 

Stygius stuxbergi (Westerlund, 1876) T 

[= Helix verna Westerlund, 1897; Helix czekanowskii Westerlund, 1897] 

Helix stuxbergi Westerlund, 1876b: 98. 

● Type locality: Siberia, Khantaiskoye lake, 68°5’ N. 

● Lectotype (Schileyko, 1978b): SMNH No. 1654. 

● Distribution: from Khamardaban Ridge to Dudinka along Angara and Yenisei rivers (Likharev, Schileyko, 

MS). 

Urticicola Lindholm, 1927 

Type species: Helix umbrosa C. Pfeiffer, 1828 (OD) 

Urticicola umbrosa (Partsch in C. Pfeiffer, 1828) T 

Helix umbrosa Partsch – Pfeiffer, 1828: 27-28, pl. 6, fig. 7. 

● Type locality: islands on Danube river near Vienna. 



ARCHAICINAE Schileyko, 1978 

Archaica Schileyko, 1970 

Type species: Helix apollinis Martens, 1882 (OD) 

Archaica apollinis (Martens, 1882) T 

[= Helix diaphora Westerlund, 1896; Cathaica apollinis nubila Lindholm, 1931] 

Helix apollinis Martens, 1882b: 105. 

● Type locality: «Andidjan in valle fluvii Naryn». 


● Distribution: Ferghana and Alai ridges (Likharev, Schileyko, MS).

Archaica heptapotamica (Lindholm, 1927) T 

[= Cathaica tianschanica Lindholm, 1927; Cathaica kokpoktasica Tzvetkova, 1950; Cathaica heptapotamica 

dshambuli Tzvetkova, 1950; Zenobiella tzwetkovi Matiokin, 1966] 

Cathaica heptapotamica Lindholm, 1927d: 265. 

● Type locality: Kuldja and Issyk-Kul. 

● Syntypes: ZIN, ZMMU No. 14410 

● Distribution: northwestern Ferghana and Chatkal and central Talas ridges westward to westernmost part of 

Tien-Shan; further along Kirgiz Ridge eastward to Issyk-Kul and adjacent regions of China (Likharev, 


Archaica labianix Schileyko, 1978 T 

Schileyko, 1978b: 261, pl. 14, fig. 139, textfigs. 323-327. 

● Type locality: upper reaches of Pskem River. 


● Distribution: upper reaches of Pskem River (Likharev, Schileyko, MS). 

Archaica papanica Pazylov et Schileyko, 1992 T 

Pazylov, Schileyko, 1992: 61-62, fig. 2. 

● Type locality: Papan village on Ak-Buura River, Alai Ridge. 



Archaica suspecta Schileyko et Moisseeva, 1989 T 

Schileyko, Moisseeva, 1989: 141, figs. 2, I; 3, I. 

● Type locality: Yangyskyr Mountains, valley of Pchan River (right tributary of Ala-Buka River, Naryn river 

basin). 

● Holotype: ZMMU No. Lc-14440, 19084. 


EUOMPHALIINAE Schileyko, 1978 

Euomphalia Westerlund, 1889 

Type species: Helix strigella Draparnaud, 1801 (SD Kobelt, 1904) 

Euomphalia appeliana (Mousson, 1876) T 

[= Helix narzanensis var. appeliusi f. depressa O. Boettger, 1883] 

Helix appeliana Mousson, 1876a: 32-33, pl. 2, fig. 3. 

● Type locality: Kislovodsk. 


● Distribution: horseshoe area around western wing of Great Caucasus from Kislovodsk to Novorossiisk, 

along coast of Black Sea and Abkhazia (Likharev, Schileyko, MS). 

Euomphalia aristata (Krynicki, 1836) T 

[= ?Helix (Trichia) chrysotricha Boettger, 1888] 

Helix aristata Krynicki, 1836: 179. 

● Type locality: «In Caucaso (Narzania ???)». 


● Distribution: central and western parts of North Caucasus and Georgia, southward to Imereti Ridge (Likharev, 


Euomphalia strigella (Draparnaud, 1801) T 

[= Helix podolica Westerlund, 1897]

Helix strigella Draparnaud, 1801: 84-85. 



● Distribution: East-European Plain to 61º N in the north, to the Black and Caspian seas in the south, to left 

bank of the Volga in the east; isolated findings in Ural (Belaya River basin and vicinities of Ekaterinburg) 


Karabaghia Lindholm, 1927 

Type species: Perforatella bituberosa Lindholm, 1927 (OD) 

Karabaghia bituberosa (Lindholm, 1927) T 

[= Helix bifrons Rosen, 1914 non Lowe, 1833] 

Perforatella bituberosa Lindholm, 1927b: 124. 

● Type locality: Nagornyi Karabakh, Zindzhirlu Ridge. 


● Distribution: highland of Nagornyi Karabakh, Pambak and Sevan ridges in Armenia (Likharev, Schileyko, 

MS). 

Monacha Fitzinger, 1833 

Type species: Helix cartusiana Müller, 1774 

Monacha cartusiana (Müller, 1774) T 

Helix cartusiana Müller, 1774: 15. 

● Type locality: «in Gallia». 


● Distribution: southern Ukraine, Crimea, vicinities of Novorossiisk (Likharev, Schileyko, MS); several 

probably introduced populations in western Ukraine (Sverlova, 2006). 

Monacha ciscaucasica Hausdorf, 2000 T 

Hausdorf, 2000: 1586-1587, figs. 5, 14. 

● Type locality: Pyatigorsk, Mashuk Mount. 


● Distribution: surroundings of Pyatigorsk in North Caucasus (Hausdorf, 2000). 

Monacha claussi Hausdorf, 2000 T 

Hausdorf, 2000: 1576-1580, figs. 1, 10. 

● Type locality: Russia: gorge between Monastyrka and Kepsh. 


● Distribution: Mzymta valley (Hausdorf, 2000). 

Monacha fruticola (Krynicki, 1833) T 

Helix fruticola Krynicki, 1833: 429-430. 

● Type locality: "Habitat in montosis fruticetis Tauriae meridionalis (Simferopol, Shulya, Baidarskaya 

dolina, Yuzhnyi bereg)". 


● Distribution: Crimea, introduced to Odessa, Nikolaev, Kherson regions, found in Zaporozhje (Likharev, 


Monacha kuznetsovi Hausdorf, 2000 T 

Hausdorf, 2000: 1581-1583: figs. 2. 11. 

● Type locality: Russia: SW slope of the Markoth Gora NE of Gelendzhik. 


● Distribution: type locality (Hausdorf, 2000).

Monacha perfrequens (Hesse, 1914) T 

[= Helix perfrequens O. Boettger, 1883 nom. nud.; Helix frequens var. obscura Mousson, 1863 non Müller, 

1774; Helix frequens f. minor O. Boettger, 1884] 

Monacha samsunensis perfrequens Hesse, 1914: 259. 

● Type locality: «Kutais». 

● Lectotype (Hausdorf, 2000): SMF No. 6572. 

● Distribution: western Caucasus (Hausdorf, 2000). 

Monacha roseni (Hesse, 1914) T 

[= Helix carascaloides O. Boettger, 1884 partim non Bourguignat, 1856; Theba elevata Hesse, 1931; 

Monacha caucasicola Schileyko, 1978; Monacha tschegemica Schileyko, 1988] 

Helix orientalis roseni Hesse, 1914: 263. 

● Lectotype (Hausdorf, 2000:1584): SMF No. 6606a. 

● Distribution: western Caucasus (Hausdorf, 2000). 

Monacha samsunensis (L. Pfeiffer, 1868) T 

[= Helix bifaria Westerlund, 1889; Helix perfrequens O. Boettger, 1891; Theba scrobiculosa Lindholm, 

1922; Theba scrobiculosa var. perforata Lindholm, 1922] 

Helix samsunensis L. Pfeiffer, 1868: 480. 

● Type locality: Samsun, Northern Turkey. 


● Distribution: along the coast of the Black Sea between Gelendzhik and Psou valley (Hausdorf, 2000). 

Monacha subcarthusiana (Lindholm, 1913) T 

[= Helix carascaloides O. Boettger, 1884 non Bourguignat, 1856] 

Helix subcartusiana Lindholm, 1913: 139. 

● Type locality: Sochi, on slope of mountain with lighthouse. 


● Distribution: western part of North Caucasus from Pyatigorsk to Black Sea and southward along the coast 

to Colchis lowland (Likharev, Schileyko, MS). 

Monacha talischana (Martens, 1880) T 

[= Theba longiflagellata Schileyko, 1968; Theba maxima Schileyko, 1968] 

Helix talischana Martens, 1880: 154. 

● Type locality: Lenkoran. 


● Distribution: Lenkoran lowland and zone of mountain forests of Talysh (Likharev, Schileyko, MS). 

Oscarboettgeria Lindholm, 1927 

Type species: Helix euages O. Boettger, 1883 (OD) 

Oscarboettgeria euages (O. Boettger, 1883) T 

[= Helix gagriensis Rosen, 1911] 

Helix euages O. Boettger, 1883: 161, pl. 4, fig. 2, pl. 6, fig. 1. 

● Type locality: Psyrtzkha near Novyi Afon. 


● Distribution: western slopes of Great Caucasus and adjacent maritime parts (Likharev, Schileyko, MS). 

Platytheba Pilsbry, 1894 

Type species: Helix nummus Ehrenberg, 1831 (OD)

Platytheba mingrelica (Hesse, 1921) T 

[= Helix jasonis Mousson, 1861 non Mayer, 1856; Helix argonautarum Lindholm, 1922] 

Circassina mingrelica Hesse, 1921: 67. 

● Type locality: Nakelakevi, Georgia. 


● Distribution: western and southwestern slopes of Great Caucasus (Likharev, Schileyko, MS). 

Platytheba prometheus (O. Boettger, 1883) T 

Helix prometheus Boettger, 1883: 159, pl. 4, fig. 6. 

● Type locality: southern slopes of Great Caucasus, Muri village on Tzkhenistzkali River. 


● Distribution: southern slopes of Great Caucasus between basins of Tzkhenistzkali and Aragvi rivers (Likharev, 


Prostenomphalia Baidashnikov, 1985 

Type species: Prostenomphalia carpathica Baidashnikov, 1985 (OD) 

Prostenomphalia carpathica Baidashnikov, 1985 T 

Baidashnikov, 1985: 206, figs. 1, 2. 

● Type locality: Zakarpatskaya Region, Rakhov District, fir tree forest on bank of the Strogovetz river 12.5 

km from Luga village. 


● Distribution: Zakarpatskaya Region, Rakhov District (Likharev, Schileyko, MS). 

Stenomphalia Lindholm, 1927 

Type species: Helix selecta Klika, 1894 (OD) 

Stenomphalia maiae (Hudec et Lezhawa, 1969) T 

Euomphalia maiae Hudec, Lezhawa, 1969: 45, figs. 3, 7. 

● Type locality: Zeleny Mys, 10-15 km from Batumi. 


● Distribution: Zeleny Mys, Batumi Botanical garden, mouth of Machakheli-Tzkali river (Likharev, 


Stenomphalia pisiformis (L. Pfeiffer, 1846) T 

[= Helix arpatschaiana Mousson, 1873; Helix pseudoglobula Mousson, 1873; Euomphalia pisiformis var. 

achtaensis Minasyan, 1947; Euomphalia assadovi Likharev et Rammelmeier, 1952] 

Helix pisiformis L. Pfeiffer, 1846: 95. 



● Distribution: Caucasus, probably Kopet-Dagh (Likharev, Schileyko, MS). 

Stenomphalia ravergiensis (Férussac, 1835) T 

[= Helix ravergii Krynicki, 1836 nom emend. pro Helix ravergiensis Férussac, 1835; Helix ravergieri auct. 

nom emend. pro Helix ravergiensis Férussac, 1835; Helix caucasica L. Pfeiffer, 1845; Helix ravergiensis var. 

transcaucasica Mousson, 1863] 

Helix ravergiensis Férussac, 1835: 21. 



● Distribution: widely but sporadically distributed in North Caucasus, Daghestan and Transcaucasia (Likharev, 

Schileyko, MS); southeastern Ukraine (Sverlova, 2006).

Stenomphalia selecta (Klika, 1894) T 

[= Helix tschorochensis Lindholm, 1913; Theba chrysomallis Lindholm, 1922; Helix globula Kaleniczenko, 

1853 non Lea, 1837; Helix? inflata Mousson, 1863 non Deshayes, 1830] 

Helix selecta Klika, 1894: 2, 4. 

● Type locality: Dilizhan, Armenia. 


● Distribution: North Caucasus, Transcaucasia (absent in Talysh and Kuro-Araks lowland) (Likharev, 


PAEDHOPLITINAE Schileyko, 1978 

Angiomphalia Schileyko, 1978 

Type species: Helix regeliana Martens, 1882 (OD) 

Angiomphalia almalensis Schileyko et Uvalieva in Schileyko, 1978 T 

Schileyko, 1978b: 297-298, pl. 17, fig. 167, textfigs. 398-399. 

● Type locality: farm Almaly, Chu River valley near Tokmak village. 


● Distribution: Chu River valley (Likharev, Schileyko, MS). 

Angiomphalia caelestimontana (Tzvetkov, 1940) T 

Cathaica caelestimontana Tzvetkov, 1940a: 390. 

● Type locality: Zailijskij Ridge, western slope of Karagalinsky Canyon. 

● Syntypes: ZMMU No. Lc-3943, Lc-14411. 

● Distribution: ridges of Tien-Shan, to south, west and north from Issyk-Kul Lake (Likharev, Schileyko, 

MS). 

Angiomphalia copiosa Schileyko, 1978 T 

Schileyko, 1978b: 292-293, pl. 17, fig. 162, textfigs. 382-384. 

● Type locality: floodland in upper reaches of Chatkal River. 


● Distribution: upper reaches of Chatkal River (Likharev, Schileyko, MS). 

Angiomphalia exasperata Schileyko et Uvalieva in Schileyko, 1978 T 

Schileyko, 1978b: 295-296, pl. 17 fig. 165, textfigs. 390-392. 

● Type locality: vicinities of Bolshoye Almaatinskoye lake. 


● Distribution: Zailijskij, Kungei and eastern part of Kirgiz ridges (Likharev, Schileyko, MS). 

Angiomphalia regeliana (Martens, 1882) T 

Helix rubens var. regelianus Martens, 1882a: 9, pl. 2, fig. 4. 



● Distribution: Tien-Shan and northern offshoots of Pamir (Likharev, Schileyko, MS). 

Angiomphalia seductilis (Westerlund, 1898) T 

Helix seductilis Westerlund, 1898: 157. 

● Type locality: «Kaschkara». 

● Syntypes: NMG No. 577. 

● Distribution: Zailijskij and Kungei ridges, western part of Terskei Ridge and eastern part of Kirgiz Ridge 

(Likharev, Schileyko, MS).

Paedhoplita Lindholm, 1927. 

Type species: Paedhoplita laminata Lindholm, 1927 (OD) 

Paedhoplita buamica (Tzvetkov et Tzvetkova, 1943) T 

Euomphalia buamica Tzvetkov et Tzvetkova, 1943: 118. 

● Type locality: Buam Canyon of Chu river, 59 km from Rybachje village. 

● Syntypes: ZMMU No. Lc-14471, 14419, 14422. 

● Distribution: canyons of Chu and Arasan rivers in eastern part of Kirgiz Ridge (Likharev, Schileyko, MS). 

Paedhoplita kirgisensis Likharev et Rammelmeier, 1952 T 

Likharev, Rammelmeier, 1952: 411, fig. 342. 

● Type locality: Kirgiz Ridge, upper reaches of Alamedin River. 



Paedhoplita laminata Lindholm, 1927 T 

Lindholm, 1927d: 266. 

● Type locality: Kirgiz Ridge. 


● Distribution: eastern part of Kirgiz Ridge (Likharev, Schileyko, MS). 

Paedhoplita lentina (Martens, 1885) T 

[= Paedhoplita mesasiana Lindholm, 1927] 

Helix lentina Martens, 1885: 18. 

● Type locality: Chatkal Ridge. 


● Distribution: northern part of Ferghana Ridge and eastern slopes of Chatkal Ridge (Likharev, Schileyko, 

MS). 

Paedhoplita lindholmi Tzvetkov et Tzvetkova, 1943 T 

Tzvetkov, Tzvetkova, 1943: 119. 

● Type locality: Kirgiz Ridge, Karabalty Canyon, rocks near Uzun-Bulak River, 1900-2400 m above sea 

level. 

● Syntypes: ZMMU No. Lc-14412, 14402. 

● Distribution: northern slopes of middle and eastern parts of Kirgiz Ridge, Zailijskij Ridge (Likharev, 


HESSEOLINAE Schileyko, 1990 

Hesseola Lindholm, 1927 

Type species: Helix adshariensis Lindholm, 1913 (= Helix solidior Mousson, 1873) (OD) 

Hesseola bactriana (Hutton, 1849) T 

[=Helix transcaspia Boettger, 1889] 

Helix bactriana Hutton, 1849: 651. 

● Type locality: Kandagar. 


● Distribution: Kopet-Dagh (Likharev, Schileyko, MS). 

Hesseola solidior (Mousson, 1873) T 

[= Helix christophori Rosen, 1911; Helix adshariensis Lindholm, 1913; Hesseola adshariensis antontomashevici 

Hudec, 1972; Metafruticicola pratensis Akramowski, 1949 partim non L. Pfeiffer, 1845]

Helix narzanensis var. solidior Mousson, 1873: 200-201. 

● Type locality: «Kisikoparan, Aschich-Dade» (Armenia). 


● Distribution: western and central parts of Lesser Caucasus, a single finding in North Caucasus (Urushten 

River valley) (Likharev, Schileyko, MS). 

METAFRUTICICOLINAE Schileyko, 1972 

Caucasocressa Hesse, 1921 

Type species: Helix joannis Mortillet, 1854 (OD) 

Caucasocressa ibera Hausdorf, 2003 T 

[= Helix pratensis var. depressa Kobelt, 1877; Helix pratensis var. unicingulata Retowski, 1914; Metafruticicola 

& Caucasocressa pratensis auctt., non L. Pfeiffer, 1846] 

Hausdorf, 2003: 2634-2635, figs. 2, 6. 

● Type locality: Georgia, Borzhomi. 


● Distribution: Georgia (Hausdorf, 2003). 

Caucasocressa joannis (Mortillet, 1854) T 

[= Helix joannis var. minor Mortillet, 1854; Helix delabris Mousson, 1863; Helix joannis var. andronakii 

Lindholm, 1913; Helix joannis f. pellucidepunctata Retowski, 1914; Fruticocampylaea pratensis karabaghensis 


Helix joannis Mortillet, 1854: 8, pl. 1, fig. 9. 

● Type locality: middle reaches of Chorokh River, vicinities of Ispira village, Turkey. 


● Distribution: Adzharia (Hausdorf, 2003). 

POLYGYRIDAE Pilsbry, 1895 

Triodopsis Rafinesque, 1819 

Type species: Triodopsis lunula Rafinesque, 1831 (= Helix tridentata Say, 1816) (SD Herrmannsen, 1849) 

Triodopsis subpersonatum (Middendorff, 1851) T 

Helix subpersonatum Middendorff, 1851: 306, pl. 30, figs. 27-29. 

● Type locality: vicinities of Udskoe (near of the Okhotsk Sea coast). 

● Holotype: ZIN (probably lost). 

● Distribution: type locality and Ayan (Likharev, Schileyko, MS). 

Class BIVALVIA 

Subclass PALEOHETERODONTA 

Order UNIONOIDA 

UNIONOIDEA

MARGARITIFERIDAE Henderson, 1929 


Dahurinaia Starobogatov, 1970 

Type species: Unio dahuricus Middendorff, 1850 (OD) 

● Remark: Graf (2007) considered all species assigned hereign to the genus as a mere synonyms of Margaritifera 

dahurica (Middendorff, 1850). 

Dahurinaia dahurica (Middendorff, 1850) F 

Unio dahuricus Middendorff, 1850: 3; 1851: 275, Taf. XXVI, fig. 3-5. 

● Type locality: Argun River at the place of its joining with the Shylka River. 

● Lectotype(selected by Bogatov et al., 2003): ZIN No. 7a in systematic catalogue. 

● Distribution: basin of Amur River, Hanka Lake (Zatrawkin, Bogatov, 1987). 

Dahurinaia komarovi Bogatov, Prozorova et Starobogatov, 2003 F 

Bogatov et al., 2003: 45-46, fig. 3 A,D. 

● Type locality: Komarovka (Suputinka) River near the Komarov's house, Ussurijsky Nature Reserve, Razdolnaya 

(Sujfun) River drainage, Primorye Territory 


● Distribution: small rivers of the basin of Razdolnaya [Sujfun] River (Bogatov et al., 2003; Starobogatov et 

al., 2004). 

● Remark: Graf (2007) considered this species as a synonyms of Margaritifera dahurica (Middendorff). 

Dahurinaia prozorovae Bogatov et Starobogatov in Bogatov, Prozorova et Starobogatov, 2003 F 

Bogatov et al., 2003: 48, fig. 3 C,I. 

● Type locality: Bolshaya Ussurka (Iman) River 30 km downstream of the Melnichnoye settlement, Ussuri 

River drainage, Primorye Territory. 


● Distribution: rivers of the basin of Ussuri River (Bogatov et al., 2003). 


Dahurinaia sujfunensis Moskvicheva, 1973 F 

Moskvicheva, 1973b: 1468, fig. 3, ä-ж [d-zh]. 

● Type locality: Suputinka (Komarovka) River (left tributary of the river Razdolnaya). 


● Distribution: Type locality; small rivers of the basin of Razdolnaya [Suifun] River (Moskvicheva, 1973b; 

Zatrawkin, Bogatov, 1987; Starobogatov et al., 2004). 


Dahurinaia tiunovae Bogatov et Zatrawkin, 1988 F 

Dahurinaia (Dahurinaia) tiunovae Bogatov, Zatrawkin, 1988:156-158, fig. 1. 

● Type locality: Komissarovka River near vilage Dvoryanka. 


● Distribution: small rivers mostly of the basin of Ussuri River (Starobogatov et al., 2004). 


Dahurinaia transbaicalica Klishko, 2008 F 

Klishko, 2008: 292-296, Figs. 1-6. 

● Type locality: Arei Lake, Uletovskyi Region, Chita Area, Transbaikalia. 

● Holotype: ZIN № 1/504-2007.

● Distribution: Arei Lake, Ingoda and Onon rivers, Transbaikalia, possibly in other rivers of upper Amur basin 

(Klishko, 2008). 

Dahurinaia ussuriensis Bogatov, Prozorova, Starobogatov, 2003 F 

Bogatov, Prozorova, Starobogatov, 2003: 47-48, fig. 3 B,G. 

● Type locality: Komissarovka River (Sintukha) near the Barabash-Levada settlement, Khanka Lake drainage, 

Primorye Territory. 


● Distribution: rivers of the basin of Amur River (Bogatov et al., 2003). 


Kurilinaia Zatrawkin et Bogatov in Bogatov et Zatrawkin, 1988 

Type species: Dahurinaia kurilensis Zatrawkin et Starobogatov, 1984 (OD) 

Kurilinaia kamchatica Bogatov, Prozorova et Starobogatov, 2003 F 

Bogatov, Prozorova, Starobogatov, 2003: 48, fig. 4 A, C. 

● Type locality: Golygina River, Kamchatka Peninsula. 


● Distribution: small rivers of south and south-west of Kamchatka (Bogatov et al., 2003; Starobogatov et al., 

2004). 

● Remark: Graf (2007) considered this species as a synonyms of Margaritifera middendorffi (Rosen, 1926). 

Kurilinaia kurilensis (Zatrawkin et Starobogatov, 1984) F 

[ = Dahurinaia shigini Zatrawkin et Bogatov, 1987] 

Dahurinaia kurilensis Zatrawkin, Starobogatov, 1984: 1789-1790, fig. 11-14. 

● Type locality: Shikotan Island [South Kurile Islands], Krabovaja Bay, river mouth near the sea tidal zone. 


● Distribution: southern Kurile Islands, possibly southern Sakhalin (Zatrawkin, Bogatov, 1987; Bogatov et 


● Remark: Graf (2007) considered this species as a synonyms of Margaritifera laevis (Haas, 1910). 

Kurilinaia laevis (Haas, 1910) F 

[= Margaritana sachalinensis Shadin, 1938] 

Ptychorhynchus laevis Haas, 1910: 498. 

● Type locality: “Saghalin Island”. 


● Distribution: Tym’ River, Sakhalin (Bogatov et al., 2003). 

Kurilinaia middendorffi (Rosen, 1926) F 

[= Unio complanatus Middendorff, 1851, Dillwyn, 1817, nor Barnes, 1823] 

Margaritana middendorffi Rosen, 1926: 269. 

● Type locality: Mekeshino Lake, Kamchatka. 

● Lectotype (Bogatov et al., 2003): ZIN No. 6 in systematic catalogue. 

● Distribution: small rivers and lakes of south and south-west of Kamchatka (Zatrawkin, Bogatov, 1987; 


Kurilinaia zatravkini Bogatov, Prozorova et Starobogatov, 2003 F 

Bogatov et al., 2003: 49-50, fig. 4 B, F. 

● Type locality: spring near Golovnino settlement, south part of Kunashir Island. 


● Distribution: lakes and small rivers of southern Kurile Islands and Sakhalin (Bogatov et al., 2003; Starobogatov 

et al., 2004). 

● Remark: Graf (2007) considered this species as a synonyms of Margaritifera laevis (Haas, 1910).

Margaritifera Schumacher, 1816 

Type species: Mya margaritifera Linnaeus, 1758 (monotypy) 

Remark: Bogatov et al. (2003) and later Starobogatov et al. (2004) by the means of comparatory method recognized 

three species of the genus: Margaritifera margaritifera (Linnaeus, 1758), Margaritifera borealis 

(Westerlund, 1871), and Margaritifera elongata (Lamarck, 1819). Sergeeva et al. (2008) on the basis of morphometry 

of more than 300 shells from Russian north (including material from ZIN, identified as Margaritifera 

elongata) came to the conclusion that only one species can be recognized, Margaritifera margaritifera. 

This conclusion concurs with the generally accepted viewpoints to taxonomy of the genus. 

Margaritifera margaritifera (Linnaeus, 1758) F 

Mya margaritifera Linnaeus, 1758: 671. 

● Type locality: “in totius orbis arctici cataractis”. 

● Probable syntypes: UUZM Nos. 811, 1419. 

● Distribution: rivers of northern Europe (Bogatov et al., 2003; Starobogatov et al., 2004). 

UNIONIDAE Rafinesque, 1820 


PSILUNIONINAE Starobogatov, 1970 

Crassiana Servain, 1882 

Type species: Unio crassus Philipsson in Retzius, 1788 (fixation of the type species not traced) 

● Remark: Drouët (1881) described Unio sieversi from Mingrelia (Georgia), which was later considered as a 

subspecies of Unio crassus. The subspecies was thought to inhabit rivers and brooks of Transcaucasia 

(Shadin, 1952; Akramowski, 1976). In later publications, eg. Zatrawkin, 1983, Antonova, Starobogatov, 1988 

and Graf, 2007, this subspecies was not considered as valid. 

Crassiana crassa crassa (Philipsson in Retzius, 1788) F 

[= Unio litoralis Rossmaessler, 1835; Unio pseudolitoralis Clessin, 1884] 

Unio crassus Philipsson – Retzius, 1788: 17. 

● Type locality: “in fluviis Europaea”. 


● Distribution: European part of Russia, Ukraine, Ural River (Stadnichenko, 1984; Pirogov, Tarasov, Kazantseva, 

1994). 

Crassiana cyprinorum (Bourguignat in Locard, 1882) F 

Unio cyprinorum Bourguignat – Locard, 1882: 289, 358. 

● Type locality: "La Seine а Chatou, dans Seine-et-Oise". 


● Distribution: rivers of Europe, except northern part, Ural river (Pirogov, Tarasov, Kazantseva, 1994; Starobogatov 

et al., 2004). 

● Remark: Graf (2007) considered this species as a synonym of Unio crassus crassus Philipsson in Retzius, 

1788. 

Crassiana fuscula (Ziegler in Rossmässler, 1835) F 

Unio fusculus Ziegler – Rossmässler, 1835b: 21. 

● Type locality: “Österreich: Kaltengang”. 


● Distribution: Europe, except northern part, Ural river (Pirogov, Tarasov, Kazantseva, 1994; Starobogatov et 

al., 2004).


1788. 

Crassiana gregorii (Kobelt, 1912) 

Unio gregorii Kobelt, 1912: 30, Taf. XDIV, fig. 2632. 

● Type locality: "Aufenthalt im Küstenflüsschen Schebs bei Grigoreffsk südlich von der Rionmündung". 


● Distribution: Caucasian Black Sea Coast (Antonova, Starobogatov, 1988). 

● Remark: Graf (2007) recognized this as a subspecies Unio crassus gontieri distributed in Ukraine to 

Transcaucasia. 

Crassiana irenjensis (Kobelt, 1912) F 

Unio kungurensis irenjensis Kobelt, 1912: 26-27, pl. 491, fig. 2626. 

● Type locality: "in Flüsschen Irenj bei Kungur". 


● Distribution: Europe, Ukraine (Stadnichenko, 1984). 


1788. 

Crassiana irgizlaica (Lindholm, 1904) F 

Unio ater var. irgizlaicus Lindholm, 1904: 343. 

● Type locality: "Fluss Irgizla" (Irgizla River, near Orenburg). 

● Syntypes: ZIN?. 

● Distribution: lower Ural, basins of Ural and Volga rivers (Pirogov et al., 1994; Starobogatov et al., 2004). 


1788. 

Crassiana mengrelica (Drouët, 1881) F 

Drouët, 1881: 16. 

● Type locality: "la Mingrélie". 


● Distribution: small rivers of Transcaucasia (Shadin, 1952). 



Crassiana musiva musiva (Spengler, 1793) F 

Unio musivus Spengler, 1793: 57. 

● Type locality: “i Floredne i Tydskland” [Germany]. 


● Distribution: Europe (Pirogov et al., 1994; Starobogatov et al., 2004). 


1788. 

Crassiana musiva gontieri (Borguignat, 1857) F 

Unio gontieri Borguignat, 1857: 16, pl. 4, figs. 10-12. 

● Type locality: "la Crimée, dans la Tchernaia". 





Crassiana nana nana (Lamarck, 1819) F 

Unio nana Lamarck, 1819: 76.

● Type locality: "dans la France – Comté". 


● Distribution: Europe, except northern part, Ural river (Pirogov, Tarasov, Kazantseva, 1994; Starobogatov et 

al., 2004). 


1788. 

Crassiana nana carnea (Küster, 1854) F 

Unio carneus Küster, 1854: 103-104, pl. 28, figs. 1, 2. 

● Type locality: "Montenegro, in einem Bach, einem Zuflusse des Moraka-flusses". 


● Distribution: Europe, except northern part, Ukraine (Stadnichenko, 1984). 

Crassiana sobriewskii (Rosén, 1925) 

Unio sobriewskii Rosén, 1925: 116-121. 






Crassiana stevenianiformis (Rosén, 1925) 

Unio mingrelicus var. stevenianiformis Shadin, 1938: 96, fig. 28. 

● Type locality: Gubis-Tshali River near Kutaisi, Geogria. 

● Types: not traced, may be in ZIN? 




Eolymnium Prashad, 1919 

Type species: Unio terminalis Bourguignat, 1852 (fixation of the type species not traced) 

● Remark: Graf (2007) considered all species assigned hereign to the genus as mere synonyms of Unio crassus 

s.lato. 

Eolymnium araxenum (Drouёt, 1881) F 

Unio araxenus Drouёt, 1881: 18 

● Type locality: Arax River. 


● Distribution: Arax River (Antonova, Starobogatov, 1988). 

Eolymnium byzantinum (Drouёt, 1879) F 

Unio byzantinus Drouёt, 1879: 171. 

● Type locality: Sperchios, Greece (fide Westerlund, 1890). 


● Distribution Western Georgia (Antonova, Starobogatov, 1988). 

Eolymnium colchicum (Drouёt, 1881) F 

Unio colchicus Drouёt, 1881: 18 

● Type locality: “la Mingrélie” (Georgia). 


● Distribution: Western Georgia (Antonova, Starobogatov, 1988). 

Eolymnium koutaisianum (Kobelt, 1886) F

Unio raddei var. koutaisiana Kobelt, 1886: 40, Taf. LII, fig. 283. 

● Type locality: “aus Bächen bei Koutais erhalten” (Georgia). 

● Types: SMF ? 

● Distribution: Western Georgia (Antonova, Starobogatov, 1988). 

Eolymnium raddei (Drouët, 1881) F 

Unio raddei Drouёt, 1881: 17 

● Type locality: “le Rion, en Mingrélie” (Georgia). 


● Distribution: Western Georgia (Antonova, Starobogatov, 1988), vicinities of Odessa, Ukraine (Drouёt, 

1881). 

Potomida Swainson, 1840 

Type species: Unio semirugatus Lamarck, 1799 (fixation of the type species not traced) 

● Remark. Graf (2007) considered all species assigned hereign to the genus as mere synonyms of Unio littoralis 

(Cuvier, 1789). 

Potomida armeniaca (Kobelt, 1912) F 

Unio armeniacus Kobelt, 1912:29, Taf. XDIII, fig. 2631. 

● Type locality: “Aufenthalt in dem Bach Karasu in der Nähe des Dorfer Aralucha, kreis Surmalin, Gouv. 

Eriwan” (Armenia). 


● Distribution: Arax River drainage basin (Shadin, 1938; Antonova, Starobogatov). 

Potomida komarowi (O. Boettger, 1880) F 

Unio rothi var. komarowi O. Boettger, 1880c: 160, Taf. V, fig. 1a-b.. 

● Type locality: “im Fluss von Kars in Russisch-Armenien”. 


● Distribution: Arax River drainage basin (Shadin, 1938; Antonova, Starobogatov). 

UNIONINAE Rafinesque, 1820 

Lanceolaria Conrad, 1853 

Type species: Unio grayanus Lea, 1834 (monotypy) 

Lanceolaria bogatovi Zatrawkin et Starobogatov, 1984 F 

Zatrawkin, Starobogatov, 1984: 1785-1787, fig. 1-4. 

● Type locality: Amur River, Sakhaljan Island, against Amursk city, Khabarovsk Territory [Far East]; inner 

lake connected with the Amur River. 


● Distribution: Amur River, its tributaries, Ussuri River, Khanka Lake (Zatrawkin, Bogatov, 1987). 

● Remark: Graf (2007) considered this species as a synonym of Lanceolaria cylindrica (Simpson, 1900). 

Lanceolaria chankensis Moskvicheva, 1973 F 

Moskvicheva, 1973b: 1466, fig. 2, к, л [k, l]. 

● Type locality: Mo (Melgunovka) River. 


● Distribution: Khanka Lake and rivers, received by it (Moskvicheva, 1973b; Zatrawkin, Bogatov, 1987). 


Lanceolaria maacki Moskvicheva, 1973 F 

[= Unio grayanus sensu Schrenck, 1867, non Lea, 1834; Nodularia cylindrica Simpson, 1900, part.]

Moskvicheva, 1973b: 1465, fig. 2, е-ж [e-zh] (nom. nov. pro Unio grayanus sensu Schrenck, 1867, non Lea, 

1834). 

● Type locality: Ussuri River. 

● Lectotype (Moskvicheva, 1973b): ZIN. 

● Distribution: Ussuri River, basin of Khanka Lake (Moskvicheva, 1973b; Zatrawkin, Bogatov, 1987). 


Lanceolaria ussuriensis Moskvicheva, 1973 F 

[= Lanceolaria grayana sensu Shadin, 1952, non Lea, 1834] 

Moskvicheva, 1973b: 1465-1466, fig. 2, з, è [z, i] (nom. nov. pro Lanceolaria grayana sensu Shadin, 1952, 

non Lea, 1834). 

● Type locality: Khanka Lake near village Troizkoe. 


● Distribution: Khanka Lake, Ussuri river (Moskvicheva, 1973b; Zatrawkin, Bogatov, 1987). 


Tumidiana Servain, 1882 

Type species: Unio tumidus Philipsson in Retzius, 1788 (fixation of the type species not traced) 

Tumidiana conus conus (Spengler, 1793) F 

[= Unio ovalis Montagu, 1845] 

Unio conus Spengler, 1793: 60-61. 

● Type locality: “fra Tranquebar”. 


● Distribution: rivers and lakes of Europe, except north and north-east, Ural river (Pirogov, Tarasov, Kazantseva, 


● Remark: Graf (2007) considered this species as a synonym of Unio tumidus Philipsson in Retzius, 1788. 

Tumidiana conus borysthenensis (Kobelt, 1880) F 

Unio tumidus var. borysthenensis Kobelt, 1880: 32, pl. 192, fig. 1950. 

● Type locality: "im Dniepr bei Ekatherinoslaw". 


● Distribution: Ukraine, Belorussia (Stadnichenko, 1984). 


Tumidiana muelleri (Rossmässler, 1838) F 

Unio muelleri Rosmässler, 1838: 41, pl. 40, fig. 541. 

● Type locality: "im Schulensee bei Kiel". 


● Distribution: rivers of Europe, except north and north-east, Irtysh River basin (within Chelyabinsk district) 

(Stadnichenko, 1984; Starobogatov et al., 2004; ; Andreeva et al., 2009). 


Tumidiana tumida tumida (Philipsson in Retzius, 1788) F 

Unio tumidus Philippson – Retzius, 1788: 17. 

● Type locality: “in fluviis Europaea”. 


● Distribution: rivers and lakes of Europe, except north and north-east, Ural River (Pirogov, Tarasov, Kazantseva, 

1994; Kornuishin et al., 2002; Starobogatov et al., 2004). 

Tumidiana tumida falcatulus (Drouët, 1881) F 

Unio tumidus var. falcatulus Drouët, 1881: 9. 

● Type locality: "la rivière de Kharkov".


● Distribution: Ukraine (Stadnichenko, 1984). 


Unio Philipsson, 1788 

Type species: Mya pictorum Linnaeus, 1758 (SD ICZN Opinion 495, 1957) 

Unio limosus limosus Nilsson, 1822 F 

Nilsson, 1822: 110-111. 

● Type locality: “in Scania vulgaris”. 


● Distribution: rivers of western and southern Europe (basins of Baltic, Northern and Black seas), lower 

Volga (Starobogatov et al., 2004). 

● Remark: Graf (2007) considered this species as a synonym of Unio pictorum (Linnaeus, 1758). 

Unio limosus behningi Starobogatov et Pirogov, 1970 [1971] F 

Starobogatov et Pirogov, 1970 [1971]: 229-230, fig. 2. 

● Type locality: Kazan near Kirovskij settlement [the Volga delta]. 




Unio limosus graniger Ziegler in Schmidt, 1840 F 

Unio limosus graniger Ziegler – Schmidt, 1840: 438-439. 

● Type locality: "Aus dem Loschza-Bache, der den Moorgrund durchsschneidet, unweit Laibach". 


● Distribution: Europe, Ukraine, Ural and Don rivers basin, Caucasus (Stadnichenko, 1984). 


Unio pictorum pictorum (Linnaeus, 1758) F 

Mya pictorum Linnaeus, 1758: 671. 


● Probable syntypes: UUZM Nos. 808, 809. 

● Distribution: lakes and rivers of Europe (except northern and north-eastern), western Transcaucasia, Ural 

river (Pirogov, Tarasov, Kazantseva, 1994; Starobogatov et al., 2004). 

Unio pictorum ponderosus Spitzi in Rossmässler, 1844 F 

"Unio pictorum var. Syn. U. ponderosus Spitzi"-- Rossmässler, 1844: 31, pl. 59, fig. 767. 

● Type locality: "in einem Mühlbache bei St. Leonhard in Steyermark". 


● Distribution: central Europe, Danube basin, Ukraine (Stadnichenko, 1984). 

“Unio protractus Lindholm, 1932” F 

Lindholm, 1932: 15-16, pl. 5, fig. 1, 2, 4-7. 

● Type locality: ???? 


● Distribution: basin of Ural river and south-east of European part of Russia (Pirogov, Tarasov, Kazantseva, 


● Remark: this species is usually referred to 1932 or 1922 (the latter, eg. Graf, 2007). We were not able to 

find its description in the publications of 1922. In the publication of 1932 Lindholm cited his owm manuscript 

dated 1930. The publication of 1932 did not contaqin formal description and diagnosis. Therefore the 

name is not available according to Article 13.1 of ICZN. Nevertheless in the reference part Linholm cited: 

«Линдгольм В.А. Моллюски из среднеплиоценовых отложений юго-западной Сибири (Mol-

lusken aus mittelpliozäen Ablagerungen Südwest-Sibiriens). Рукопись, принятая к печати в марте 

1930 г. и содержащая первоописания следующих упомянутых в статье новых видов: ... U. protractus 

....». [“Lindholm V.A. Molluscs of the Middle Pliocene deposits of south-western Siberia (Mollusken 

aus mittelpliozäen Ablagerungen Südwest-Sibiriens). The manuscript accepted in March 

1930 and contains descriptions of the following mentioned in the paper new species: ... U. protractus 

....”] We were not able to find any references to the corresponding publication and we are not 

aware whether the descriptipon of Unio protractus was formally published. It is possivle that the 

name is not yet validated. 

Graf (2007) considered this species as a synonym of Unio pictorum (Linnaeus, 1758). 

Unio rostratus rostratus Lamarck, 1819 F 

[= Unio longirostris Rossmässler, 1836; Unio schrenkianus Clessin, 1880] 

Lamarck, 1819: 77. 

● Type locality: "dans le Rhфne et les grandes rivières de l'Allemagne, de la Silésie, etc." 


● Distribution: slow rivers and lakes of Europe, except north-east, Irtysh River basin (within Chelyabinsk 

district) (Starobogatov et al., 2004; Andreeva et al., 2009). 


Unio rostratus gentilis Haas in Hesse, 1911 F 

Unio gentilis Haas – Hesse, 1911: 151-152. 

● Type locality: "Maritza bei Philippopel". 



ANODONTINAE Rafinesque, 1820 

Amuranodonta Moskvicheva, 1973 

Type species: Amuranodonta kijaensis Moskvicheva, 1973 (OD) 

Amuranodonta boloniensis Zatrawkin et Bogatov, 1987 F 

Buldowskia boloniensis Zatrawkin, Bogatov, 1987: 86-87, fig. 18 з-к [z-k]. 

● Type locality: Bolon Lake basin of the lower Amur River. 


● Distribution: type locality (Starobogatov et al., 2004), Arejskoye Lake, Transvbaikalia (Klishko, 2009). 

● Remark: Graf (2007) considered this species as a synonym of Anodonta arcaeformis (Heude, 1877). 

Amuranodonta kijaensis Moskvicheva, 1973 F 

Amuranodonta kijaensis Moskvicheva, 1973a: 831-832, fig. 2 (5). 

● Type locality: Zarechnoye Lake, basin of the River Kiya (Khabarovsk Region). 


● Distribution: basin of the Ussuri and lower Amur rivers (Starobogatov et al., 2004). 


Klishko (2009) recorded this species in Arejskoye Lake, Transvbaikalia, and failed to distinguish it from A. 

sitaensis Bogatov, Starobogatov, 1996 от the basis of conchological and morphological characters. 

Amuranodonta lomakini (Zatrawkin et Bogatov, 1987) F 

[= Amuranodonta inflata Bogatov et Starobogatov, 1996] 

Buldowskia lomakini Zatrawkin, Bogatov, 1987: 86, fig. 18 д-л [d-l]. 

● Type locality: Amur River near village Susanino. 

● Holotype: ZIN.

● Distribution: basin of the Ussuri and lower Amur rivers (Zatrawkin, Bogatov, 1987; Starobogatov et al., 

2004), Arejskoye Lake, Transvbaikalia (Klishko, 2009).. 


Amuranodonta parva (Moskvicheva, 1973) F 

[= Anemina (Buldowskia) lacustris Labay et Shulga, 1999] 

Amuranodonta parva Moskvicheva, 1973a: 832, fig. 2 (6). 

● Type locality: lower Amur River. 


● Distribution: basin of the lower Amur River and Sladkoe Lake (northwestern Sakhalin) (Zatrawkin, Bogatov, 




pulchra Bogatov et Starobogatov, 1996 оn the basis of conchological and morphological characters. 

Amuranodonta pulchra Bogatov et Starobogatov, 1996 F 

Bogatov, Starobogatov, 1996a: 974-975, textfig. (3, 4, 6b). 

● Type locality: lake in old river bed, 0.5 km upstream of Vostretsovo settlement, right bank of Bolshaya Ussurka 

River. 


● Distribution: Bolshaya Ussurka River basin (Primorje) (Starobogatov et al., 2004). 



parva Moskvicheva, 1973 оn the basis of conchological and morphological characters. 

Amuranodonta sitaensis Bogatov et Starobogatov, 1996 F 

[= Anemina (Buldowskia) adotymensis Labay et Shulga, 1999] 

Buldowskia (Amurbuldowskia) sitaensis Bogatov, Starobogatov, 1996a: 973-974, textfig. (1, 2). 

● Type locality: inlet of Sita River near bridge of motor road Khabarovsk – Vladivostok. 


● Distribution: Ussuri basin, lower Amur basin, Krivushka Lake (northern Sakhalin) (Bogatov, 2001; Starobogatov 

et al., 2004). 



kijaensis Moskvicheva, 1973 от the basis of conchological and morphological characters. 

Anemina Haas, 1969 

Type species: Anodonta arcaeformis Heude, 1877 (OD) 

Anemina buldowskii (Moskvicheva, 1973) F 

Sinanodonta (Anemina) buldowskii Moskvicheva, 1973a: 829-830, fig. 2 (3). 

● Type locality: Ulakhe (Ussuri) River. 


● Distribution: Ussuri River, basin of the Khanka (Zatrawkin, Bogatov, 1987). 


Anemina fuscoviridis (Moskvicheva, 1973) F 

[= Anodonta euscaphus sensu Shadin, 1938, non Anodon euscaphus Heude, 1879; Anemina shadini deflexa 

Martynov et Tshernyshev, 1992] 

Sinanodonta fuscoviridis Moskvicheva, 1973a: 830 (nom. nov. pro Anodonta euscaphus sensu Shadin, 1938, 

non Anodon euscaphus Heude, 1879). 

● Type locality: Ulakhe (Ussuri) River. 

● Lectotype (Moskvicheva, 1973a): ZIN.

● Distribution: Ussuri River, Trosnikovoye Lake, introduced in vicinities of Vladivostok (Zatrawkin, Bogatov, 


● Remark: Graf (2007) considered this species as a synonym of Anodonta euscaphys (Heude, 1879). 

Anemina shadini (Moskvicheva, 1973) F 

[= Anodonta arcaeformis sensu Shadin, 1938, non Anodon arcaeformis Heude, 1877] 

Sinanodonta (Anemina) shadini Moskvicheva, 1973a: 828-829 (nom. nov. pro Anodonta arcaeformis sensu 

Shadin, 1938, non Anodon arcaeformis Heude, 1877). 

● Type locality: Mandzhurka (Troizkaya) River, basin of the Khanka Lake. 

● Lectotype (Moskvicheva, 1973a): ZIN. 

● Distribution: Shilka, Keryleya, Zeya, Amur (lower), and Troizkaya rivers (Zatrawkin, Bogatov, 1987). 


Anodonta Lamarck, 1799 

Type species: Mytilus cygneus Linnaeus, 1758 (monotypy) 

Anodonta cygnea (Linnaeus, 1758) F 

Mytilus cygneus Linnaeus, 1758: 706. 



● Distribution: Europe, except northern part, Ural river (Stadnichenko, 1984; Pirogov, Tarasov, Kazantseva, 


Anodonta stagnalis (Gmelin, 1791) F 

Mytilus stagnalis Gmelin, 1791: 3362. 

● Type locality: "in Schwansee" (Germany?). 


● Distribution: lakes and ponds of Europe, Ural river (Stadnichenko, 1984; Pirogov, Tarasov, Kazantseva, 


● Remark: Graf (2007) considered this species as a synonym of Anodonta cygnea (Linnaeus, 1758). 

Anodonta zellensis zellensis (Gmelin, 1791) F 

[= Anodonta cellensis auctt., nom. err. pro zellensis] 

Mytilus zellensis Gmelin, 1791: 3362. 

● Type locality: "moenia urbis Cellensis circumdante" (Germany?). 


● Distribution: lakes of Europe except north and north-east, Ural river (Pirogov, Tarasov, Kazantseva, 1994; 


● Remark: Graf (2007) considered this species as a synonym of Anodonta cygnea (Linnaeus, 1758). 

Anodonta zellensis micheli Modell, 1945 F 

Modell, 1945: 81, 97-99, textfig. 26. 

● Type locality: "Dnjepr bei Kijew". 


● Distribution: Ukraine, except Crimea and Karpatian mountains (Stadnichenko, 1984). 

Beringiana Starobogatov in Zatrawkin, 1983 

Type species: Anodonta beringiana Middendorff, 1851 (OD) 

Beringiana beringiana (Middendorff, 1851) F 

Anodonta cellensis var. beringiana Middendorff, 1851: 284-287, pl. 29, figs. 1-3. 

● Type locality: Aleutian Islands, Unalashka, Kenai Lake. 

● Holotype: ZIN.

● Distribution: Kamchatka, Chukchi Peninsula, Kunashir аnd Paramushir islands (Kurile Islands) (Zatrawkin, 

Bogatov, 1987; Starobogatov et al., 2004). 

Beringiana chereshnevi Bogatov et Starobogatov, 2001 F 

Bogatov, Starobogatov, 2001: 26-27, figs. 2 a, 2 б (b), 4 ж (zh). 

● Type locality: Elergytgyn Lake, Khatanga River basin. 


● Distribution: Chukchi Peninsula, Kamchatka (Starobogatov et al., 2004). 

● Remark: Graf (2007) considered this species as a synonym of Anodonta beringiana Middendorff, 1851. 

Beringiana compressa Sayenko et Bogatov, 1998 F 

Sayenko, Bogatov, 1998: 1414-1417, fig. 2, 3 б (b), 4. 

● Type locality: Pernatoe Lake, Paramushir Island, 50º03.1' N, 155º22.87' E (North Kurile Islands). 




Beringiana derzhavini Bogatov et Starobogatov, 2001 F 

Bogatov, Starobogatov, 2001: 27, figs. 2 в (v), 2 г (g), 4a. 

● Type locality: Blizhnyaya River, source of Blizhneje Lake, vicinities of Petropavlovsk-Kamchatsky. 


● Distribution: Kamchatka (Starobogatov et al., 2004). 


Beringiana kamchatica Bogatov et Starobogatov, 2001 F 

Bogatov, Starobogatov, 2001: 27-28, figs. 2ä (d), 2 е (e), 4 г (g). 

● Type locality: Blizhnyaya River, source of Blizhneje Lake, vicinities of Petropavlovsk-Kamchatsky. 


● Distribution: Kamchatka, Paramushir Island (North Kurile Islands), Magadan Region (Starobogatov et al., 

2004). 


Beringiana youconensis (Lea, 1867) F 

[= Anodonta youkanensis Lea, 1868; Anodonta beringiana sensu Shadin, 1938, non Middendorff, 1851] 

Anodonta youconensis Lea, 1867: 81. 

● Type locality: Upper Youkan River (Alaska). 

● Lectotype (Johnson, 1974): USNM No. 86624. 

● Distribution: Yakutia, Magadan Region, Kamchatka, Chukchi Peninsula, basin of Anadyr’ River 

(Zatrawkin, Bogatov, 1987; Starobogatov et al., 2004). 


Buldowskia Moskvicheva, 1973 

Type species: Anodonta arcaeformis sujfunica Lindholm, 1925 (OD) 

Buldowskia cylindrica Moskvicheva, 1973 F 

Moskvicheva, 1973b: 1467-1468, fig. 3, в, г [v, g]. 

● Type locality: Batalyanza River downstream of El-Pauza Lake. 


● Distribution: lakes in the vicinities of Artem (Primorje) (Starobogatov et al., 2004 


Buldowskia flavotincta (Martens, 1905) F 

Anodonta arcaeformis var. flavotincta Martens, 1905: 64, pl. 2, fig. 4.

● Type locality: "Söul, Provinz Kyöngkwido, Keumgang bei Konju, Provinz Chhungchhöngdo". 


● Distribution: Lug Lake in the basin of Komarovka River, Razdolnaya River, Krolevezkoye Lake and ajacent 

portions of the river Kievhanka (Zatrawkin, Bogatov, 1987). 


Buldowskia koreana Bogatov et Starobogatov, 1996 F 

Bogatov, Starobogatov, 1996b: 1331, fig. 2 в, г (v, g). 

● Type locality: lake of former bed of Gladkaya River in 2 km of Gvozdevo settlement, Khasan District, 

Primorje Territory. 


● Distribution: southernmost part of Primorje, lakes and former river beds (Starobogatov et al., 2004). 


Buldowskia possietica Bogatov et Starobogatov, 1996 F 

Bogatov, Starobogatov, 1996b: 1329-1330, fig. 3 а, б (a, b). 

● Type locality: lake on Boisman Bay shore, Ryazanovka River basin, Khasan District, Primorje Territory. 


● Distribution: southernmost part of Primorje, lakes (Starobogatov et al., 2004). 


Buldowskia starobogatovi (Moskvicheva, 1973) F 

Amuranodonta starobogatovi Moskvicheva, 1973a: 831, fig. 2 (4). 

● Type locality: small lake on the Krasnoyarskaya hill, vicinities of city Ussuriisk (formerly Nikolsk). 


● Distribution: type locality, Khasan Lake, Lug Lake, lakes of the basin of Razdolnaya River (Zatrawkin, 

Bogatov, 1987). 


Buldowskia sujfunensis (Shadin, 1938) F 

[= Anemina (Buldowskia) zatrawkini Martynov et Tshernyshev, 1992] 

Anodonta beringiana var. sujfunensis Shadin, 1938: 132, fig. 48. 

● Type locality: Bokhiiskoe lake, basin of Razdolnaya (Sujfun) River. 


● Distribution: lakes of the basin of Razdolnaya River, Khasan Lake, lakes on Pyutatin Island, Barabashevka 

River (Zatrawkin, Bogatov, 1987). 


Buldowskia sujfunica (Lindholm, 1925) F 

Anodonta arcaeformis var. sujfunica Lindholm, 1925b: 137. 

● Type locality: Razdolnaya (Sujfun) River, near city of Ussuriisk (formerly Nikolsk). 


● Distribution: Razdolnaya, Komarovka, and Kievchanka rivers (Zatrawkin, Bogatov, 1987). 


Buldowskia suputinensis Moskvicheva, 1973 F 

Moskvicheva, 1973a: 833, fig. 2 (8). 

● Type locality: Lug Lake, basin of Komarovka River, vicinities of city Ussuriisk. 


● Distribution: type locality, Kroveletskoye Lake (basin of thew river Kievchanka, tributary of the river Artemovka), 

Kievchanka River (Zatrawkin, Bogatov, 1987). 

● Remark: Graf (2007) considered this species as a synonym of Anodonta arcaeformis (Heude, 1877).

Colletopterum Bourguignat, 1880 

Type species: Anodonta letourneuxi Bourguignat, 1880 (= Anodonta subcircularis Clessin, 1873) (SD 

Zatrawkin, 1983) 

Colletopterum anatinum (Linnaeus, 1758) F 

[= Anodonta minima Millet, 1833; Anodonta sedakovi Siemaschko, 1848; Anodonta seisanensis Kobelt, 

1911; Anodonta piscinalis var. sorica W. Dybowski in B. Dybowski, 1913] 

Mytilus anatinus Linnaeus, 1758: 706. 


● Probable syntype: UUZM No. 825. 

● Distribution: rivers of Europe, except extreme north-east, Ural river, Siberia, eastward to Baikal (Pirogov, 

Tarasov, Kinazantseva, 1994; Starobogatov et al., 2004; Bogatov et al., 2005). 

Colletopterum apollonicum (Bourguignat, 1880-1881) F 

[=Anodonta piscinalis var. volgensis Zhadin, 1938, part.] 

Anodonta apollonica Bourguignat, 1880-1881: 91-95, pl. 15, figs. 1-3. 

● Type locality: Apollonia Lake, Anatolia, Turkey (Westerlund, 1890). 


● Distribution: South Europe and Asia Minor (Starobogatov et al., 2004; Bogatov et al., 2005 ). 

● Remark: Graf (2007) considered this species as a synonym of Anodonta anatina (Linnaeus, 1758). 

Colletopterum bactrianum (Rolle, 1897) F 

Anodonta bactriana Rolle, 1897: 172. 

● Type locality: “Serafschan bei Buchara”. 


● Distribution: Central Asia – Zeravshan and Syrdarya basins (Starobogatov, Izzatullaev, 1984; Izzatullaev, 

2002). 

● Remark: Bogatov et al. (2005) did not mention this species in their revision of the genus. Graf (2007) considered 

this species as a synonym of Anodonta cyrea Drouët, 1881. 

Colletopterum baeri Bogatov, Starobogatov and Prozorova, 2005 F 

Bogatov et al., 2005: 1055-1056, figs. 3л, 3м, 4д [3l, 3m, 4d]. 

● Type locality: Volga River near Astrahan’. 


● Distribution: lower part of Volga River (Bogatov et al., 2005). 

Colletopterum convexa (Drouët, 1888) F 

[= Anodonta piscinalis var. sorica W. Dybowski in B. Dybowski, 1913, part.; Anodonta piscinalis var. volgensis 

Zhadin, 1938, part. 

Anodonta convexa Drouët, 1888: 110 

● Type locality: Saône near Charrey, France (fide Westerlund, 1890). 


● Distribution: southern Europe, finding of the species in Baikal is doubtful and the specimen of W. Dybowski 

from Bolshoi Sor of Baikal might have a wrong label (Bogatov et al., 2005). 

Colletopterum cyreum cyreum (Drouët, 1881) F 

Anodonta cyrea Drouët, 1881: 29. 

● Type locality: "le Kour ...; les cours d'eau du Caucase et de la mer Caspienne" (Kura River). 

● Syntype: ZIN ???? 

● Distribution: Caucasus, Transcaucasia (Shadin, 1952; Bogatov et al., 2005). 

Colletopterum cyreum sogdianum (Kobelt, 1896) F 

[= Anodonta retteri Drouët, 1899; A. samarkandensis Kobelt, 1913]

Anodonta sogdiana Kobelt, 1896: 102-103. 

● Type locality: “Serafschan bei Buchara”. 


● Distribution: Central Asia (Starobogatov, Izzatullaev, 1984; Izzatullaev, 2002) 

● Remark: Bogatov et al. (2005) did not mention this species in their revision of the genus. 

Colletopterum depressum (Bourguignat, 1880-1881) F 

Anodonta depressa Bourguignat, 1880-1881: 221-222. 

● Type locality: "dans les lacs de la Carniole et de la Carinthe, notamment dans ceux des environs de Klagenfurt". 


● Distribution: southern Europe, basin of the Black Sea, possibly in the lower Don and Volga (Starobogatov 

et al., 2004). 


this species as a synonym of Anodonta anatina (Linnaeus, 1758). 

Colletopterum kokandicum Starobogatov et Izzatullaev, 1984 F 

Starobogatov, Izzatullaev, 1984: 78, fig. 4, 

● Type locality: Pechkoran Lake, northward from Kokand city [Uzbek SSR]. 


● Distribution: Central Asia, type locality (Starobogatov, Izzatullaev, 1984; Izzatullaev, 2002). 


this species as a synonym of Anodonta cyrea Drouët, 1881. 

Colletopterum milaschevichi Bogatov, Starobogatov and Prozorova, 2005 F 

[=Anodonta ponderosa var. volgensis .Zhadin, 1938, part.] 

Bogatov et al., 2005: 1056, figs. 3н-3р, 4в [3n-3r, 4v]. 

● Type locality: Volga Delta, Kabanyi Il’men’. 

● Holotype: ZIN (it is one of the paralectotypes of Anodonta ponderosa var. volgensis .Zhadin, 1938. 

● Distribution: lower part of Volga River (Bogatov et al., 2005). 

Colletopterum nilssonii (Küster, 1842) F 

[= Anodonta piscinalis var. sorica W. Dybowski in B. Dybowski, 1913, part.] 

Anodonta nilssonii Küster, 1842: 61. 

● Type locality: “in der Regnitz bei Erlangen” (Germany). 


● Distribution: Europe and Siberia, eastward to Baikal (Starobogatov et al., 2004; Bogatov et al., 2005). 


Colletopterum oblongum (Millet, 1831) F 

Anodonta oblonga Millet, 1831: 242, pl. 12, fig. 1. 

● Type locality: "la Mayenne". 


● Distribution: basins of Baltic and Northern seas (Starobogatov et al., 2004). 

● Remark: Bogatov et al. (2005) did not mention this species in their revision of the genus. 

Colletopterum ostiarium (Drouët, 1881) F 

[=Anodonta piscinalis var. volgensis Zhadin, 1938, part.; Anodonta (Colletopterum) piscinalis shadini Starobogatov, 

Pirogov, 1970[1971]] 

Anodonta ostiaria Drouët, 1881: 26. 

● Type locality: “le liman du Dniéper”. 


● Distribution: southern part of Central and eastern Europe (Bogatov et al., 2005).

● Remark: Stadnichenko (1984) recorded this species in Black Sea Maritinme territory under the name Colletopterum 

subcirculare plattenicum (Servain, 1881). 

Colletopterum piscinale piscinale (Nilsson, 1822) F 

[=Anodonta cellensis var. selengensis W. Dybowski in B. Dybowski, 1913; Anodonta nova B. Dybowski, 

1913; Anodonta piscinalis sibirica Krovosheina et Starobogatov, 1970] 

Anodonta piscinalis Nilsson, 1822: 116-117. 

● Type locality: “in piscinis & fossis Scaniae passim”. 


● Distribution: Europe, except extreme north-east, southern part of western Sibiria, Pribaikal’e (eastward to 

Baikal), northern Kazakhstan; Armenia (Akramowski, 1976; Pirogov, Tarasov, Kazantseva, 1994; Starobogatov 

et al., 2004; Bogatov et al., 2005). 


Colletopterum piscinale falcatum (Drouët, 1879) F 

Anodonta falcata Drouët, 1879: 332-333. 

● Type locality: "le Dniéper, а Wordkodniéprowsk". 


● Distribution: Don river basin waterbodies (Stadnichenko, 1984). 

● Remark: Bogatov et al. (2005) did not mention this subspecies in their revision of the genus. 

Colletopterum ponderosum ponderosum (C. Pfeiffer, 1825) F 

[=Anodonta piscinalis var. volgensis Zhadin, 1938, part; Anodonta anatina var. petshorica Shadin, 1938; 

Anodonta anatina var. lenae Shadin, 1938, part.; Anodonta ponderosa altaica Krivosheina et Starobogatov, 

1970] 

Anodonta ponderosa C. Pfeiffer, 1825: 31-32, pl. 4, figs. 1-6. 

● Type locality: "Aus flusse des Radsieker Teiches in den Wörntebach, in der Gegend von Pyrmont". 


● Distribution: Europe and Siberia, including Yakutia and Transbaikalia, Mongolia (Starobogatov, Izzatullaev, 

1984; Starobogatov et al., 2004; Bogatov et al., 2005). 


Colletopterum ponderosum rumanicum (Letourneux in Bourguignat, 1880-1881) F 

Anodonta rumanica Letourneux – Bourguignat, 1880-1881:199-200. 

● Type locality: "dans le Danube aux environs de Giurgewo, en Valachie". 


● Distribution: Black and Azov seas basin (Stadnichenko, 1984). 

● Remark: Bogatov et al. (2005) did not mention this subspecies in their revision of the genus. 

Colletopterum rostratum (Rossmaessler, 1836) F 

[Anodonta anatina var. lenae Shadin, 1938, part.] 

Anodonta rostrata Rossmaessler, 1836b: 25, pl. 20, fig. 284 

● Type locality: "im Wörthsee bei Klagenfurth und in dessen beiden Ausflüssen, dem Lendkanale und dem 

Glanfurtbachem und im Salzsee im Mannsfeldischen”. 


● Distribution: Europe and Siberia, eastward to Lena River (Bogatov et al., 2005). 

Colletopterum sorensianum (W. Dybowski in B. Dybowski, 1913) F 

Anodonta complanata var. sorensiana W. Dybowski in B. Dybowski, 1913: 214, pl. 6, No. 34. 

● Type locality: Bolshoi Sor, Baikal Lake. 

● Holotype: ZIN (the only known specimen). 

● Distribution: Baikal Lake (Bogatov et al., 2005).

Colletopterum subcirculare (Clessin, 1873) F 

[=Anodonta piscinalis var. volgensis Zhadin, 1938, part.; Anodonta (Colletopterum) subcircularis astrachanica 

Starobogatov, Pirogov, 1970 [1971]] 

Anodonta subcirculare Clessin, 1873b: 87-88, pl. 22, fig. 2. 

● Type locality: "Europäische Türkei, im See von Skutari". 


● Distribution: rivers of the western and southern Europe (Starobogatov et al., 2004; Bogatov et al., 2005). 


Cristaria Schumacher, 1817 

Type species: Cristaria tuberculata Schumacher, 1817 (monotypy) 

Cristaria herculea (Middendorff, 1847) F 

Anodonta herculea Middendorff, 1847: 302-303, 1851: 278, pl. 21, fig. 5, pl. 22, figs. 1, 2. 

● Type locality: Onon River, near village Ust’-Alyatui. 

● Lectotype: ZIN (selected by??) 

● Distribution: lakes on Iturup Island (Kuriles), basin of Khanka Lake, Onon, Nercha, Argun, Shilka, Zeya, 

and lower Amur rivers (Zatrawkin, Bogatov, 1987). 

● Remark: Graf (2007) considered this species as a synonym of Cristaria plicata (Leach, 1815). 

Cristaria tuberculata Schumacher, 1817 F 

Schumacher, 1817: 107, pl. 20, fig. 2. 



● Distribution: basin of Khanka Lake, Ussuri River (Zatrawkin, Bogatov, 1987). 

● Remark: Graf (2007) considered this species as a synonym of Cristaria plicata (Leach, 1815). 

Sinanodonta Modell, 1945 

Type species: Symphyncta woodiana Lea, 1834 (OD) 

Sinanodonta amurensis Moskvicheva, 1973 F 

Moskvicheva, 1973a: 826-827, fig. 1 (11). 

● Type locality: vicinities of Khabarovsk, channel Malye Chepchiki. 


● Distribution: channels of the Amur River, Ussuri, Bolshaya Ussurka rivers, Ulil’, Petropavlovskoye, Dzhalynskoye, 

Khanka lakes (Zatrawkin, Bogatov, 1987). 

● Remark: Graf (2007) considered this species as a synonym of Anodonta woodiana woodiana (Lea, 1834). 

Sinanodonta crassitesta Moskvicheva, 1973 F 

[= Sinanodonta renzini Bogatov et Zatrawkin, 1988; Sinanodonta ovata Bogatov et Starobogatov, 1996] 


● Type locality: Suputinka (Komarovka) River. 

● Lectotype: ZIN. 

● Distribution: Razdolnaya, Komarovka rivers, Soldatskoye Lake (Zatrawkin, Bogatov, 1987). 


Sinanodonta fukudai Modell, 1945 F 

[= Anodonta woodiana var. eliptica sensu Shadin, 1938, non Anodon eliptica Heude, 1878] 

Sinanodonta woodiana fukudai Modell, 1945: 96, textfig. 24. 

● Type locality: "Zenra Dф, Fuan Gun, Turuppo, Nordkorea". 

● Holotype: Modell collection, No. 3420. 

● Distribution: basin of the Razdolnaya river (Zatrawkin, Bogatov, 1987). 

● Remark: Graf (2007) considered this species as a synonym of Anodonta woodiana woodiana (Lea, 1834).

Sinanodonta gibba (Benson, 1855) F 

Anodonta gibbum Benson, 1855: 135. 

● Type locality: Tangtse Kaing (China). 

● Syntypes: MCZ No. 177314. 

● Distribution: central Asia, Tashkent Region, ponds [introduced]; Zeravshan River basin (Starobogatov, Izzatullaev, 

1984; Izzatullaev, 2002; Boimurodov, 2004). 


Sinanodonta likharevi Moskvicheva, 1973 F 


● Type locality: Shilka River in the vicinities of the town Sretensk-Zabaikalskii. 


● Distribution: Shilka, Nercha, Zeya, lower Amur River, Ussuri, Bolshaya Ussurka, Ilistaya rivers, Udyl, 

Trostnikovoe (Zatrawkin, Bogatov, 1987). 


Sinanodonta manchurica Bogatov et Starobogatov, 1996 F 

Bogatov, Starobogatov, 1996b: 1329, fig. 2 в, г (v, g). 

● Type locality: lake of former bed of Gladkaya River in 2 km of Gvozdevo settlement, Khasan District, 



● Distribution: southernmost part of Primorje (Starobogatov et al., 2004). 


Sinanodonta orbicularis (Heude, 1880) F 

Anodon orbicularis Heude, 1880b: pl. 42, fig. 82, caption on opposite unnumbered page. 

● Type locality: "Lac de Ma-tang, district de Peng-tse (Kiang-si), au bord du fleuve Bleu". 


● Distribution: Central Asia, Tashkent Region, ponds [introduced]; Zeravshan River basin (Starobogatov, 

Izzatullaev, 1984; Izzatullaev, 2002; Boimurodov, 2004). 


Sinanodonta primorjensis Bogatov et Zatrawkin, 1988 F 

Sinanodonta (Cristariopsis) primorjensis Bogatov, Zatrawkin, 1988: 162-163, fig. 4. 

● Type locality: Soldatskoye Lake in the basin of Razdolnaya River, Primorje. 


● Distribution: lakes in the basin of Razdolnaya [Sujfun] River (Starobogatov et al., 2004). 


Sinanodonta puerorum (Heude, 1880) F 

Anodon puerorum Heude, 1880b: pl. 47, fig. 90, caption on opposite unnumbered page. 

● Type locality: "les fosses de la plaine de Tch'ao-hien, sue les bords du Yang-tze". 

● Syntype: USNM No. 126509. 

● Distribution: central Asia, Tashkent Region, ponds [introduced]; Zeravshan River basin (Starobogatov, Izzatullaev, 

1984; Izzatullaev, 2002; Boimurodov, 2004). 


Sinanodonta schrencki Moskvicheva, 1973 F 

[= Anodonta magnifica sensu Schrenck, 1867, non Symphinota magnifica Lea, 1834; Anodonta woodiana 

sensu Shadin, 1938, non Symphinota woodiana Lea, 1834] 

Moskvicheva, 1973a: 826 (nom. nov. pro Anodonta magnifica sensu Schrenck, 1867, non Symphinota magnifica 

Lea, 1834).

● Type locality: Ussuri River near mouth of the river Khor ("Poor"). 


● Distribution: lower Amur, Ussuri, Arsenievka rivers, basin of Khanka Lake (Zatrawkin, Bogatov, 1987). 


Sinanodonta woodiana (Lea, 1834) F 

Symphinota woodiana Lea, 1834: 42, pl. 5, fig. 13. 

● Type locality: China. 

● Types: lost (Johnson, 1974). 

● Distribution: southwestern Ukraine, introduced (Korniushin et al., 2002). 

NODULARIINAE Starobogatov et Zatrawkin in Zatrawkin et Bogatov, 

1987 

● Remark: Chernyshev (2004) considered establishing the separate subfamily Nodulariinae for two genera – 

Nodularia Conrad, 1857 and Middendorffinaia Moskvicheva et Starobogatov, 1973 – as premature. His opinion 

was based on the absence of the comparison of the diagnosis of the new subfamily with that of Unioninae. 

Nevertheless he himself did not formally synonymized Nodulariinae with any other group of Unionidae 

and until the formal taxonomic decision will be published we prefer to consider this subfamily as valid. 

Middendorffinaia Moskvicheva et Starobogatov, 1973 

Type species: Unio mongolicus Middendorff, 1851 (OD) 

Middendorffinaia arsenievi Moskvicheva et Starobogatov, 1973 F 

Moskvicheva, Starobogatov, 1973: 29, fig. 3 m-o. 

● Type locality: Pashino River, near Dormidontovka village, Khabarovsk Territory [Far East] 


● Distribution: basins of Ussuri and lower Amur rivers (Zatrawkin, Bogatov, 1987). 

● Remark: Graf (2007) considered this species as a synonym of Unio crassus mongolicus Middendorff, 

1851. 

Middendorffinaia dulkeitiana Moskvicheva et Starobogatov, 1973 F 

Moskvicheva, Starobogatov, 1973: 33. fig. 5 г, ä [g, d]. 

● Type locality: tributary of the Suputinka River [Far East]. 


● Distribution: basins of Suputinka and Razdolnaya rivers (Zatrawkin, Bogatov, 1987). 

● Remark: Graf (2007) considered this species as a synonym of Inversidens pantoensis (Neumayr, 

1899). 

Middendorffinaia mongolica (Middendorff, 1851) F 

Unio mongolicus Middendorff, 1851: 272, pl. 42, figs. 7-8. 

● Type locality: mountain spring near village Gorbiza [eastern part of Chita Region]. 

● Holotype: lost (Moskvicheva, Starobogatov, 1973). 

● Distribution: Type locality, upstreams of the rivers Zeya, Arga, Ussuri, Khor, Arsenievka, Sungacha 

(Zatrawkin, Bogatov, 1987). 

Middendorffinaia ochotica Bogatov, 2000 F 

Bogatov, 2000: 861-862, textfig. (а-г) (a-g). 

● Type locality: Kukhtui River northward of Okhotsk, Khabarovsk Region. 


● Distribution: Kukhtui River basin (Okhotsk Sea basin), possibly Amur basin (Starobogatov et al., 2004).


1851. 

Middendorffinaia shadini Moskvicheva et Starobogatov, 1973 F 

[= Unio continentalis Shadin, 1938, non Haas, 1910; Middendorffinaia martensi Moskvicheva et Starobogatov, 

1973] 

Moskvicheva, Starobogatov, 1973: 31-32, fig. 5 a-в [a-v] [nom. nov. pro Unio continentalis Shadin, 1938, 

non Haas, 1910]. 

● Type locality: Rakovka River, tributary of the river Razdolnaya [Far East]. 

● Lectotype (Moskvicheva, Starobogatov, 1973?): ZIN. 

● Distribution: basin of the Razdolnaya River (Zatrawkin, Bogatov, 1987). 


1899). 

Middendorffinaia sujfunensis Moskvicheva et Starobogatov, 1973 F 

Moskvicheva, Starobogatov, 1973: 30, fig. 4. 

● Type locality: Sujfun [Razdolnaya] River, 1.5 km southward from Razdol'noe station, Primorje Territory 

[Far East]. 


● Distribution: Razdolnaya, Artemovka, Kievchanka, and Partizanskaya rivers (Moskvicheva, Starobogatov, 

1973; Zatrawkin, Bogatov, 1987). 


1899). 

Middendorffinaia ussuriensis Moskvicheva et Starobogatov, 1973 F 

Moskvicheva, Starobogatov, 1973: 28-29, fig. 3 è-м [i-m]. 

● Type locality: Daubikhe River, near Jakovlevka village [Far East]. 


● Distribution: rivers Arsenievka [Daubikhe], Pashino, Hor; basin of Ussuri River (Zatrawkin, Bogatov, 



1851. 

Middendorffinaia weliczkowskii Moskvicheva et Starobogatov, 1973 F 

[= Middendorffinaia hassanica Moskvicheva et Starobogatov, 1973; Middendorffinaia maihensis Moskvicheva, 

1973] 

Moskvicheva, Starobogatov, 1973: 33-34, fig. 5 e, ж [e, zh]. 

● Type locality: Olenevka River (tributary of the Sujfun [Razdolnaya] River [Far East]. 


● Distribution: Type locality (Moskvicheva, Starobogatov, 1973; Zatrawkin, Bogatov, 1987). 


1899). 

Nodularia Conrad, 1857 

Type species: Unio douglasiae Griffith et Pidgeon, 1833 (monotypy) 

Nodularia abbreviata (Westerlund, 1897) F 

Unio abbreviatus Westerlund, 1897b: 132. 

● Type locality: Amur River, near Nikolaevsk. 


● Distribution: Amur, Ussuri, Pilda, Ilistaya, Melgunovka rivers, Khanka Lake (Zatrawkin, Bogatov, 1987).

● Remark: Graf (2007) considered this species as a synonym of Nodularia douglasiae (Griffith and Pidgeon, 

1834). 

Nodularia amurensis (Mousson, 1887) F 

[= Unio douglassiae sensu Shadin, 1938, part. non Griffith et Pidgeon, 1833] 

Unio pictorum var. amurensis Mousson, 1887: 26-27, 35, pl.1, fig. 8. 

● Type locality: "Chabarofka..., Nicolajewsk". 

● Syntypes: SMF 3663/2 (fide Zilch, 1967). 

● Distribution: Amur, Ussuri, Milgunovka rivers, Khanka, Udil’, Petropavlovskoye lakes (Zatrawkin, Bogatov, 

1987). 


1834). 

Nodularia extremalis Martynov et Tshernyshev, 1992 F 

Nodularia (Magadaninaia) extremalis Martynov, Tshernyshev, 1992: 19, fig.1-3 

● Type locality: Magadan Region, Olskii district, a river near village Motykleyka. 

● Holotype: 10602/Bv - 1484, Zoological Museum of Far-Eastern State University (No. 254 was erroneously 

mentioned in the original publication). 

● Distribution: type locality, basin of Tayiskaya guba, River Motyleika (Martynov, Tshernyshev, 1992; Starobogatov 

et al., 2004). 


1834). 

Nodularia flavoviridis (Haas, 1911) F 

Nodularia douglassiae var. flavoviridis Haas, 1911: 71, pl. 6, fig. 2. 

● Type locality: "Buir-nur in Amurgebiet" (Buir-Nur Lake, Mongolia). 

● Lectotype (Zilch, 1967): SMF No. No. 3654. 

● Distribution: Amur, Ussuri, Milgunovka, Mandzhurka rivers, Khanka, Petropavlovskoye lakes (Zatrawkin, 

Bogatov, 1987). 


1834). 

Nodularia lebedevi Zatrawkin et Starobogatov, 1984 F 

Zatrawkin, Starobogatov, 1984: 1787-1788, fig. 5-8. 

● Type locality: Sakhalin Island on the Amur River across city of Amursk, inner lake, connected to the Amur 

river, Khabarovsk Territory [Far East]. 


● Distribution: Type locality (Zatrawkin, Bogatov, 1987). 


1834). Chernyshev (2004) cited the personal communication of Starotobogatov, that the specimen originated 

from fresh-water aquarium in Amursk city. The collection data were provided by the owner of the aquarium. 

Chernyshev consider this information as erroneous and stated that the species probably has different origin 

from Amur river and is extremely similar to European species Unio tumidus Philippson, 1788. 

Nodularia middendorffi (Westerlund, 1890) F 

[= Unio pictorum var. longirostris Middendorff, 1851] 

Unio middendorffi Westerlund, 1890: 113. 

● Type locality: Nercha River, near Nerchinsk. 


● Distribution: Amur, Nercha, Ingoda, Shilka, Onon, Zeya, and Ussuri rivers (Zatrawkin, Bogatov, 1987). 


1834).

Nodularia moskvichevae Bogatov et Starobogatov, 1992 F 

Bogatov, Starobogatov, 1992: 134, fig. 1 (5, 6). 

● Type locality: inlet of Razdolnaya River in 1 km from railroad station Razdolnoe, Nadezhdinsky District, 



● Distribution: basin of Razdolnaya [Sujfun] River (Starobogatov et al., 2004). 


1834). 

Nodularia sakhalinensis Bogatov, 2001 F 

Nodularia sakhalinensis Bogatov, 2001: 75-77, fig. 2 A-C. 

● Type locality: Langry River, northern Sakhalin. 


● Distribution: type locality (Bogatov, 2001). 


1834). 

Nodularia schrencki (Westerlund, 1897) F 

Unio schrencki Westerlund, 1897b: 131-132. 

● Type locality: Amur River. 


● Distribution: Amur, Ussuri rivers (Zatrawkin, Bogatov, 1987). 


1834). 

Nodularia vladivostokensis Moskvicheva, 1973 F 

[= Nodularia sujfunica Moskvicheva, 1973] 

Moskvicheva, 1973b: 1463, fig. 1 п, р [p, r]. 

● Type locality: Sujfun (Razdolnaya) River, near station Baranovskyi. 


● Distribution: Razdolnaya, and Komarovka rivers (Zatrawkin, Bogatov, 1987). 


1834). 

PSEUDANODONTINAE Jaeckel, 1962 

Kunashiria Starobogatov in Zatrawkin, 1983 

Type species: Anodonta japonica Clessin, 1874 (OD) 

Kunashiria compressa (Bogatov et Starobogatov, 1996) F 

Arsenievinaia compressa Bogatov, Starobogatov, 1996b: 1332, fig. 4 в, г (v, g). 

● Type locality: small lake Zerkalnoe, Rudnaya River basin, Dalnegorsk District, Primorje Territory. 


● Distribution: Rudnaya River basin (Primorje) (Starobogatov et al., 2004). 

● Remark: Graf (2007) considered this species as a synonym of Anodonta woodiana japonica (von Martens 

in Clessin 1874). 

Kunashiria coptzevi Zatrawkin et Bogatov, 1987 F 

[= Arsenievinaia alimovi Bogatov et Zatrawkin, 1988] 

Arsenievinaia coptzevi Zatrawkin, Bogatov, 1987: 104-105, fig. 20, б [b], 24 в, г [v, g]. 

● Type locality: Vas’kovskoye Lake near vellage Rudnaya Pristan’, Dalnegorsky area, Primorje Region.


● Distribution: eastern slope of Sihkhote-Alin and northern Sakhalin [Primorje] (Zatrawkin, Bogatov, 1987; 




Kunashiria haconensis (Ihering, 1893) F 

[= Amuranodonta sihotealinica Zatrawkin et Starobogatov, 1984; Kunashiria japonica boreosakhalinensis 

Labay et Shulga, 1999] 

Anodonta haconensis Ihering, 1893: 153. 

● Type locality: Hakone Lake (Japan). 


● Distribution: southern Sakhalin, Kurile Islands (Zatrawkin, Bogatov, 1987). 



Kunashiria iturupica Bogatov, Sayenko et Starobogatov, 1999 F 

Bogatov et al., 1999: 60-62, fig. 4. 

● Type locality: Kuibyshevskoye Lake, Iturup Island, Kurile Islands. 

● Holotype: ZIN?. 

● Distribution: Kunashir and Iturup Islands, Kurile Islands (Bogatov et al., 1999). 



Kunashiria iwakawai (Suzuki, 1939) F 

Anodonta iwakawai Suzuki, 1939: 130-138 (in Japanese), 139-143 (in English), pl. 5, 6, textfigs. 1-7. 

● Type locality: Tarukawa-numa, n-w of Sapporo. 

● Holotype: Geological Institute, University of Tokyo, LF1. 

● Distribution: Lesser Kurile Islands (Zelenyi, Tanfilyeva, Yurii islands), South Sakhalin (Starobogatov et 

al., 2004) 



Kunashiria japonica (Martens in Clessin, 1874) F 

Anodonta japonica Martens – Clessin, 1874a: 144-145, pl. 47, figs. 3-4. 

● Type locality: "Yokohama in Japan". 


● Distribution: lakes of southern Sakhalin and southern Kurile Islands (Zatrawkin, Bogatov, 1987; Starobogatov 

et al., 2004). 

Kunashiria sinanodontoides Bogatov, Sayenko et Starobogatov, 1999 F 

Bogatov et al., 1999: 59-60, fig. 1, 3. 

● Type locality: Dobroye Lake, Dobroye Nachalo Bay Basin, Iturup Island, Kurile Islands. 

● Holotype: ZIN?. 




Kunashiria taranetsi (Shadin, 1938) F 

[= Arsenievinaia zatravkini Bogatov et Starobogatov, 1996] 

Anodonta beringiana var. taranetsi Shadin, 1938: 133, fig. 49. 

● Type locality: Sakhalin, lake on the flood plain of the river Tym’ near village Kotik. 

● Lectotype: ZIN (selected by ???.

● Distribution: east slope of Sikhote-Alin and Sakhalin (Starobogatov et al., 2004). 



Kunashiria zarjaensis (Bogatov et Zatrawkin, 1988) F 

Arsenievinaia zarjaensis Bogatov, Zatrawkin, 1988: 166-167, fig. 6. 

● Type locality: Zarya Lake, Primorje. 





Kunashiria zimini (Zatrawkin et Bogatov, 1987) F 

Arsenievinaia zimini Zatrawkin, Bogatov, 1987: 102-104, fig. 24, а, б [a, b] 

● Type locality: Zarya Lake [coast of the Japan Sea] 30 km SW from the village Valentin. 


● Distribution: eastern slope of Sihkhote-Alin’ [Primorje] (Zatrawkin, Bogatov, 1987; Starobogatov et al., 

2004). 



Pseudanodonta Bourguignat, 1877 

Type species: Anodonta complanata Ziegler in Rossmässler, 1835 (SD Westerlund, 1902) 

Pseudanodonta complanata (Ziegler in Rossmässler, 1835) F 

[= Pseudanodonta rossmaessleri sensu Shadin, 1952, non Bourguignat, 1870; Pseudanodonta kletti sensu 

Stadnichenko, 1984, non Rossmässler, 1835; Pseudanodonta elongata tanousi sensu Stadnichenko, 1984, 

non Bourguignat, 1880] 

Anodonta complanata Ziegler – Rossmässler, 1835a: 112, pl. 3, fig. 68. 

● Type locality: “Elbe”. 


● Distribution: delta of Volga River, rivers and lakes of the basin of the Baltic Sea, Ukraine, ?Ural river (Pirogov, 

Tarasov, Kazantseva, 1994; Korniushin et al., 2002; Starobogatov et al., 2004). 

Pseudanodonta elongata (Holandre, 1836) F 

Anodonta elongata Holandre, 1836: 54. 

● Type locality: "sur les bords de la Moselle en Chambières, au-dessous du Poutiffroy". 


● Distribution: basin of the Baltic Sea and in delta of Volga River (Starobogatov et al., 2004). 

● Remark: Graf (2007) considered this species as a synonym of Pseudanodonta complanata (Ziegler in 

Rossmässler, 1835). 

Pseudanodonta kletti (Rossmässler, 1835) F 

Anodonta kletti Rossmässler, 1835a: 112. 

● Type locality: “Oder-Gebiet: Ohlau Fluss bei Breslau”. 


● Distribution: northern Europe, except its extreme north-east, rivers and lakes (Starobogatov et al., 2004). 



Pseudanodonta nordenskioldi Bourguignat, 1880-1881 F 

Bourguignat, 1880-1881: 41 (nom. nov. pro Anodonta complanata sensu Nordenskiold et Nylander, 1856, 

non Ziegler in Rossmässler, 1835).

● Type locality: "Frugеrd i Mäntsälä" (Finland). 


● Distribution: rivers and lakes of the basin of the Baltic Sea (Starobogatov et al., 2004). 



Subclass HETERODONTA 

Order VENEROIDEA 

CARDIOIDEA 

CARDIIDAE Lamarck, 1809 


● Remark: treated here genera are characterized by the presence of both marine (especially from the Caspian 

Sea) and brackishwater species. In order not to split them we deal with entire genera. 

Adacna Eichwald, 1838 

Type species: Glycimeris laeviuscula Eichwald, 1829 

Adacna acuticosta (Logvinenko et Starobogatov, 1967) М 

Hypanis acuticosta Logvinenko, Starobogatov, 1967: 232. 

● Type locality: central part of the slope of the northern Caspian Sea, 22 m. 


● Distribution: Middle Caspian Sea; 0-50 m (Logvinenko, Starobogatov, 1968; Starobogatov et al., 2004). 

Adacna angusticostata (Borcea, 1926) B 

[= Monodacna colorata var. razelmiana Borcea, 1926 (part.)] 

Monodacna colorata var. angusticostata Borcea, 1926: ??? 

● Type locality: ???. 


● Distribution: limans, estuaries and near-coast lakes of northern Black Sea maritime area (Scarlato, Starobogatov, 

1972). 

Adacna colorata (Eichwald, 1829) B 

[= Monodacna colorata var. tanaisiana Milachevitch, 1916; Monodacna colorata var. razelmiana Borcea, 

1926 (part.)] 

Glycimeris colorata Eichwald, 1829: 279. 

● Type locality: “Hab. in ostio Hypanis prope Nicolajeff, in Tanai et Miotide, ubique vivam abservani; In 

Volga anme adesse quoque videtun”. 


● Distribution: waterbodies of the Danube delta, Dniester and Dnieper-Bug limans (northern Black Sea and 

Azov Sea maritime area), Don and Volga deltas, Kuban mouth, water reservoirs of lower Don and Volga, 

northern Caspian Sea, intoduced to Balkhash Lake ((Logvinenko, Starobogatov, 1968; Scarlato, Starobogatov, 


Adacna fragilis Milaschewitsch, 1908 B 

Milaschewitsch, 1908: 992-993.

● Type locality: Dniester liman and Katlabukh lake (northern Black Sea maritime area). 


● Distribution: fresh and brackish-water waterbodies of Danube Delta, Dniester and Dnieper-Bug limans 

(northern Black Sea maritime area) (Scarlato, Starobogatov, 1972; Starobogatov et al., 2004). 

Adacna glabra Ostroumoff, 1905 B 

[= Amphidesma caspia Krynicki, 1837 non Eichwald, 1829; Adacna aspera Ostroumoff et Rosen in Ostroumoff, 

1905] 

Adacna glabra Ostroumoff, 1905: 18-19. 

● Type locality: Dniester Liman. 


● Distribution: northern Caspian Sea near Volga delta and Ural mouth (Logvinenko, Starobogatov, 1968; 


Adacna grossui (Scarlato et Starobogatov, 1972) B 

Hypanis grossui Scarlato, Starobogatov, 1972: 214, pl. 4, fig. 1 A, B. 

● Type locality: Seymeny, Dniester liman, northern Black Sea. 

● Holotype: ZIN, uncataloged. 

● Distribution: limans and lakes of the Danube delta and low-salinity parts of Dniester liman, northern Black 

Sea maritime area (Scarlato, Starobogatov, 1972). 

Adacna jalpugensis (Borcea, 1926) B 

Monodacna colorata var. jalpugensis Borcea, 1926: ??? 



● Distribution: waterbodies of Danube Delta and Inguletz River mouth, northern Black Sea maritime area 

(Scarlato, Starobogatov, 1972). 

Adacna laeviscula (Eichwald, 1829) М 

Glycimeris laeviscula Eichwald, 1829: 279. 



● Distribution: Caspian Sea; 30-60 m (Starobogatov et al., 2004). 

Adacna lucida (Borcea, 1926) B 

Monodacna colorata var. lucida Borcea, 1926: ??? 



● Distribution: north-western Black Sea maritime area (Starobogatov et al., 2004, as A. "luciea"). 

Adacna polymorpha (Logvinenko et Starobogatov, 1967) B 

Hypanis angusticostata polymorpha Logvinenko, Starobogatov, 1967: 232. 

● Type locality: central part of northern Caspian Sea, 3 m. 

● Holotype: ZIN ??? 

● Distribution: northern Caspian Sea except most freshened areas (Logvinenko, Starobogatov, 1968; Starobogatov 

et al., 2004). 

Adacna pontica (Eichwald, 1838) B 

[= Cardium pseudocardium Deshayes, 1838; Cardium ponticum Ostroumoff, 1893] 

Monodacna pontica Eichwald, 1838: 168-169. 

● Type locality: «ostio Tyrae versus Pontum Euxinum, prope Businowatajam». 


● Distribution: limans and lakes of the Danube delta and low-salinity parts of Dniester and Dnieper-Bug li-

mans, northern Black Sea maritime area (Scarlato, Starobogatov, 1972). 

Didacna Eichwald, 1838 

Type species: Cardium trigonoides Pallas, 1771 (SD Stoliczka, 1870) 

Didacna baeri (Grimm, 1877) М 

[= Cardium crassum sensu Eichwald, 1829 non Gmelin, 1791; Cardium eichwaldi Krynicki, 1837, nom. nov. 

pro Cardium crassum Eichwald, 1829 non Gmelin, 1791] 

Cardium baeri Grimm, 1877: 51-54, pl. 8, figs. 2a-c, 3a-d. 

● Type locality: three localities in the Caspian Sea, from 40º04' to 40º41' N and from 02º29' to 02º33' E, 19- 

23 fathoms. 



● Remark: The name Cardium eichwaldi Krynicki, 1837 has priority, so additional investigation is needed to 

reveal whether it can be considered a nomen oblitum. 

Didacna barbotdemarnyi (Grimm, 1877) М 

Cardium barbot-de-marnyi Grimm, 1877: 56-58, pl. 8, figs. 5a-c, 6a-d. 

● Type locality: five localities in the Caspian Sea, from 40º13' to 44º22' N and from 2º33' E to 0º 22'30" W, 

7-20 fathoms. 



Didacna longipes (Grimm, 1877) М 

Cardium longipes Grimm, 1877: 54-56, pl. 8, fig. 4a-c. 

● Type locality: 2 two localities in the Caspian Sea: 40º39' N, 0º26' E, 6 fathoms, and 40º33' N, 0º22' E, 10 

fathoms. 



Didacna parallela Bogatschev, 1932 М 

Bogatschev, 1932: 



● Distribution: southern Caspian Sea, western coast of middle Caspian Sea; 50-85 m (Logvinenko, Starobogatov, 

1968). 

Didacna profundicola Logvinenko et Starobogatov, 1966 М 

Logvinenko, Starobogatov, 1966a: 13-14, fig. 1. 

● Type locality: 39°38' N, 52°02' E (central Caspian Sea), 94 m. 

● Holotype: ZIN /10300. 


Didacna protracta protracta (Eichwald, 1841) М 

Adacna protracta Eichwald, 1841: 280, pl. 40, figs. 10, 11 (not 9, 10, as indicated in the text). 

● Type locality: "cum antecendente fossilis in argillaceo deserto circa lacum Elto nensem". 


● Distribution: Middle Caspian Sea; 25-50 m (Logvinenko, Starobogatov, 1968). 

Didacna protracta submedia Andrusov, 1911 М 

[= Didacna protracta var. media Andrusov, 1911; Didacna protracta var. grimmi Andrusov, 1911] 

Didacna protracta var. submedia Andrusov, 1911: 

● Type locality: “Hab/ cum ancedente fossilis in argillaceo deserto circa lacum Eltonensem”.



Didacna pyramidata (Grimm, 1877) М 

Cardium pyramidatum Grimm, 1877: 46-49, pl. 8, fig. 1a-d. 

● Type locality: five localities in the Caspian Sea, from 38º14'30" to 40º53' N and from 2º33' E to 0º28' E, 

20-60 fathoms. 



Didacna trigonoides trigonoides Pallas, 1771 М 

Cardium trigonoides Pallas, 1771: 478. 



● Distribution: northern Caspian Sea (Logvinenko, Starobogatov, 1968). 

Didacna trigonoides praetrigonoides Nalivkin et Anisimov, 1914 М 

Nalivkin, Anisimov, 1914: 5-6, 16-17, pl. 1, figs. 1-2. 

● Type locality: post-Pliocene of the Apsheron Peninsula. 



Hypanis Pander in Menetries, 1832 

Type species: Glycymeris plicata Eichwald, 1829 (monotypy) 

Hypanis albida Logvinenko et Starobogatov, 1967 М 

[= Cardium pseudocatillus "Abich" Grimm, 1877 non Barbot de Marny, 1869] 

Logvinenko, Starobogatov, 1967: 232. 

● Type locality: southeastward from Derbent (western Caspian Sea), 56 m. 


● Distribution: Middle and southern Caspian Sea; 0-150 m (Logvinenko, Starobogatov, 1968). 

Hypanis caspia caspia (Eichwald, 1829) B 

Corbula caspia Eichwald, 1829: 281, pl. 5, fig. 6. 

● Type locality:Caspian Sea. 


● Distribution: eastern part of northern Caspian Sea (Logvinenko, Starobogatov, 1968). 

Hypanis caspia filatovae Logvinenko et Starobogatov, 1967 М 

[= Cardium caspium sensu Grimm, 1876, non Corbula caspia Eichwald, 1829] 


● Type locality: Gulf of Baku (western Caspian Sea). 


● Distribution: southern Caspian Sea; 50 m (Logvinenko, Starobogatov, 1968). 

Hypanis caspia knipowitschi Logvinenko et Starobogatov, 1966 М 

Logvinenko, Starobogatov, 1966a: 15, fig. 2. 

● Type locality: 39°38' N, 52°08' E (central Caspian Sea), 71 m. 

● Holotype: ZIN /10560. 


Hypanis minima minima (Ostroumoff, 1907) B 

Ostroumoff, 1907: 23, textfig.



● Distribution: Aral Sea (was not found since 1970 th ) (Logvinenko, Starobogatov, 1968; Starobogatov, 1974; 

Andreeva, Andreev, 2003). 

Hypanis minima ostroumovi Logvinenko et Starobogatov, 1967 М 

[= Hypanis minima Ostroumoff, 1907 (partim)] 


● Type locality: near Ogurchinsky Island (Caspian Sea), 23. 



Hypanis plicata (Eichwald, 1829) M 

[= Adacna grimmi Andrusov, 1923] 

Glycimeris plicata Eichwald, 1829: 279. 




Hypanis regularis Starobogatov in Starobogatov, Prozorova, Bogatov et Sayenko, 2004 B 

Starobogatov et al., 2004: 48, pl. 39, fig. 6 (nom. nov. pro Adacna plicata Grimm, 1877: 66, pl. 8, fig. 14, non 

Glycymeris plicata Eichwald, 1829). 

● Type locality: Caspian Sea: Srednii Ashur Island; 39º51' N, 0º03' E, 13 fathoms; 40º14' N, 0º37' E, 8 fathoms. 


● Distribution: Caspian Sea shallows (Starobogatov et al., 2004). 

Hypanis relicta (Milaschewitsch, 1916) B 

Adacna relicta Milaschewitsch, 1916: 274-276, pl. 8, figs. 10-12. 

● Type locality: Dniester liman, Berezan liman (northern Black Sea maritime area). 


● Distribution: waterbodies of Danube Delta, Dniester and Dnieper-Bug limans (northern Black Sea maritime 

area) (Scarlato, Starobogatov, 1972). 

Hypanis semipellucida Logvinenko et Starobogatov, 1967 М 

[= Adacna edentula Grimm, 187 (partim), non Mya edentula Pallas, 1771] 

Logvinenko, Starobogatov, 1967: 232-233. 

● Type locality: off Tokmak Cape, near shoreline (Caspian Sea). 



Hypanis sidorovi Starobogatov, 1974 B 

[= Adacna laeviuscula Martens, 1882 non Eichwald, 1829] 

Hypanis minima sidorovi Starobogatov, 1974: 246, fig. 213. 

● Type locality: near well Kara-Kibir. 


● Distribution: Aral Sea; 1-10 m (probably extinct since 1970 th ) (Starobogatov, 1974; Andreeva, Andreev, 

2003). 

Hypanis vitrea vitrea (Eichwald, 1829) М 

Glycimeris vitrea Eichwald, 1829: 279. 

Type locality: 

Holotype:


Hypanis vitrea bergi Starobogatov, 1974 B 

Starobogatov, 1974: 245-246, fig. 212. 

● Type locality: near well Kara-Kibir (western coast of Aral Sea). 


● Distribution: Delta of Amu Darya River, north-western part of Aral Sea (probably extinct since 1980 th ) 

(Starobogatov, 1974; Andreeva, Andreev, 2003). 

DREISSENOIDEA 

DREISSENIDAE Gray, 1840 

(Kantor Yu.I., Vinarski M.V.) 

Dreissena Van Beneden, 1835 

Type species: Mytilus polymorphus Pallas, 1771 (monotypy) 

Dreissena bugensis (Andrusov, 1897) B, F 

Dreissensia bugensis Andrusov, 1897: 285-286, pl. 15, figs. 31-37. 

● Type locality: Bug Liman near Nikolaev City, at Ingul River mouth (across Nikolaev). 

● Lectotype (Rosenberg, Ludyansky, 1994): Paleontological Institute, Moscow, No. 451/77. 

● Distribution: western Black Sea maritime area, Dnieper River Basin (except Far North) (Starobogatov, 

1994; Starobogatov, Andreeva, 1994). 

Dreissena caspia caspia Eichwald, 1855 B 

Eichwald, 1855: 311, pl. 10, figs. 19-21. 

● Type locality: Chistyi Bank and Cheleken Island (Caspian Sea). 

● Types: possibly in the Geological Museum of SPb University. 

● Distribution: Caspian Sea (now probably extinct) (Starobogatov, 1994; Starobogatov, Andreeva, 1994). 

Dreissena caspia pallasi (Andrusov, 1897) B 

Dreissensia pallasi Andrusov, 1897: 671-672, pl. 20, figs. 33-35. 

● Type locality: Aral Sea, near St. Nicholas Island. 

● Syntypes (Rosenberg, Ludyansky, 1994): Paleontological Institute, Moscow, No. 451/113, 451/115. 

● Distribution: Aral Sea. It was last recorded in 1980 (Rosenberg, Ludyansky, 1994) and is probably extinct 

since 1990th (Starobogatov, 1994; Starobogatov, Andreeva, 1994; Andreeva, Andreev, 2003). 

Dreissena elata elata (Andrusov, 1897) B 

[= Dreissena polymorpha var. lata Andrusov, 1897; Dreissena polymorpha var. marina auctt. non Pallas, 

1771] 

Dressensia polymorpha var. elata Andrusov, 1897: 353, pl. 20, fig. 25. 

● Type locality: Kuuli Cape, Dazmyk, Apsheron (western Caspian Sea). 

● Syntype (Rosenberg, Ludyansky, 1994): Paleontological Institute, Moscow, No. 451/184. 

● Distribution: Caspian Sea (now probably extinct); down to 30 m (Starobogatov, 1994; Starobogatov, Andreeva, 

1994). 

Dreissena obtusecarinata (Andrusov, 1897) B 

Dreissensia polymorpha var. obtusecarinata Andrusov, 1897: 354. 

● Type locality: Aral Sea. 


● Distribution: Aral Sea (probably extinct since 1980th) (Starobogatov, 1994; Starobogatov, Andreeva, 1994;

Andreeva, Andreev, 2003). 

Dreissena polymopha polymorpha (Pallas, 1771) B 

[= Mytilus volgae Chemnitz, 1795; Mytilus volgensis Gray, 1825; Mytilus hagenii Baer, 1825; Mytilus arca 

Kickx, 1834; Dythalmia danubii Jay, 1835; Tichogonia chemnitzi Rossmaessler, 1835; Dreissena fluviatilis 

Bourgignat, 1856; Dreissensia belgrandi Locard, 1893; Dreissensia complanata Locard, 1893; Dreissensia 

curta Locard, 1893; Dreissensia eximia Locard, 1893; Dreissensia lacunosa Locard, 1893; Dreissensia lutetiana 

Locard, 1893; Dreissensia occidentalis Locard, 1893; Dreissensia recta Locard, 1893; Dreissensia 

servaini Locard, 1893; Dreissensia sulcata Locard, 1893; Dreissensia tumida Locard, 1893; Dreissensia ventrosa 

Locard, 1893; Dreissensia westerlundi Locard, 1893; Dreissena polymorpha var. angustiformis 

Kolesnikov, 1950; Dreissena cornuliformis Ali-zade, 1961; Dreissena gubkini Ali-zade, 1961; Dreissena 

kambai Ali-zade, 1961; Dreissena kopetdagica Ali-zade, 1961; Dreissena lunata Ali-zade, 1961; Dreissena 

nalivkini Ali-zade, 1961; Dreissena vestae Ali-zade, 1961] 

Mytilus polymorphus Pallas, 1771: 368, 435, 478. 

● Type locality: Volga and Yaik rivers, Caspian Sea. 


● Distribution: low-salinity areas of Black and Azov seas, Aral Sea, fresh waters of European Russia, eastward 

to Ural Mountains and northward to 62° N (Starobogatov, 1994; Starobogatov, Andreeva, 1994). 

Dreissena polymopha andrusovi (Brusina in Andrusov, 1897) B 

[= Dreissensia latro Andrusov, 1897] 

Dreissensia andrusovi Brusina – Andrusov, 1897: 374-375, pl. 18, figs. 21-23. 


● Lectotype (Rosenberg, Ludyansky, 1994): Paleontological Institute, Moscow, No. 451/97. 

● Distribution: northern Caspian Sea (Starobogatov, 1994; Starobogatov, Andreeva, 1994). 

Dreissena polymopha aralensis (Andrusov, 1897) B 

Dreissensia polymorpha var. aralensis Andrusov, 1897: 354-355, pl. 18, fig. 40. 

● Type locality: Aral Sea. 


● Distribution: waterbodies adjacent to Aral Sea (Starobogatov, 1994; Starobogatov, Andreeva, 1994). 

Dreissena rostriformis compressa Logvinenko et Starobogatov, 1966 B 

Logvinenko, Starobogatov, 1966a: 15-16, fig. 3. 

● Type locality: off Samur River mouth (Caspian Sea), 90 m. 


● Distribution: Caspian Sea; 70-300 m (Starobogatov, 1994; Starobogatov, Andreeva, 1994). 

Dreissena rostriformis distincta (Andrusov, 1897) B 

Dreissensia rostriformis var. distincta Andrusov, 1897: 273-276, pl. 14, figs. 18-24. 

● Type locality: numerous localities of Volga River, Caspian, Black and Marmara seas (Recent and fossil). 


● Distribution: northern and northeastern Caspian Sea; 20-50 m (Starobogatov, 1994; Starobogatov, Andreeva, 

1994). 

Dreissena rostriformis grimmi (Andrusov, 1890) B 

Dreissensia grimmi Andrusov, 1890: 41. 

● Type locality: Caspian Sea, 43°35’ N, 0°30’30» E; 1°3’ E; 30-35 fathoms. 


● Distribution: Middle Caspian Sea; 45-80 m (Starobogatov, 1994; Starobogatov, Andreeva, 1994). 

Dreissena rostriformis pontocaspica (Andrusov, 1897) B 

Dreissensia tschaudae var. pontocaspica Andrusov, 1897: 294-297, pl. 9, figs. 27-32, pl. 15, figs. 29-30.

● Type locality: several localities on Apsheron Peninsula (Caspian Sea), eastern Caspian Sea coast, Black 

Sea at 30-101 fathoms (fossil and subfossil). 

● Syntype (Rosenberg, Ludyansky, 1994): Paleontological Institute, Moscow, No. 451/41-44. 

● Distribution: southwestern and southeastern Caspian Sea; 20-80 m (Starobogatov, 1994; Starobogatov, Andreeva, 

1994). 

Mytilopsis Conrad, 1857 

Type species: Mytilus leucophaeatus Conrad, 1831 (SD) 

Mytilopsis leucophaeata (Conrad, 1831) B, F 

Mytilus leucophaeatus Conrad, 1831: p. 263-264, pl. 11, fig. 13. 

● Type locality: “Southern coast of the [eastern] United States” (Rosenberg, 2009). 

● .Types: Not traced. Most probably, in the Academy of Natiral Sciences, Philadephia (USA). 

● Distribution: USA (Gulf of Mexico, Hudson River, Upper Mississipi, New England), introduced to the 

North Sea and recently was revealed as a cryptic invader to the Dniester Liman, Black Sea (Therriault et al., 

2004). 

● Remark: Some authors consider Mytilopsis to be a subgenus of the genus Congeria (Andrusov, 1897; Starobogatov, 

1970; Babak, 1983) and subsequently the species is sometimes cited as Congeria leucophaeata 

(see Glöer & Meier-Brook, 2003). Mytilopsis was elevated to genus level by (Nuttall, 1990) and subsequent 

authors accepted this taxonomic solution (Rosenberg & Ludyansky, 1994; Gittenberger et al., 1998; Falkner 

et al., 2001). 

CORBICULOIDEA 

CORBICULIDAE Gray, 1847 


Corbicula Megerle von Mühlfeld, 1811 

Type species: Tellina fluminalis Müller, 1774 (monotypy; ICZN Opinion 335) 

Corbicula amurensis Bogatov et Starobogatov, 1994 F 

Bogatov, Starobogatov, 1994: 149, fig. 1 D-F. 

● Type locality: lower Amur near Balbinski cliff [22 km upstream of Kalinovka settlement and 337 km from 

the mouth]. 


● Distribution: lower Amur River (Bogatov, Starobogatov, 1994; Starobogatov et al., 2004). 

Corbicula consobrina (Cailliaud, 1827) F 

[= Corbicula fluminalis var. minor Kobelt, 1897; Corbicula fickeli Clessin, 1897; Corbicula heuglini Clessin, 

1897] 

Cyrena consobrina Cailliaud, 1827: 263. 



● Distribution: Azerbaijan (Kursalova, Starobogatov, 1971). 

Corbicula cor (Lamarck, 1818) F 

[= Corbicula hohenackeri Clessin, 1879; Corbicula feliciani Locard, 1883] 

Cyrena cor Lamarck, 1818: 526. 


● Types: not traced.

● Distribution: eastern Transcaucasia and planes of Anterior and Central Asia (Starobogatov et al., 2004). 

Corbicula elatior Martens, 1905 F 

[= Corbicula sujfunensis var. sujfunensis Lindholm, 1927] 

Martens, 1905: 65-66, pl. 2, fig. 5. 

● Type locality: "Provinz Kyöngsangdo, zwischen Tongnai und Kimhai. Imjingang bai Imjin, Provinz 

Kyöngkwido". 


● Distribution: southern Primorje (Razdolnaya River and other rivers of the Japan Sea basin) (Zatrawkin, 


Corbicula ferghanensis Kursalova et Starobogatov, 1971 F 

Kursalova, Starobogatov, 1971: 95. 

● Type locality: Ferghana [Uzbekistan], large aryk behind the railway station. 


● Distribution: Central Asia (Starobogatov et al., 2004) 

Corbicula finitima Lindholm, 1927 B 

[= Corbicula fluminea var. sujfunensis sensu Golikov, Scarlato, 1967 non Lindholm, 1927] 

Corbicula sujfunensis var. finitima Lindholm, 1927e: 552, Taf. 32, fig. 2. 

● Type locality: estuary of the river Artemovka. 


● Distribution: southern Primorje, brackish water waterbodies (Zatrawkin, Bogatov, 1987; Starobogatov et 

al., 2004). 

Corbicula fluminalis (Müller, 1774) F 

[= Corbicula crassula Mousson, 1854] 

Tellina fluminalis Müller, 1774: 204-205. 

● Type locality: “in fluvio Asiae Euphrat”. 

● Types: Possibly, in ZMUC (fide Dance, 1986). 

● Distribution: eastern Transcaucasia and planes of Central Asia (Kursalova, Starobogatov, 1971; Starobogatov 

et al., 2004). 

Corbicula japonica Prime, 1864 B 

[= Corbicula bifrons Reinhardt, 1877; Corbicula ovalis Reinhardt, 1877; Corbicula transversa Martens, 

1877; Cyrena yokohamensis Sowerby, 1877; Corbicula doenitziana Clessin, 1878; Corbicula fuscata atrata 

Reinhardt, 1878; Corbicula martensi Clessin, 1878; Corbicula sadoensis Pilsbry, 1901; Corbicula nipponensis 

Pilsbry, 1907; Corbicula nipponensis delicata Pilsbry, 1907; Corbicula sujfunensis Lindholm, 1927; Corbicula 

fluminalis var. extrema Lindholm, 1929; Corbicula fluminea sensu Shadin, 1952 (part.), non Müller, 

1774] 

Prime, 1864: 68, fig. 15. 

● Type locality: Japan. 

● Holotype: MCZ No. 152904. 

● Distribution: Japan Sea, South Sakhalin, South Kurile Islands, also lower reaches of Amur River and some 

lakes and rivers near the sea; 0-10 m (Zatrawkin, Bogatov, 1987; Kafanov, 1991; Starobogatov et al., 2004). 

Corbicula lindholmi Kursalova et Starobogatov, 1971 F 

Kursalova, Starobogatov, 1971: 94. 

● Type locality: lower part of Pachikheza River [Far East]. 



Bogatov, 1987; Starobogatov et al., 2004).

Corbicula nevelskoyi Bogatov et Starobogatov, 1994 F 

Bogatov, Starobogatov, 1994: 148-149, fig. 1 A-C. 

● Type locality: lower Amur near Balbinski cliff 22 km upstream of Kalinovka settlement and 337 km from 

the mouth. 



Corbicula producta Martens, 1905 F 

[= Corbicula fluminea sensu Shadin, 1952 (part.), non Müller, 1774] 

Martens, 1905: 66, pl. 2, fig. 8. 

● Type locality: "Keumgang bei Kongju, Provinz Chhungchhöngdo". 




Corbicula purpurea Prime, 1864 F 

Prime, 1864: 77, fig. 26. 

● Type locality: Antioch, Syria. 

● Lectotype (Johnson, 1959, as "holotype"): MCZ No. 176914. 

● Distribution: eastern Transcaucasia, planes of Anterior and Central Asia (Kursalova, Starobogatov, 1971; 


Corbicula sirotskii Bogatov et Starobogatov, 1994 F 

Bogatov, Starobogatov, 1994: 149-1150, fig. 1 G-I. 

● Type locality: lower Amur 7 km upstream of Maxim Gorky settlement and 397 km from the mouth. 



Corbicula tibetensis Prashad, 1929 F 

Prashad, 1929: 61-62, pl. 8, figs. 3-5. 

● Type locality: Tibet. 

● Holotype: Zoological Survey of India No. M 12837/2. 

● Distribution: mountain areas of Central Asia, basin of Syr-Darya River, eastern Kazakhstan (Kursalova, 

Starobogatov, 1971; Starobogatov et al., 2004). 

SPHAERIIDAE Jeffreys, 1862 


SPHAERIINAE Jeffreys, 1862 

Amesoda Rafinesque, 1820 

Type species: Cyclas similis Say, 1816 (SD Starobogatov, Streletzkaja, 1967) 

Amesoda asiatica (Martens, 1864) F 

Cyclas asiatica Martens, 1864b: 349, fig. 5. 

● Type locality: “am Ufer des Irtyschflusses bei Omsk” (Irtysh near Omsk, West Siberia). 

● Lectotype (Korniushin, Glaubrecht, 2001): ZMB No. 102816 – fossil (subfossil?) specimen. 

● Distribution: basins of Ob (except Irtysh) and Yenissei rivers (Starobogatov et al., 2004). 

Amesoda caperata (Westerlund, 1897) F 

Sphaerium (Cyclas) asiaticum var. caperatum Westerlund, 1897b: 130. 

● Type locality: "Sibirien. Werchojansk, Mündung des Obj".


● Distribution: Yana river basin (Westerlund, 1897b), basins of Ob (except Irtysh) and Pechora rivers (Starobogatov 

et al., 2004). 

● Remark: In Korniushin's (2001) opinion, this species is a synonym of S. solidum. 

Amesoda draparnaldii (Clessin in Westerlund, 1873) F 

Sphaerium draparnaldii Clessin – Westerlund, 1873: 512 (nom. nov. pro Cyclas lacustris Draparnaud, 1805 

[= Cyclas ovalis Férussac, 1807], non Tellina lacustris Müller, 1774) 

● Type locality: "Habite dans les lacs et les marais" (France – from title; Draparnaud, 1805: 130). 


● Distribution: Europe (except extreme north and north-east), Ural River (Stadnichenko, 1984; Pirogov et al., 


Amesoda falsinucleus Novikov in Starobogatov et Korniushin, 1986 [1987] F 

Amesoda falsinucleus Novikov – Starobogatov, Korniushin, 1986 [1987]: 37. 

● Type locality: middle Ob River, near Pershin (Tomsk Region). 


● Distribution: middle Ob River (Starobogatov et al., 2004). 

● Remark: In Korniushin's (2001) opinion, this species is a synonym of Sphaerium radiatum. 

Amesoda galitzini (Clessin, 1875) F 

Sphaerium galitzini Clessin, 1875a: 40-41, pl. 2, fig. 6. 

● Type locality: "im Auswurfe der Oka, nahe der Einmündung in die letztere zwischen Wladimir und 

Nischney-Nowgorod". 


● Distribution: Europe except extreme north and north-east (Starobogatov et al., 2004). 

● Remark: There is another spelling of the specific epithet: galitzinianum (as in caption to fig. 6 in Clessin, 

1875a); we prefer to use here the original spelling. 

Amesoda korotniewii (W. Dybowski, 1902) F 

Sphaerium korotniewii W. Dybowski, 1902: 92. 

● Type locality: south-eastern Baikal Lake. 


● Distribution: Baikal Lake, 0-3 m (Slugina et al., 1994; Slugina, Starobogatov, 2004). 

Amesoda scaldiana (Normand, 1844) F 

Cyclas scaldiana Normand, 1844: 5, figs. 1, 2. 

● Type locality: “Habite l'Escaut, а Valenciennes" (Escaut (Scheldt) River, France). 


● Distribution: Europe, except extreme north-east, basin of Irtysh (Starobogatov et al., 2004). 

● Remark: In Korniushin's (2001) opinion, this species is a synonym of Sphaerium corneum. 

Amesoda solida (Normand, 1844) F 

Cyclas solida Normand, 1844: 6, figs. 3, 4. 

● Type locality: "Habite l'Escaut canalisé, а Valenciennes" (Escaut (Scheldt) River, France). 


● Distribution: Europe (except extreme north and north-east), Ural River (Pirogov et al., 1994; Starobogatov 

et al., 2004). 

Amesoda subsolida (Clessin, 1888) F 

Shaerium subsolidum Clessin, 1888: 339. 

● Type locality: Alnarp, Skеne (Sweden) (subfossil). 


● Distribution: basin of the Baltic Sea: Ladozshskoye Lake and freshwater parts of the Finnish Gulf (Starobogatov 

et al., 2004). 

● Remark: In Korniushin's (2001) opinion, this species is a synonym of Sphaerium solidum. 

Amesoda transversalis (Westerlund, 1898) F 

Sphaerium (Corneola) transversale Westerlund, 1898: 179-180. 

● Type locality: "Sibirien. See Puiko, aus dem Magen eines Störs". 

● Lectotype (Starobogatov, Korniushin, 1986 [1987]): ZIN. 

● Distribution: basin of Ob (except Irtysh), Pechora, and Mezen rivers (Starobogatov et al., 2004). 

● Remark: In Korniushin's (2001) opinion, this species is a synonym of S. solidum. 

Nucleocyclas Alimov et Starobogatov, 1968 

Type species: Cyclas nucleus Studer, 1820 (OD) 

Nucleocyclas marisminus (Starobogatov in Starobogatov et Korniushin, 1986 [1987]) F 

Sphaerium marisminus Starobogatov – Starobogatov, Korniushin, 1987[1987]: 36, fig. 18. 

● Type locality: Mukhor Bay, Baikal Lake, depth 5-6 m. 


● Distribution: Baikal Lake, 5-6 m (Slugina et al., 1994). 

● Remark: In Korniushin's (2001) opinion, this species is a synonym of Sphaerium baicalense. 

Nucleocyclas nucleus (Studer, 1820) F 

Cyclas nucleus Studer, 1820: 93. 

● Type locality: “Gegend von Yverdon”(Yverdon-les-Bains, Switzerland). 


● Distribution: Europe, Ukraine, bogs, snall streams and ditches in swamps, wet meadows (Stadnichenko, 

1984; Korniushin, 1994). 

Nucleocyclas ovale (Férussac, 1807) F 

[= Sphaerium nitidum sensu Stadnichenko, 1984, non Clessin, 1876; Sphaerium radiatum sensu Korniushin, 

1996, 2001, non Westerlund, 1897] 

Cyclas ovalis Férussac, 1807: 128. 


● Lectotype (Falkner, 2000): NHMV, Coll. Draparnaud No. 1820.XXVI.156. 

● Distribution: Ukraine (Korniushin et al., 2002). 

Nucleocyclas radiata (Westerlund, 1897) F 

Sphaerium (Cyclas) levinodis var. radiatum Westerlund, 1897b: 131. 

● Type locality: "Sibiriens. Tomsk". 


● Distribution: Europe and western Siberia, eastward to Yenissei basin (Starobogatov et al., 2004). 

Parasphaerium Alimov et Starobogatov, 1968 

Type species: Sphaerium rectidens Starobogatov et Streletzkaja, 1967 (OD) 

Parasphaerium nitida (Clessin in Westerlund, 1876) F 

[= Sphaerium radiatum sensu Starobogatov et Streletzkaya, 1967, non Westerlund, 1897] 

Sphaerium nitida Clessin in Westerlund, 1876a: 66-67, pl., fig. 18. 

● Type locality: “Jenissei, Dudino (lat. 69º15'); norr om Lusino (lat. 68º40')". 

● Lectotype (???): SMNH. – Korniushin, 1996 

● Distribution: Europe and Siberia, 0-3 m (Stadnichenko, 1984; Slugina et al., 1994; Korniushin, 1994; Starobogatov 

et al., 2004). 

● Remark: see remark to Sphaerium levinodis.

Parasphaerium rectidens (Starobogatov et Streletzkaja, 1967) F 

Sphaerium rectidens Starobogatov, Streletzkaja, 1967: 245-246, fig. 42. 

● Type locality: lower Kolyma River, Levoe Topolskoe lake 


● Distribution: small lakes of northern Eurasia, Scandinavia to Chukchi Peninsula (Korniushin, 1994; Starobogatov 

et al., 2004). 

Rivicoliana Servain, 1888 

Type species: Cyclas rivicola Leach in Lamarck, 1818 (fixation of type species not traced) 

Rivicoliana boettgeriana (Bourguignat in Servain, 1882) F 

Sphaerium boettgerianum Bourguignat – Servain, 1882: 18. 

● Type locality: "en Pologne dans la Vistule près de Varsovie". 


● Distribution: Europe, except northern part (Starobogatov et al., 2004). 

Rivicoliana bourguignati (Lallement et Servain, 1869) F 

Sphaerium bourguignati Lallement, Servain, 1869: 46-47. 

● Type locality: "Dans la Marne, а Jaulgonne". 


● Distribution: Europe, except northern part and possibly Volga and Ural basins, Ukraine (Stadnichenko, 


Rivicoliana morini (Servain, 1882) F 

Sphaerium morini Servain, 1882: 11-12. 

● Type locality: "Mein" (vicinity of Frankfurt on Mein). 


● Distribution: Europe, except northern part (Starobogatov et al., 2004). 

Rivicoliana rivicola (Leach in Lamarck, 1818) F 

Cyclas rivicola Leach – Lamarck, 1818: 558. 

● Type locality: "France, Tamise". 


● Distribution: Europe, except northern part, Ukraine, Irtysh basin (Stadnichenko, 1984; Starobogatov et al., 

2004). 

Sphaerium Scopoli, 1777 

Type species: Tellina cornea Linnaeus, 1758 (SD Gray, 1847) 

Sphaerium baicalense W. Dybowski, 1902 F 

Sphaerium baicalense W. Dybowski, 1902: 92. 

● Type locality: SE of Baikal, off mouths or rivers. 


● Distribution: Baikal Lake, 1.5-60 m (Slugina et al., 1994; Slugina, Starobogatov, 2004). 

Sphaerium capiduliferum Lindholm, 1909 F 

Sphaerium capiduliferum Lindholm, 1909: 81, pl. 2, fig. 51. 

● Type locality: Kruglaya Inlet, Chivyrkyiski Bay, Baikal Lake. 


● Distribution: lakes of eastern Siberia (Yenissei and Lena basins), Baikal Lake, 1-5 m (Slugina et al., 1994). 

● Remark: In Korniushin's (2001) opinion, this species is a synonym of S. corneum.

Sphaerium corneum (Linnaeus, 1758) F 

[= Spaerium mamillatum var. tomentosum Westerlund, 1871] 

Tellina cornea Linnaeus, 1758: 678. 



● Distribution: small lakes and ponds of European part of Russia, Ukraine, central Asia (Stadnichenko, 1984; 

Korniushin, 1994; Izzatullaev, 2002). 

Sphaerium dybowskii Lindholm, 1909 F 

[ = Sphaerium westerlundi sensu W. Dybowski, 1902, non Clessin, 1873] 

Sphaerium dybowskii Lindholm, 1909: 93. 



● Distribution: western and eastern Siberia, Baikal, 0-3 m (Slugina et al., 1994; Slugina, Starobogatov, 2004). 


Sphaerium kozhovi Starobogatov in Starobogatov et Korniushin, 1986 [1987] F 

Starobogatov, Korniushin, 1986 [1987]: 38. 

● Type locality: Baikal Lake, Chivyrkyiski Bay, transsection Baklan — Kurbulik, depth 8 m. 

● Holotype: ZIN 



Sphaerium levinodis Westerlund, 1876 F 

Sphaerium levinodis Westerlund, 1876a: 67. 

● Type locality: Yenissei River, 62°50’N. 

● Types: not traced. Possibly, in NMG. 

● Distribution: western and eastern Siberia, Baikal, 0-5 m (Slugina et al., 1994; Starobogatov et al., 2004; 

Slugina, Starobogatov, 2004). 

● Remark: This and some other species described by Westerlund in 1876 are often considred in Russian literature 

as described in the Nachrichtsblatt der Deutschen Malakozoologishen Gesellschaft (Westerlund, 

1876b). However, the respective issue (No. 8-9) was dated August-September (i.e. September 30 for nomenclatural 

purposes), whereas the paper in the Kongliga Svenska Vetenskaps-Akademiens Handlingar (Westerlund, 

1876a) was published on April 1. 


Sphaerium mamillanum Westerlund, 1871 F 

Westerlund, 1871: 154. 

● Type locality: Sweden: Skеne, West Götland, Dalarna. 

● Syntypes: ZMB No. 21887; ZIN, SMNH, ? NMG. 

● Distribution: lakes, ponds, small rivers and running-water ditches of European part of Russia (Korniushin, 

1994). 


Sphaerium westerlundi Clessin in Westerlund, 1873 F 

[= Sphaerium ssorense Dybowski, 1902] 

Sphaerium westerlundi Clessin – Westerlund, 1873: 508. 

● Type locality: Dalarno province, Sweden. 


● Distribution: northern Europe, Siberia, Baikal Lake, northeastern Asia to Chukchi Peninsula and Kamchatka 

(Slugina et al., 1994; Starobogatov et al., 2004; Slugina, Starobogatov, 2004). 

● Remark: In Korniushin's (2001) opinion, this species is a synonym of Sphaerium corneum.

MUSCULIINAE Starobogatov in Stadnichenko, 1984 

Musculium Link, 1807 

Type species: Tellina lacustris Müller, 1774 (monotypy) 

Musculium amurense Moskvicheva in Zatrawkin et Moskvicheva, 1986 F 

Musculium amurensis Moskvicheva – Zatrawkin, Moskvicheva, 1986: 616, textfig. (1-3). 

● Type locality: small river in the Ussuri River basin in vicinities of Dormidontovka village, Khabarovsk Region. 


● Distribution: stagnant permanent waterbodies of middle and lower Amur basin, Primorje (Starobogatov et 

al., 2004). 

Musculium clessini (Paulucci in Clessin, 1882) F 

Calyculina clessini Paulucci – Clessin, 1882a: 192, pl. 4, fig. 8 [sic!]. 

● Type locality: "Ekaterinoslaw Rossiae". 


● Distribution: south of Eastern Europe, in drying waterbodies of steppes; central Asia (Izzatullaev, 2002; 


Musculium compressum (Middendorff, 1851) F 

[= Calyculina lacustris var. septentrionalis Clessin in Westerlund, 1878; Musculium lacustre auct., non 

Müller, 1774] 

Cyclas calyculata var. compressa Middendorff, 1851: 288, pl. 29, figs. 9-10. 

● Type locality: vicinities of Barnaul (Altai). 


● Distribution: shallow waters of lakes and ponds of Siberia and northern Far East (Slugina et al., 1994; Starobogatov 

et al., 2004). 

Musculium creplini (Dunker, 1845) F 

Cyclas creplini Dunker, 1845: 20. 

● Type locality: "in kleinen Lehmgruben bei Hohenkirschen, unfern Cassel". 


● Distribution: Eurasia, mountain Tajikistan, Kazakhstan (Izzatullaev, 1977; Stadnichenko, 1984; Starobogatov 

et al., 2004). 

Musculium heterodon (Pilsbry, 1895) F 

Sphaerium heterodon Pilsbry, 1895: 159, pl. 3, figs. 15-17. 

● Type locality: "Kishiu, Hizen, Japan". 


● Distribution: permanent waterbodies of south Sakhalin (Starobogatov et al., 2004). 

Musculium hungaricum (Hazay, 1881) F 

Calyculina hungaricum Hazay, 1881: 95. 

● Type locality: Budapest. 


● Distribution: Ukraine, central Asia (Stadnichenko, 1984; Izzatullaev, 2002). 

Musculium johanseni Tscheremnov, 1972 F 

Tscheremnov, 1972: 1888, textfig. 

● Type locality: lake in floodland of the Tuba River, 6 km westward of Mayaki village (upper Yenissei basin).

● Holotype: Chair of Zoology, Abakan Pedagogical Institute. 

● Distribution: lakes of the upper basin of Yenissei (Starobogatov et al., 2004). 

Musculium jurievi Zatrawkin in Zatrawkin et Moskvicheva, 1986 F 

Zatrawkin, Moskvicheva, 1986: 618, textfig. (13-15). 

● Type locality: Bolshoe Kizi Lake, 10-12 km from Marjino settlement, Khabarovsk Region. 


● Distribution: lakes of the lower Amur basin (Starobogatov et al., 2004). 

Musculium kafanovi Starobogatov in Starobogatov et Korniushin, 1987 F 

Musculium (Parvimusculium) kafanovi Starobogatov – Starobogatov, Korniushin, 1986 [1987]: 39-40, fig. 

19. 

● Type locality: Malaya Tymka River, Tymovskoe settlement (northern Sakhalin). 


● Distribution: rivers of central Sakhalin (Tym River basin) (Starobogatov et al., 2004). 

Musculium lacustre (Müller, 1774) F 

Tellina lacustris Müller, 1774: 204. 



● Distribution: basins of the Baltic and Black seas (Starobogatov et al., 2004). 

Musculium likharevi Moskvicheva in Zatrawkin et Moskvicheva, 1986 F 

Zatrawkin, Moskvicheva, 1986: 617-618, textfig. (4-6). 

● Type locality: Razdolnaya (Suifun) River basin near Ussuriisk City. 


● Distribution: rivers of southern Primorje (Starobogatov et al., 2004). 

Musculium miyadii (Mori, 1933) F 

Sphaerium miyadii Mori, 1933: 155, fig. 1D. 

● Type locality: 12 localities on Iturup and Hokkaido; "Kimon-numa (Etorohu Island, Kurile Group)" marked 

as the "type locality", though the types were not designated nor mentioned. 


● Distribution: southern Kurile Islands including Urup Island (Prozorova, 1996 [1997]a; Starobogatov et al., 

2004). 

Musculium morii Starobogatov et Budnikova, 1985 [1986] F 

Starobogatov, Budnikova, 1985 [1986]: 96-97, fig. 2. 

● Type locality: Lagunnoe Lake, Kunashir Island (Kurile Islands). 



2004). 

“Musculium moussoni Starobogatov, 1996” F 

Musculium moussoni "Starobogatov, 1996" – Starobogatov et al., 2004: 56 (nom.nov. pro Sphaerium compressum 

Mousson, 1887 non Cyclas calyculata var. compressa Middendorff, 1851). 

● Distribution: stagnant permanent waterbodies of middle and lower Amur basin, Primorje (Starobogatov et 

al., 2004). 

● Remark: this species name attributed to Starobogatov, 1996, was published in Starobogatov et al. (2004). 

To our best knowledge, this name was not proposed in any publication of Starobogatov in 1996. Publication 

of the name in Starobogatov et al. (2004) did not comply with the requirements of ICZN (primarily with Article 

16) and therefore the name Musculium moussoni is not available.

Musculium mucronulatum (Moquin-Tandon, 1855) F 

Cyclas ryckholtii var. mucronulata Moquin-Tandon, 1855: 595, pl. 53, fig. 42. 

● Type locality: "près de Mézières". 


● Distribution: south of eastern Europe, Ukraine (Stadnichenko, 1984; Starobogatov et al., 2004). 

Musculium ryckholtii (Normand, 1844) F 

Cyclas ryckholtii Normand, 1844: 7, figs. 5, 6. 

● Type locality: "eaux tranquilles des fфrets de Raismes et de Vicoigne". 


● Distribution: basin of the Baltic Sea (Starobogatov et al., 2004). 

Musculium steinii (A. Schmidt, 1850) F 

Cyclas steinii Schmidt, 1850: 118-119. 

Type locality:"Weissensee bei Berlin". 


● Distribution: basin of the Baltic Sea (Starobogatov et al., 2004). 

Musculium strictum (Normand, 1854) F 

Sphaerium strictum Normand, 1854: 3. 

● Type locality: "les fossés de forкts de Raismes et de Vicoigne, et ceux des marais d'Hélesmes" (France). 


● Distribution: lakes of south of Europe, Transcaucasia, basin of the Baltic Sea (Stadnichenko, 1984; Starobogatov 

et al., 2004). 

Musculium terverianum Dupuy, 1851-1852 F 

Cyclas terveriana Dupuy, 1851-1852: 674-675, pl. 29, fig. 9. 

● Type locality: "les mares des environs d'Auch, а la Boubée et а la Hourre" [France]. 


● Distribution: basin of the Baltic Sea, Ukraine (Stadnichenko, 1984; Starobogatov et al., 2004). 

Paramusculium Alimov et Starobogatov, 1968 

Type species: Cyclas transversa Say, 1829 (OD) 

Paramusculium inflatum (Middendorff, 1851) F 

Cyclas calyculata forma inflate Middendorff, 1851: 287-288. 

● Type locality: “Südsibirischen Kirgisensteppe abbilden lassen”. 

● Lectotype (Starobogatov, Streletzkaya, 1967): ZIN, No. 1 in systematic catalog. 

● Distribution: Altai and Yenissei River basin (Starobogatov et al., 2004). 

● Remark. Korniushin (1996) places this species in the genus Amesoda. 

Paramusculium limanicum (Moskvicheva in Zatrawkin et Moskvicheva, 1986) F 

Sphaerinova limanica Moskvicheva – Zatrawkin, Moskvicheva, 1986: 618-619, textfig. (10-12). 

● Type locality: Amur River near Nikolaevsk City. 


● Distribution: basin of Amur River (downstream of Bureya Ridge), including basin of Ussuri, liman of 

Amur (Starobogatov et al., 2004). 

PISIDIIDAE Gray, 1857 

(Kantor Yu. I., Vinarski M..V., Sysoev A.V.)

Europisidium Starobogatov in Stadnichenko, 1984 

Type species: Pisidium tenuilineatum Stelfox, 1918 (OD) 

Europisidium alpinum (Odhner, 1938) F 

Pisidium tenuilineatum f. alpina Odhner, 1938: 234, figs. 1-2, 5-8. 

● Type locality: "Wippach-Fluss bei Dornberg (Monte Spino), 10 km östlich von Görz". 


● Distribution: Europe, south of western Siberia (Stadnichenko, 1984; Starobogatov et al., 2004). 

Europisidium stelfoxi (Pirogov et Starobogatov in Timm, 1976) F 

Neopisidium stelfoxi Pirogov et Starobogatov in Timm, 1976: 49-50, fig. 5. 

● Type locality: North Caspian, in the Kirovski canal. 


● Distribution: rivers of western and southern Europe (Starobogatov et al., 2004). 

Europisidium tenuilineatum (Stelfox, 1918) F 

Pisidium tenuilineatum Stelfox, 1918: 296-298, pl. 8, figs. 1, 4-13. 

● Type locality: "Grand Junction Canal, near Marsworth Church, Bucks., and other places". 


● Distribution: Europe, except northern part, Ukraine, rivers and large lakes (Stadnichenko, 1984; Starobogatov 

et al., 2004). 

Kuiperipisidium Izzatullaev et Starobogatov, 1986 

Type species: Odhneripisidium sogdianum Izzatullaev et Starobogatov, 1986 (OD) 

Kuiperipisidium issykkulense (Izzatullaev et Starobogatov, 1986) F 

Odhneripisidium (Kuiperipisidium) issykkulense Izzatullaev, Starobogatov, 1986: 68-69, fig. 17. 

● Type locality: Kirgizia, Issyk-Kul Lake, Tonskij Bay, near Kutarga settlement. 


● Distribution: Issyk-Kul Lake, Hazarati-sho Ridge (Kirgizstan); river plains of Obi-Vazor and Zeravshan 

rivers (Tajikistan) (Izzatullaev, Starobogatov, 1986). 

Kuiperipisidium khorense (Izzatullaev et Starobogatov, 1986) F 

Odhneripisidium (Ussuripisidium) khorense Izzatullaev, Starobogatov, 1986: 69-70, fig. 18. 

● Type locality: Khabarovsk Territory [Far East], Khor River, spring. 


● Distribution: Ussuri River basin, in springs (Starobogatov et al., 2004). 

Kuiperipisidium polytimeticum (Izzatullaev et Starobogatov, 1986) F 

Odhneripisidium (Kuiperipisidium) polytimeticum Izzatullaev, Starobogatov, 1986: 66-67, fig. 15. 

● Type locality: Nuratin Ridge, Minishkor kishlak near Dzhush, spring (Uzbekistan). 


● Distribution: possibly northern Caucasus, in springs (Starobogatov et al., 2004). 

Kuiperipisidium sogdianum (Izzatullaev et Starobogatov, 1986) F 

Izzatullaev, Starobogatov, 1986: 65-66, fig. 14. 

● Type locality: Uzbekistan, northern slopes of Turkestan Ridge, Dzhizakh, Bakhmal, spring 


● Distribution: from Italy to Central Asia, possibly northern Caucasus, in springs (Starobogatov et al., 2004). 

Kuiperipisidium terekense (Kazannikov in Izzatullaev et Starobogatov, 1986) F 

Odhneripisidium (Kuiperipisidium) terekense Kazannikov – Izzatullaev, Starobogatov, 1986: 67-68, fig. 16. 

● Type locality: North Osetia, Bekan Lake in vicinities of Kardzhin village.


● Distribution: northern Caucasus, in springs; Zeravshan River Basin (Central Asia) (Starobogatov et al., 

2004; Boimurodov, 2004). 

Neopisidium Odhner, 1921 

Type species: Pisidium torquatum Stelfox, 1918 (SD Habe, 1951) 

Neopisidium moitessierianum (Paladilhe, 1866) F 

[= Pisidium punctatum Sterki, 1895; Pisidium punctatum var. armatum Sterki, 1905] 

Pisidium moitessierianum Paladilhe, 1866: 172, pl. 13, figs. 11-17. 

● Type locality: "fossé d'irrigation des praires de Maurin, au-dessous de Montpellier". 


● Distribution: rivers of Europe, except northern part (Stadnichenko, 1984; Starobogatov et al., 2004). 

Neopisidium ovatotrigonum Krivosheina, 1979 F 

Krivosheina, 1979: 604, textfig. (B). 

● Type locality: Pennoe bog near Leninogorsk City (East Kazakhstan), 2 m. 


● Distribution: type locality (Krivosheina, 1979). 

Neopisidium torquatum (Stelfox, 1918) F 

Pisidium torquatum Stelfox, 1918: 299. 



● Distribution: rivers of Europe (Stadnichenko, 1984; Starobogatov et al., 2004). 

Neopisidium trigonum (Bourguignat in Locard, 1893) F 

Pisidium trigonum Bourguignat – Locard, 1893: 148-149. 

● Type locality: "Jaulgonne (Aisne)". 


● Distribution: south of Europe, Ukraine, Ural River (Stadnichenko, 1984; Pirogov et al., 1994; Starobogatov 

et al., 2004). 

Odhneripisidium Kuiper, 1962 

Type species: Pisidium stewarti Preston, 1909 (OD) 

Odhneripisidium behningi Izzatullaev et Starobogatov, 1986 F 

Izzatullaev, Starobogatov, 1986: 58. fig. 6. 

● Type locality: Iskander-Kull Lake (Tajikistan], northern slope of Hissar Ridge. 


● Distribution: type locality; Son-Kull Lake (Kirgizstan); Zeravshan River basin (Izzatullaev, Starobogatov, 

1986; Boimurodov, 2004). 

Odhneripisidium chatyrkulense Izzatullaev et Starobogatov, 1986 F 


● Type locality: Kirgizia, Chatyr-Kul Lake. 


● Distribution: type locality; Pamir (lakes Karakul and river Kara-dzhilga), Nuraginskyi Ridge (Uzbekistan) 

(Izzatullaev, Starobogatov, 1986). 

Odhneripisidium dancei (Kuiper, 1962) F 

Pisidium dancei Kuiper, 1962: 11-14, figs. 1-4. 

● Type locality: "Busetun (Bisoutoum) dans la province de Kermanshab, en Iran".


● Distribution: springs in Pamir, vicinities of Kulyaba; (Tajikistan) springs on the shores of the lakes Issyk- 

Kul’ and Son-Kul’ (Kirgizstan) (Izzatullaev, Starobogatov, 1986). 

Odhneripisidium gafurovi Izzatullaev et Starobogatov, 1986 F 

Izzatullaev et Starobogatov, 1986: 59, fig. 7 

● Type locality: Tajikistan, eastern Pamir Mountains, Suluktysaj Lake, 9 km from Kyzyl-Rabat. 


● Distribution: central Asia (Izzatullaev, 2002). 

Odhneripisidium kungejense (Butenko et Starobogatov in Butenko, 1967) F 

Pisidium kungejense Butenko et Starobogatov – Butenko, 1967: 209-210, fig. 1 

● Type locality: Kungei Ala-Tau [Kazakhstan], mountain lake Kulsai, 2600 m above sea level. 


● Distribution: mountain areas of the Central Asia, probably in mountain Altai (Starobogatov et al., 2004). 

Odhneripisidium pamirense Izzatullaev et Starobogatov, 1986 F 


● Type locality: Eastern Pamir (Tajikistan), water body in the valley of Kara-Dzhilga River. 


● Distribution: eastern Pamir (Izzatullaev, Starobogatov, 1986). 

Odhneripisidium popovae Starobogatov et Streletzkaja, 1967 F 

Starobogatov, Streletzkaja, 1967: 257, fig. 71 

● Type locality: Tuva, Terekhol Lake (SW Siberia). 


● Distribution: Tuva (upper reaches of Yenissei to southern regions) (Starobogatov et al., 2004). 

Odhneripisidium prashadi (Odhner, 1937) F 

Pisidium prashadi Odhner, 1937a: 117-123, textfigs. 1-10, 13-14. 

● Type locality:"Zentralasiens, in der Provinz Cham. Tal des Flusses Deha-tehu (Quellgebiet des Flusses Yang-tze-kiang), 

12 300 Fuss über Meeresniveau, unter 33º N, 97º Ost". 

● Syntypes: ZIN ??? 

● Distribution: mountain areas of Central Asia, Altai (Tenga Lake) (Starobogatov et al., 2004). 

Odhneripisidium stewarti (Preston, 1909) F 

Pisidium stewarti Preston, 1909: 116, fig. 3. 

● Type locality: High Hill Gompa, Gyantse valley, in a small hill stream, 14500 feet [Tibet]. 

● Holotype: Zoological Survey of India, Calcutta. 

● Distribution: Pamir, lakes Suluktysai and Zorkul (Tajikistan); lake Son-kul (Kirgizstan) (Izzatullaev, Starobogatov, 

1986). 

Odhneripisidium terekholicum Izzatullaev et Starobogatov, 1986 F 




● Distribution: type locality (Starobogatov et al., 2004) 

Odhneripisidium tuvaense Izzatullaev et Starobogatov, 1986 F 




● Distribution: Tuva (upper reaches of Yenissei in Altai foothills to southern regions) (Starobogatov et al.,

2004). 

Pisidium Pfeiffer, 1821 

Type species: Cyclas obliqua Lamarck, 1818 [= Pisidium amnicum (Müller, 1774)] (SD Gray, 1847) 

Pisidium amnicum (Müller, 1774) F 

[= Cyclas obliqua Lamarck, 1818] 

Tellina amnica Müller, 1774: 205. 



● Distribution: Europe, Kazakhastan, Siberia, upper Yenissey Basin (Tuva, Azas Lake), Baikal and vicinities, 

basin of the Amur River (Slugina et al., 1994; Starobogatov et al., 2004; Slugina, Starobogatov, 2004; Prozorova, 

Zasypkina, 2008). 

Pisidium amurense Moskvicheva in Zatrawkin, 1985 F 

Zatrawkin, 1985a:61-62, textfig. 

● Type locality: channel Piksa in 2 km from the mouth (basin of Gassi Lake) [Far-East]. 

● Holotype: ZIN, No. 1 in systematic catalog. 

● Distribution: Amur basin includinh Ussuri (Zatrawkin, 1985a; Starobogatov et al., 2004). 

Pisidium baicalense W. Dybowski, 1902 F 

[= Pisidium dubium sensu Lindholm, 1909, non Say, 1816] 

W. Dybowski, 1902: 93 

● Type locality: Baikal Lake, off mouth of Selenga River. 



Pisidium decurtatum Lindholm, 1909 F 

[= Pisidium baicalense auct., non W. Dybowski, 1902] 

Lindholm, 1909: 83, pl. 2, F. 43-44. 

● Type locality: Baikal Lake, off mouth of Selenga River. 

● Syntypes: ZIN, №№ 3,4 in systematic catalogue (Prozorova, Zasypkina, 2008). 

● Distribution: Baikal, upper Yenissei basin (Tuva, Azas Lake), Bolin River (lower Amur basin) (Zatrawkin, 

1985a; Slugina et al., 1994; Slugina, Starobogatov, 2004; Prozorova, Zasypkina, 2008). 

Pisidium inflatum Megerle von Mühlfeld in Porro, 1838 F 

Pisidium inflatum Megerle von Mühlfeld – Porro, 1838: 121-122, pl. 2, fig. 13. 

Type locality:"Nel fiume Bevera in Brianza". 


● Distribution: Europe, except extreme north-east (Stadnichenko, 1984; Starobogatov et al., 2004). 

Pisidium orientale Prozorova, 1995 F 

Prozorova, 1995: 34-35, textfig. 

● Type locality: Spasovka River mouth (Khanka Lake Basin). 


● Distribution: rivers of Ussuri and Razdolnaya (Suifun) basins (Starobogatov et al., 2004). 

EUGLESIDAE Pirogov et Starobogatov, 1974 

EUGLESINAE Pirogov et Starobogatov, 1974

Amureuglesa Kohniushin, 1996 

Type species: Euglesa khurbaense Zatrawkin, 1987 (OD) 

Amureuglesa khurbaensis (Zatrawkin, 1987) F 

Euglesa (Casertiana) khurbaensis Zatrawkin, 1987: 49-51, textfig. (I). 

● Type locality: first floodland lake of Bureya River near Chekunda settlement (old settlement), Verkhnebureinsky 

District, Khabarovsk Territory. 


● Distribution: lower Priamurye, rivers and lakes (Starobogatov et al., 2004). 

Amureuglesa kruglovi (Zatrawkin, 1987) F 

Euglesa (Cingulipisidium) kruglovi (Zatrawkin, 1987: 52, textfig. (III). 

● Type locality: lake of floodland of Amur channel Ukhta, left shore upstream of Kolchem village, Ulcha 

District of Khabarovsk Territory. 


● Distribution: lower Priamurye, rivers and lakes (Starobogatov et al., 2004). 

Cingulipisidium Pirogov et Starobogatov, 1974 

Type species: Pisidium nitidum Jenyns, 1832 (OD) 

Cingulipisidium cingulatum (Anistratenko et Starobogatov, 1990 [1991]) F 

Euglesa (Cingulipisidium) cingulata Anistratenko, Starobogatov, 1990 [1991] 

● Type locality: Germany, Bodensee near Nonnenhorn. 


● Distribution: Europe and western Siberia, lakes and rivers (Starobogatov et al., 2004). 

Cingulipisidium crassum (Stelfox, 1918) F 

Pisidium nitidum var. crassa Stelfox, 1918: 295-296, pl. 7, figs. 5-8. 

● Type locality: "Grand Junction Canal near Marsworth Church, Bucks.". 


● Distribution: lakes of north of Europe, Siberia, Ukraine, Ural River (Stadnichenko, 1984; Pirogov et al., 


Cingulipisidium depressinitidum (Anistratenko et Starobogatov, 1990[1991]) F 

Euglesa (Cingulipisidium) depressinitida Anistratenko, Starobogatov, 1990[1991]: 14, fig. 1. 

● Type locality: Ukraine, Cherkassk Region, spring in the settlement Radivanovka. 


● Distribution: Europe and western Siberia, lakes and rivers (Starobogatov et al., 2004). 

Cingulipisidium fedderseni (Westerlund, 1890) F 

Pisidium nitidum var. fedderseni Westerlund, 1890: 24. 

● Type locality: "Island". 


● Distribution: Europe and south of western Siberia, central Asia; lakes and rivers (Stadnichenko, 1984; Izzatullaev, 


Cingulipisidium feroense Korniushin, 1991 F 

Korniushin, 1991: 21-22, fig. 1 (17-20). 

Type locality:Iceland, Fero. 


● Distribution: Europe, western Siberia, Central Asia, lakes and rivers (Izzatullaev, 2002; Starobogatov et al.,

2004). 

● Remark: The authorship of the species was credited by Korniushin to "Mörch in coll.", which was based on 

the label written by Westerlund. Korniushin gave a "redescription" of the species, stating that "the species 

probably has never been described". No type material was selected, though the description was based on a 

single specimen. 

Cingulipisidium juliae (Stadnichenko, 1980) F 

Euglesa (Cingulipisidium) juliae Stadnichenko, 1980: 29. 

● Type locality: spring near Kiparisnoye village (Crimea). 

● Holotype: Zoological Institute of Ukranian Academy of Sciences. 

Distribution: Crimea (Stadnichenko, 1984). 

● Remark: this species was not listed by Korniushin (1996) in his comprehensive monograph on Palaearctic 

Pisidioidea and its reality should be proven. 

Cingulipisidium khurdinensis (Pirogov et Starobogatov in Stadnichenko, 1984) F 

Euglesa (Cingulipisidium) khurdinensis Pirogov et Starobogatov in Stadnichenko, 1984: 255-256, fig. 82. 

● Type locality: Bolshoj Karabulak bayou (Astrakhan Region, the Volga delta). 


● Distribution: south of Europe, eastward to Volga delta, lakes and протоках (Starobogatov et al., 2004). 

Cingulipisidium kozhovi (Starobogatov et Streletzkaja, 1967) F 

Galileja kozhovi Starobogatov, Streletzkaja, 1967: 254, fig. 64. 

● Type locality: Malyi Posolskyi Sor, Baikal Lake. 


● Distribution: Baikal and vicinities, 0-5 m (Slugina et al., 1994; Starobogatov et al., 2004; Slugina, Starobogatov, 

2004). 

Cingulipisidium nitidum (Jenyns, 1832) F 

Pisidium nitidum Jenyns, 1832: 304-305, pl. 20, figs. 7-8. 

● Type locality: Cambridgeshire. 


● Distribution: rivers and lakes of Europe and Siberia; Central Asia (Izzatullaev, 2002; Starobogatov et al., 

2004). 

Cingulipisidium peipsi (Timm, 1976) F 

Euglesa peipsi Timm, 1976: 47, fig. 3, 4(IV). 

● Type locality: Lales Peipsi, Lämmijärv and Pihkva (Estonia). 

● Types: Vörtsjärv Limnological Station. 

● Distribution: Estonia, Ukraine (Stadnichenko, 1984). 


Pisidioidea and its reality should be proven. 

Cingulipisidium pulchricingulatum (Starobogatov et Budnikova, 1985 [1986]) F 

Euglesa (Cingulipisidium) pulchricingulata Starobogatov, Budnikova, 1985 [1986]: 101-102, fig. 6. 



● Distribution: southern Kurile Islands including Urup Island (Prozorova, 1996 [1997]b; Starobogatov et al., 

2004). 

Cingulipisidium rotundicostatum (Starobogatov et Streletzkaja, 1967) F 

Galileja rotundicostata Starobogatov, Streletzkaja, 1967: 253, fig. 62. 

● Type locality: Kolyma River valley (lower part), channel of the small river Pantelelkha. 


● Distribution: Kolyma basin (Starobogatov, Streletzkaja, 1967; Starobogatov et al., 2004). 

Cingulipisidium shadini (Johansen, 1937) F 

Pisidium (Eupisidium) shadini Johansen, 1937: 195-197, fig. 2. 

● Type locality: Srednee Multinskoe Lake, Multa River valley (Katun Mountain Range, Altai). 


● Distribution: Altai (Starobogatov et al., 2004). 

● Remark: In many respects this species is very similar to Euglesa tanuga Timm, 1975 from Pskovsko- 

Chudskoye Lake (Starobogatov et al., 2004). 

Cingulipisidium splendens (Baudon, 1857) F 

Pisidium nitidum var. splendens Baudon, 1857: 25-26, pl. 1, fig. B. 

● Type locality: lake in Annecy (France). 


● Distribution: Europe and Siberia, eastward to basin of Yenissei River inclusive (Stadnichenko, 1984; Starobogatov 

et al., 2004). 

Cingulipisidium tanuga Timm, 1975 F 

Euglesa tanuga Timm, 1975: 223, 225, fig. 8. 

● Type locality: Lake Vörtsjärv. 

● Holotype: Vörtsjärv Limnological Station. 

● Distribution: Lake Vörtsjärv (Timm, 1975). 

● Remark: the specific epithet of this species is usually spelled as tanugum. This is wrong, since the name is 

based on the Estonian noun (without any gender) meaning "wearing a female cap, or kokoshnik" (Tarmo 

Timm, personal communication). 

Conventus Pirogov et Starobogatov, 1974 

Type species: Pisidium conventus Clessin, 1877 (OD) 

Conventus akkesiensis (Mori, 1938) F 

Pisidium conventus akkesiense Mori, 1938b: 259, 274, pl. 11, fig. 18a-d. 

● Type locality: Akkesi, Hokkaido. 

● Holotype: Otu Hydrobiological Station. 

● Distribution: southern Kurile Islands (Iturup and Zeliyony) (Prozorova, 1996 [1997]b). 

Conventus conventus (Clessin, 1877) F 

Pisidium conventus Clessin, 1877c: 181-182, pl. 3, fig. 5. 

● Type locality: "Im Starnberger See bei Tutzing, bei c. 50 m. Tiefe". 


● Distribution: Europe, lakes of Altai, Kazakhastan, Novaya Zemlya, Yamal, Siberia eastward to Kamchatka 

(Stadnichenko, 1984; Starobogatov et al., 2004). 

Conventus dybowskii Slugina et Starobogatov in Slugina, Starobogatov et Korniushin, 1994 F 

Slugina et al., 1994: 144-145, fig. 16 G-I. 

● Type locality: Baikal Lake, Peschanaya Bay, 20 m. 



Conventus iturupensis Prozorova, 1996 [1997] F 

Prozorova, 1996 [1997]b: 43-44, fig. 1d, 2d. 

● Type locality: Iturup Island [southern Kurile Islands], spring swamp near river Glush (Slavyanskaya Bay). 


● Distribution: South and Middle Kurile Islands (Starobogatov et al., 2004).

Conventus lamuanus Slugina et Starobogatov in Slugina, Starobogatov, Korniushin, 1994 F 

Slugina et al., 1994: 144, fig. 16 D-F. 

● Type locality: Baikal Lake, Dagarskaya Inlet, 9 m. 



Conventus occultus Prozorova, 1996 [1997] F 

Prozorova, 1996 [1997]b: 42-43, fig. 1c, 2c. 

● Type locality: Zelenyi Island [southern Kurile Islands], spring swamp near Utinoye Lake. 


● Distribution: southern Kurile Islands (Iturup and Zelenyi) (Prozorova, 1996 [1997]b; Starobogatov et al., 

2004). 

Conventus raddei (W. Dybowski, 1902) F 

Pisidium raddei W. Dybowski, 1902: 95. 

● Type locality: southern part of Baikal Lake, 2-60 m. 



Conventus subtilis Prozorova, 1996 [1997] F 

Prozorova, 1996 [1997]b: 41, fig. 1b, 2b. 

● Type locality: Zelenyi Island [southern Kurile Islands], spring swamp near Utinoye Lake. 


● Distribution: southern Kurile Islands (Iturup and Zelenyi) (Prozorova, 1996 [1997]b; Starobogatov et al., 

2004). 

Conventus urinator (Clessin, 1877) F 

Pisidium urinator Clessin, 1877a: 70, pl. 8, figs. 16-18. 

● Type locality: "Europa, im Züricher See zwischen 28-50 M. Tiefe". 


● Distribution: Ukraine, northern Eurasia (eastward to Kamchatka) (Stadnichenko, 1984; Starobogatov et al., 

2004). 

Cyclocalyx Dall, 1903 

Type species: Pisidium scholtzii Clessin, 1873 (OD) 

Cyclocalyx angarensis (Slugina et Starobogatov in Slugina, Starobogatov et Korniushin, 1994) F 

Euglesa (Cyclocalyx) angarensis Slugina et Starobogatov in Slugina, Starobogatov, Korniushin, 1994: 135- 

136, fig. 13 J-L. 

● Type locality: Irkutsk reservoir off Cheremshanaya valley, 25 m. 


● Distribution: Irkutsk water reservoir (Slugina et al., 1994; Starobogatov et al., 2004). 

Cyclocalyx chaunensis (Prozorova, 1988) F 

Euglesa (Cyclocalyx) chaunensis Prozorova, 1988a: 1578-1579, fig. 2 ж, з, è (zh, z, i). 

● Type locality: large floodland-thermokarst lake in 20-km vicinity of Chaun station of the Institute of Biological 

Problems of the North, Far Eastern Branch of the USSR Academy of Sciences. 


● Distribution: north-east of Siberia (basin of Chaun River), lakes (Starobogatov et al., 2004).

Cyclocalyx cor (Starobogatov et Streletzkaja, 1967) F 

Galileja lapponica cor Starobogatov, Streletzkaja, 1967: 251, fig. 60. 

● Type locality: quaternary deposits of the river Malyi Anui. 


● Distribution: Siberia and north of Far East including Kamchatka and Baikal Lake (Slugina et al., 1994; 

Starobogatov et al., 2004; Slugina, Starobogatov, 2004). 

Cyclocalyx cyclocalyx (Starobogatov et Budnikova, 1985 [1986]) F 

Euglesa (Cyclocalyx) cyclocalyx Starobogatov, Budnikova, 1985 [1986]: 97-99, fig. 3. 




2004). 

Cyclocalyx dubrueili (Baudon, 1872) F 

Pisidium dubrueili Baudon, 1872: 30-32, pl. 3, figs. 1-4. 

● Type locality: "Saint-Félix (Oise)". 


● Distribution: western and southern part of Eastern Europe, lakes and small permanent waterbodies (Stadnichenko, 


Cyclocalyx fragilis (Clessin in Westerlund, 1873) F 

Pisidium obtusale δ solidum Clessin – Westerlund, 1873: 548. 

● Type locality: Blekinge vid Ronneby I ett skogskärrr nära landsvägen till Langasjö”. 


● Distribution: basin of the Baltic Sea, basin of Dnieper, small permanent waterbodies (Stadnichenko, 1984; 


Cyclocalyx gibbus (Prozorova, 1988) F 

Euglesa (Cyclocalyx) gibba Prozorova, 1988a: 1579, fig. 3 а, б, в (a, b, v). 

● Type locality: various floodland-thermokarst lakes in 20-km vicinity of Chaun station of the Institute of 

Biological Problems of the North, Far Eastern Branch of the USSR Academy of Sciences. 


● Distribution: north-eastern Siberia (Chaun River basin), lakes (Starobogatov et al., 2004). 

Cyclocalyx globosus (Prozorova, 1988) F 

Euglesa (Cyclocalyx) globosa Prozorova, 1988a: 1577-1578, fig. 2 г, ä, е (g, d, e). 

● Type locality: various floodland-thermokarst lakes in 20-km vicinity of Chaun station of the Institute of 

Biological Problems of the North, Far Eastern Branch of the USSR Academy of Sciences. 


● Distribution: north-eastern Siberia (Chaun River basin), lakes (Starobogatov et al., 2004). 

Cyclocalyx gurvichi (Izzatullaev et Starobogatov in Izzatullaev, 1985) F 

Euglesa (Cyclocalyx) gurvichi Izzatullaev, Starobogatov — Izzatullaev, 1985: 243-244, fig. (А). 




Cyclocalyx hinzi (Kuiper, 1975) F 

Pisidium hinzi Kuiper, 1975: 27-31, textfigs. 1-6, 28, 30. 

● Type locality: "Buetjörni (= Davggeluobal), 147 m NN, Flachsee (tiefste Stelle 9 m), Varangerhalbiusel im 

NO Norwegens". 


● Distribution: lakes of northern Eurasia, from Scandinavia to Kamchatka (Starobogatov et al., 2004).

Cyclocalyx jacuticus (Starobogatov et Streletzkaja, 1967) F 

Galtteja jacutica Starobogatov, Streletzkaja, 1967: 253-254, fig. 63. 

● Type locality: Kolyma River valley (lower part), Viska Karetovskaja below the Proryv Lake. 


● Distribution: Kolyma River basin (Starobogatov et al., 2004). 

Cyclocalyx japonicus (Pilsbry et Hirase, 1908) F 

Pisidium japonicum Pilsbry, Hirase, 1908: 35, fig. 1. 

● Type locality: Akkeshi, Kushiro, Yesso, Japan. 


● Distribution: Kurile Islands (Prozorova, 1996 [1997]a). 

Cyclocalyx johanseni (Dolgin et Korniushin, 1994) F 

Euglesa johanseni Dolgin, Korniushin, 1994: 12-13, fig. 1 г (g). 

● Type locality: Yamal, lake system Yarroto, small lake. 


● Distribution: northern Siberia, from Khatanga to Yana, lakes (Starobogatov et al., 2004). 

Cyclocalyx lapponicus (Clessin in Westerlund, 1873) F 

Pisidium scholtzi β lapponicum Clessin – Westerlund, 1873: 540. 

● Type locality: “I Sverige… i Lappland…”. 


● Distribution: north of Europe and Siberia, eastward to Lena River, in lakes (Starobogatov et al., 2004). 

Cyclocalyx magnificus (Clessin in Westerlund, 1873) F 

Pisidium obtusale β magnificum Clessin – Westerlund, 1873: 547-548. 

● Type locality: “vid Göteborg, i Upland vid Harg”. 


● Distribution: Baltic Sea basin and central part of Dnieper basin, small permanent and semi-permanent waterbodies 


Cyclocalyx mitchelli (Prashad, 1925) F 

Pisidium mitchelli Prashad, 1925: 419-4210, pl. 7, figs. 11, 11a, pl. 8, fig. 10. 

● Type locality: Manasbal Lake [India]. 

● Holotype: Zoological Survey of India, Calcutta, M 12533/2. 

● Distribution: Central Asia (Izzatullaev, 2002). 

Cyclocalyx obtusalis (Lamarck, 1818) F 

Cyclas obtusale Lamarck, 1818: 559. 

● Type locality: France. 


● Distribution: Europe and Siberia, central Asia. small permanent and semi-permanent waterbodies (Stadnichenko, 

1984; Izzatullaev, 2002; Starobogatov et al., 2004). 

Cyclocalyx ovatus (Prozorova, 1988) F 

Euglesa (Cyclocalyx) ovata Prozorova, 1988a: 1579, fig. 3 г, ä, е (g, d, e). 

● Type locality: large floodland-thermokarst lakes in 20-km vicinity of Chaun station of the Institute of Biological 



● Distribution: north-eastern Siberia (Chaun River basin), lakes (Starobogatov et al., 2004).

Cyclocalyx scholtzii (Clessin, 1873) F 

Pisidium scholtzii Clessin, 1873a: 23-24, pl. 1, fig. 1. 

● Type locality: "Strassengraben bei Breslau". 


● Distribution: Europe and western Siberia, small running and stagnant permanent waterbodies (Stadnichenko, 


Cyclocalyx solidus (Clessin in Westerlund, 1873) F 

Pisidium obtusale ε solidum Clessin – Westerlund, 1873: 548. 

● Type locality: “Fuirendal Skov, skogsdam vid Holte station, Ellemose vid Valdbyguard, torfgraf I Porsmosen 

vid Sveibaek nеra Silkeborg”. 


● Distribution: southern part of Eastern Europe, small permanent and semi-permanent waterbodies (Stadnichenko, 


Euglesa Leach in Jenyns, 1832 

Type species: Tellina henslowiana Leach in Sheppard, 1823 (monotypy) 

Euglesa altiumbonata Starobogatov et Budnikova, 1985 [1986] F 

Starobogatov, Budnikova, 1985 [1986]: 110-111, fig. 14 

● Type locality: Lagunnoe Lake, Kunashir Island (Kurile Islands) 


● Distribution: Kunashir Island (Prozorova, 1996 [1997]a; Starobogatov et al., 2004). 

Euglesa ambigua Starobogatov et Budnikova, 1985 [1986] F 

Starobogatov, Budnikova, 1985 [1986]: 107-108, fig. 11 



● Distribution: Kunashir Island (Prozorova, 1996 [1997]a). 

Euglesa buchtarmensis Krivosheina, 1978 F 

[= Euglesa dymy Stadnichenko, 1980] 

Krivosheina, 1978: 1496-1497, textfig. (12). 

● Type locality: Glatkovsky Bay, Oktyabrsky area of mountain-valley part of Bukhtarma Water Reservoir 

(upper Irtysh basin, East Kazakhstan). 


● Distribution: Europe, south of western Siberia, mountain area of the central Asia, Altai, lakes (Izzatullaev, 

Korniushin, 1993; Starobogatov et al., 2004). 

Euglesa casertana (Poli, 1791) F 

[= Pisidium cinereum Alder, 1838] 

Cardium casertanum Poli, 1791: 65. 

● Type locality: Sicilia near Palermo (fide Westerlund, 1890). 

● Lectotype: ??? 

● Distribution: lakes of southern Europe, Ukraine, Crimea, Ural River (Stadnichenko, 1984; Pirogov et al., 


Euglesa cavatica (Shadin, 1952) F 

Pisidium cavaticum Shadin, 1952: 338-339, fig. 310. 

● Type locality: waterbody of cave in ravine of upper reaches of the Khosta River. 



Euglesa crassicardo, Krivosheina, 1976 

Krivosheina, 1976: 29, fig. 2(2). F 

● Type locality: Monastyrskoye Lake, Eastern Kazakhstan Region. 

● Types: not traced. ZIN (?). 

● Distribution: lakes and rivers of southern Altai; central Asia (Izzatullaev, 2002; Starobogatov et al., 2004). 

Euglesa crimeana Stadnichenko, 1980 F 

Stadnichenko, 1980: 30. 

● Type locality: Karani-Hoba cave (Krimea) 

● Holotype: Zoological Institute of Ukranian Ac. Sci. 

● Distribution: mountains of Crimea, Carpathians (Starobogatov et al., 2004). 

Euglesa curta (Clessin, 1874) F 

Pisidium fossarinum var. curtum Clessin, 1874b: 34. 

● Type locality: "Bayern". 


● Distribution: Europe, mountain area of the central Asia (Izzatulaev, Korniushin, 1993; Starobogatov et al., 

2004). 

Euglesa depressissima Krivosheina, 1978 F 

Krivosheina, 1978: 1497-1498, textfig. (13). 

● Type locality: dam of Bobrovka River at Bobrovka village (upper Irtysh basin, East Kazakhstan). 


● Distribution: lakes of southern Altai (Starobogatov et al., 2004). 

Euglesa eichwaldi Stadnichenko, 1983 F 

Stadnichenko, 1983: 25-26, fig. 2. 

● Type locality: Simeiz settlement of Crimean Region, brook with spring source near motorway Yalta- 

Simeiz. 


● Distribution: Europe (Starobogatov et al., 2004). 

Euglesa elliptica (Mori, 1938) F 

Pisidium ellipticum Mori, 1938b: 255, 259, 270, pl. 10, fig. 13. 

● Type locality: Noga-Ike, near the summit of Mt. Kisokoma, Nagano Ken. 


● Distribution: southern Kurile Islands (Kunashir) (Prozorova, 1996 [1997]a). 

Euglesa fossarina (Clessin in Westerlund, 1873) F 

Pisidium fossarinum Clessin – Westerlund, 1873: 544-546. 

● Type locality: “I Sverige tvifselsutan I de flesta trakter, sеson tagen flerestädes i Skеne, Blekinge, WGötland, 

ÖGötland, Westmanland och Dalarne”. 

● Syntypes: SMF. 

● Distribution: Ukraine, Ural River (Stadnichenko, 1984; Pirogov et al., 1994). 

Euglesa granum (Lindholm, 1909) F 

Pisidium granum Lindholm, 1909: 87. 

● Type locality: Baikal Lake, Dagarskaya Inlet, 10.5 m. 


● Distribution: Baikal Lake (Slugina et al., 1994; Slugina, Starobogatov, 2004). 

Euglesa hissarica Izzatullaev, 1985 F 

Euglesa (Cingulipisidium) hissarica Izzatullaev et Starobogatov – Izzatullaev, 1985: 244-245, fig. (Б).



● Distribution: mountain area of the central Asia (Izzatullaev, Korniushin, 1993). 

● Remark: Probably a junior synonym of E. obliquata (Korniushin, 1996). 

Euglesa hokkaidoensis Starobogatov in Starobogatov, Prozorova, Bogatov et Sayenko, 2004 F 

Starobogatov et al., 2004: 71-72, pl. 69, figs. 13-15 (nom. nov. pro Pisidium humeriforme Mori, 1938 non 

Pisidium casertanum var. humeriforme Stelfox, 1918). 



● Distribution: Iturup Island (South Kurile Islands) (Starobogatov et al., 2004). 

Euglesa hoyeri (Clessin in Esmark et Hoyer, 1886) F 

Pisidium hoyeri Clessin – Esmark, Hoyer, 1886: 120. 

Type locality:"Tromsö, Renö". 


● Distribution: lakes of Europe (Starobogatov et al., 2004). 

Euglesa inflata (Mori, 1938) F 

Pisidium cinereum inflatum Mori, 1938b: 256, 259, 272-273, pl. 11, fig. 19. 

● Type locality: Tadeno-Umio, near Nikko, Totigi-Ken. 


● Distribution: South Kurile Islands including Urup Island (Prozorova, 1996 [1997]a; Starobogatov et al., 

2004). 

Euglesa intermedia (Gassies, 1855) F 

[= Pisidium ibericum Clessin, 1873; Pisidium heldreichi Clessin, 1877; Pisidium cedrorum Clessin, 1877; 

Euglesa solodovnikovi Stadnichenko, 1983] 

Pisidium intermedium Gassies, 1855: 11-12, pl. 1, fig. 4. 

● Type locality: "а Saint-Féréol près Agen". 


● Distribution: lakes and rivers of Southern and Central Europe and Central Asia (Stadnichenko, 1984; Izzatullaev, 

Korniushin, 1993; Korniushin, 1996; Starobogatov et al., 2004). 

Euglesa irtyschensis Krivosheina, 1976 F 

Krivosheina, 1976: 28-29, fig. 2(3). 


● Types: not traced. ZIN? 

● Distribution: lakes and rivers of southern Altai; central Asia (Izzatullaev, 2002; Starobogatov et al., 2004). 

Euglesa jaudouiniana (Gassies, 1855) F 

[= Pisidium casertanum var. humeriforme Stelfox, 1918] 

Pisidium jaudouinianum Gassies, 1855: 18-19, pl. 2, fig. 2. 

● Type locality: "dans la Garonne а Agen". 


● Distribution: lakes and rivers of Europe, mountain area of the central Asia (Izzatullaev, Korniushin, 1993; 


Euglesa klucharevae Starobogatov et Budnikova, 1985 [1986] F 




● Distribution: Kunashir Island (Prozorova, 1996 [1997]a; Starobogatov et al., 2004).

Euglesa koltschomensis Zatrawkin, 1987 F 

Zatrawkin, 1987: 51-52, textfig (II). 

● Type locality: lake of floodland of Amur channel Ukhta, left shore of the channel, downstream of Kolchem 

village, Ulcha District of Khabarovsk Territory. 


● Distribution: lower Priamurye (Starobogatov et al., 2004). 

Euglesa korotnevi (Lindholm, 1909) F 

Pisidium korotnevi Lindholm, 1909: 85, Taf. 2, fig. 47-48. 

● Type locality: Baikal Lake, Listvennichnoye. 


● Distribution: Baikal Lake, 1.5-60 m (Slugina et al., 1994; Slugina, Starobogatov, 2004). 

Euglesa latiumbonata Korniushin, 1991 F 

Euglesa (Cyclocalyx) latiumbonata Korniushin, 1991: 20, fig. 1 (8-10). 

● Type locality: floodland of Nerussa River, Smelizh village, Suzem District, Bryansk Region. 


● Distribution: southern Europe (Starobogatov et al., 2004). 

Euglesa likharevi (Korniushin, 1991) F 

Cingulipisidium (Potamopisidium) likharevi Korniushin, 1991: 17, fig. 1 (1-4). 

● Type locality: small lake westward of Slovechno town, Zhitomir Region (Ovruch Range). 


● Distribution: rivers and lakes of eastern Europe (Starobogatov et al., 2004). 

Euglesa ljovushkini (Starobogatov, 1962) F 

Pisidium ljovuschkini Starobogatov, 1962: 49-50, fig. 3. 

● Type locality: Tzebelda Cave, Caucasus. 



Euglesa minuta (Kozhov, 1936) F 

Pisidium korotniewi var. minutum (part.) Kozhov, 1936: 87. 

● Type locality: Baikal, Berhin Bay, 5 m. 



Euglesa miyadii Starobogatov in Starobogatov, Prozorova, Bogatov et Sayenko, 2004 F 

Starobogatov et al., 2004: 72, pl. 71, figs. 7-9 (nom. nov. pro Pisidium ponderosum Mori, 1938 non Pisidium 

casertanum var. ponderosa Stelfox, 1918). 



● Distribution: Iturup Island (South Kurile Islands) (Starobogatov et al., 2004). 

Euglesa montana Krivosheina, 1976 F 

Krivosheina, 1976: 30, fig. 2(4). 

● Type locality: A channel of the Irtysh River near Ust’-Kamenogorsk Town, Eastern Kazakhstan. 


● Distribution: lakes and rivers of southern Altai (Starobogatov et al., 2004). 

Euglesa nikkoensis (Mori, 1938) F 

Pisidium cinereum nikkуense Mori, 1938b: 256, 259, 272, pl. 10, figs. 14c, 16.

● Type locality: Kotoku-Numa, near Nikko, Totigi-Ken. 


● Distribution: southern Kurile Islands (Iturup and Kunashir) (Prozorova, 1996 [1997]a; Starobogatov et al., 

2004). 

Euglesa obensis Starobogatov, Anistratenko et Dolgin in Anistratenko et Starobogatov, 1990 [1991] F 

Anistratenko, Starobogatov, 1990 [1991]: 16-17, fig. 3. 

● Type locality: Tyumen’ district, lake on the left bank of River Malaya Ob' [SW Siberia]. 


● Distribution: Europe, western Siberia, rivers and lakes (Starobogatov et al., 2004). 

Euglesa obliquata (Clessin in Martens, 1874) F 

[= Pisidium hispanicum Clessin, 1877] 

Pisidium obliquatum Clessin – Martens, 1874: 36-37, pl. 3, fig. 31. 

● Type locality: "aquas prope Djam in valle Sarafschan". 


● Distribution: Europe, mountain area of the central Asia (Stadnichenko, 1984; Izzatullaev, Korniushin, 

1993; Izzatullaev, 2002; Starobogatov et al., 2004). 

Euglesa personata (Malm, 1855) F 

Pisidium personatum Malm, 1855: 107, fig. 

● Type locality: Sweden, Göteborg region, near Hofås. 

● Types: SMF. 

● Distribution: Europe (Stadnichenko, 1984; Starobogatov et al., 2004). 

Euglesa platyponderosa Starobogatov et Korniushin in Korniushin, 1991 F 

Korniushin, 1991: 20-21, fig. 1 (11-13). 

● Type locality: Boden Lake (Germany). 


● Distribution: rivers and lakes of Europe (Starobogatov et al., 2004). 

Euglesa platyvalva Slugina et Starobogatov in Slugina, Starobogatov et Korniushin, 1994 F 

Slugina et al., 1994: 139, fig. 15 A-C. 

● Type locality: Baikal Lake, Dagarskaya Inlet, 5 m. 


● Distribution: type locality (Slugina et al., 1994; Slugina, Starobogatov, 2004). 

Euglesa ponderosa (Stelfox, 1918) F 

Pisidium casertanum var. ponderosum Stelfox, 1918: 294-295, pl. 7, figs. 31-34. 

● Type locality: "Grand Junction Canal at Cowroast Lock, Dudswell, Hetrs., the Thames, the Barrow, Nore, 

and Suir in Ireland". 


● Distribution: Europe, south of western Siberia, Altai (Stadnichenko, 1984; Starobogatov et al., 2004). 

Euglesa rivularis (Clessin, 1874) F 

Pisidium rivulare Clessin, 1874b: 30-31, pl. 3, figs. 7-11. 

● Type locality: "Europa, Bayern". 


● Distribution: Europe, Irtysh basin (Stadnichenko, 1984; Starobogatov et al., 2004). 

Euglesa rotundovata Krivosheina, 1976 F 

Krivosheina, 1976: 27-28, fig. 2(1).



● Distribution: southern Altai, Ukraine (Stadnichenko, 1984; Starobogatov et al., 2004). 

Euglesa steenbuchi (Møller, 1842) F 

Cyclas steenbuchi Møller, 1842: 93. 

● Type locality: Greenland. 


● Distribution: lakes of northern Eurasia (Starobogatov et al., 2004). 

Euglesa subcinerea Starobogatov et Budnikova, 1985 [1986] F 





2004). 

Euglesa subdepressa Starobogatov et Budnikova, 1985 [1986] F 





Euglesa subfossarina Starobogatov et Budnikova, 1985 [1986] F 





Euglesa subgranum Slugina et Starobogatov in Slugina, Starobogatov, Korniushin, 1994 F 

Slugina, Starobogatov, Korniushin, 1994: 138, fig. 14 G-I. 

● Type locality: Baikal Lake, Peschanaya Bay, 20 m. 



Euglesa subinflata (Mori, 1938) F 

Pisidium cinereum subinflatum Mori, 1938b: 256, 259, 273-274, pl. 10, fig. 20. 

● Type locality: Siritori, Karahuto. 


● Distribution: lakes of southern Sakhalin (Starobogatov et al., 2004). 

Euglesa subplanata Starobogatov et Budnikova, 1985 [1986] F 





Euglesa subponderosa Krivosheina, 1976 F 

Krivosheina, 1976: 30-31, fig. 2(6). 



● Distribution: Europe, south of western Siberia, Altai (Starobogatov et al., 2004).

Euglesa subterranea (Shadin, 1932) 

Pisidium subterraneum Shadin, 1932: 13, pl. 1, fig. 2. 

● Type locality: Rioni Cave near Kutaisi, Georgia. 


● Distribution: caves of Transcaucasia (Shadin, 1952). 

Euglesa turkestanica Izzatullaev, 1974 F 

Izzatullaev, 1974: 1086-1087, textfig. (1). 

● Type locality: floodland of Surkhob River near Alaka kishlak, vicinities of Soviet farm "Muminabad" (Tajikistan). 


● Distribution: Central Asia (Izzatullaev, 2002) 

● Remark. Korniushin (1996) classified this species in the genus Roseana. 

Euglesa zugmayeri (Weber, 1910) F 

Pisidium zugmayeri Weber, 1910: 310. 

● Type locality: Kashmir (Tibet after Korniushin, 1996). 


● Distribution: central Asia (Izzatullaev, 2002) 

Henslowiana Fagot, 1892 

Type species: Pisidium henslowianum Fagot, 1892 [= Tellina henslowana Leach in Sheppard, 1823] (SD 

Boettger, 1961) 

Henslowiana baudonii (Clessin, 1873) F 

Pisidium baudonii Clessin, 1873a: 83-85 (nom. nov. pro Pisidium henslowanum sensu Baudon, 1857, non 

Leach in Sheppard, 1823) 

● Type locality: "Mouy (Oise)". 


● Distribution: rivers of Europe (Starobogatov et al., 2004). 

Henslowiana chankensis (Likharev in Shadin, 1952) F 

Pisidium (Eupisidium) chankense Likharev – Shadin, 1952: 331, fig. 300. 

● Type locality: Khanka Lake, 2-5 m. 

● Lectotype (Prozorova et al., 1996): ZIN. 

● Distribution: Khanka Lake (Starobogatov et al., 2004). 

Henslowiana conica (Baudon, 1857) F 

Pisidium conicum Baudon, 1857: 50-52, pl. 5, fig. B. 

● Type locality: "de Valenciennes et de Troyes". 


● Distribution: rivers of Baltic Sea basin, Ukraine, central Asia (Stadnichenko, 1984; Izzatullaev, 2002; Starobogatov 

et al., 2004). 

Henslowiana corbiculaeformis Korniushin, Moskvicheva et Starobogatov 

in Prozorova, Starobogatov et Korniushin, 1996 F 

Henslowiana corbiculaeformis Korniushin, Moskvicheva et Starobogatov – Prozorova et al., 1996: 1323, 

figs. 1 б (b), 2 (2). 

● Type locality: not specified; type series from: Amur liman and lower reaches of Amur River; Ussuri River 

near Lesozavodsk town. 


● Distribution: Ussuri River and, probably, middle and lower Amur basin (Starobogatov et al., 2004).

Henslowiana costifera Korniushin et Starobogatov in Prozorova, Starobogatov et Korniushin, 1996 F 

Henslowiana costifera Korniushin et Starobogatov – Prozorova et al., 1996: 1323-1324, figs. 1 г (g), 2 (4). 

● Type locality: specified; type series from: Amur liman and lower reaches of Amur River; Ussuri River near 

Lesozavodsk town. 


● Distribution: middle and lower Amur basin including Khanka Lake (Starobogatov et al., 2004). 

Henslowiana czerskii (Starobogatov et Streletzkaja, 1967) F 

[= Galileja henslowana karetovensis Starobogatov et Streletskaja, 1967] 

Galileja czerskii Starobogatov, Streletzkaja, 1967: 248. 

● Type locality: channel of the Kolyma River near Nizhnie Kresty. 


● Distribution: eastern Siberia, Baikal Lake, Kolyma River basin (Slugina et al., 1994; Starobogatov et al., 

2004; Slugina, Starobogatov, 2004). 

Henslowiana elegantula (Starobogatov et Budnikova, 1985 [1986]) F 

Euglesa (Costopisidium) elegantula Starobogatov, Budnikova, 1985 [1986]: 105. fig. 9 

● Type locality: Lagunnoe Lake, Kunashir (Kurile Islands). 


● Distribution: southern Kurile Islands (Kunashir Island) (Prozorova, 1996 [1997]a; Starobogatov et al., 

2004). 

Henslowiana dupuyana (Normand, 1854) F 

Pisidium dupuyanum Normand, 1854: 5. 

● Type locality: "Valenciennes". 


● Distribution: Europe, rivers and permanent waterbodies of the river plains (Stadnichenko, 1984; Starobogatov 

et al., 2004) , Western Siberia (Korniushin, 1996). 

Henslowiana falsicorbula Prozorova in Prozorova, Starobogatov et Korniushin, 1996 F 

Henslowiana falsicorbula Prozorova – Prozorova et al., 1996: 1322-1323, figs. 1 a, 2 (1). 

● Type locality: Ussuri River near Lesozavodsk town. 


● Distribution: Ussuri River and, probably, middle and lower Amur basin (Starobogatov et al., 2004). 

Henslowiana formosa (Prozorova, 1988) F 

Euglesa (Arctoeuglesa) formosa Prozorova, 1988a: 1576-1577, fig. 1 г, ä, е (g, d, e). 

● Type locality: small but deep floodland-thermokarst lake in 20-km vicinity of Chaun station of the Institute 

of Biological Problems of the North, Far Eastern Branch of the USSR Academy of Sciences. 


● Distribution: lakes of Chukotka (Chaun lowland) (Starobogatov et al., 2004). 

Henslowiana glaciata (Prozorova, 1988) F 

Euglesa (Arctoeuglesa) glaciata Prozorova, 1988a: 1577, fig. 2 а, б, в (a, b, v). 




● Distribution: lakes of Chukotka (Chaun lowland) (Starobogatov et al., 2004). 

Henslowiana henslowana (Leach in Sheppard, 1823) F 

Tellina henslowana Leach – Sheppard, 1823: 150-151. 

● Type locality: stream at Holbrook. 


● Distribution: rivers of Europe and Siberia, excluding Kamchatka (Stadnichenko, 1984; Starobogatov et al., 

2004). 

Henslowiana infirmicostata (Pirogov et Starobogatov, 1974) F 

Euglesa (Costopisidium) infirmicostata Pirogov et Starobogatov, 1974: 330, fig. 2, 5. 

● Type locality: the Volga River avandelta, offshore, Chernevoj Ochirkin Island. 


● Distribution: Volga River delta, Teterev river [Ukraine] (Stadnichenko, 1984; Starobogatov et al., 2004) 

Henslowiana izzatullaevi (Zatrawkin, 1987) F 

Euglesa (Arcteuglesa) izzatullaevi Zatrawkin, 1987: 52-53, textfig. (IV). 

● Type locality: first floodland lake of Bureya River, left bank near Chekunda settlement (old settlement), 

Verkhnebureinsky District, Khabarovsk Territory. 


● Distribution: lakes of lower Amur basin (Starobogatov et al., 2004). 

Henslowiana kunasiriensis (Mori, 1938) F 

Pisidium pulchellum kunasiriense Mori, 1938b: 255, 259, 269, pl. 9, fig. 11a-d. 

● Type locality: Kunasiri-Zima. 


● Distribution: lakes on Kunashir Island (Kurile Islands) (Starobogatov et al., 2004). 

Henslowiana kurilica (Starobogatov et Budnikova, 1985 [1986]) F 

Euglesa (Costopisidium) kurilica Starobogatov, Budnikova, 1985 [1986]: 102-104, fig. 7. 



● Distribution: lakes of southern Kurile Islands (Prozorova, 1996 [1997]a; Starobogatov et al., 2004). 

Henslowiana kurodai (Mori, 1938) F 

Pisidium kurodai Mori, 1938b: 255, 258, 262, pl. 7, fig. 4. 

● Type locality: Toro-Numa, Etorohu-Zima. 


● Distribution: lakes of southern Kurile Islands (Kunashir and Iturup) (Prozorova, 1996 [1997]a; Starobogatov 

et al., 2004). 

Henslowiana lilljeborgii (Clessin in Esmark et Hoyer, 1886) F 

Pisidium lilljeborgii Clessin – Esmark, Hoyer, 1886: 119-120. 

● Type locality: "Gеsö bei Grötö, Skotstinder bei Grötö, Trondenaes, Mеlselvdalen, Tromsö, Jarfjord". 


● Distribution: lakes of northern Europe (Starobogatov et al., 2004); Armenia (Akramowski, 1976); North 

Kurile Islands (Kuiper, 1998). 

Henslowiana morii (Starobogatov et Budnikova, 1985 [1986]) F 

Euglesa (Costopisidium) morii Starobogatov, Budnikova, 1985 [1986]: 104-105, fig. 8. 



● Distribution: lakes of southern Kurile Islands (Kunashir Island) (Prozorova, 1996 [1997]a; Starobogatov et 

al., 2004). 

Henslowiana nordenskioldi (Clessin in Westerlund, 1876) F 

Pisidium nordenskioldi Clessin – Westerlund, 1876a: 68-69, pl., fig. 20. 

● Type locality: "Jenissei, vid Dudino, (lat. 69º15'), Surgutskoj (lat. 62º50'), Worogowa Selo (lat. 69º50')". 


● Distribution: north of Eastern Siberia (Korniushin, 1996). 

● Remark: see remark to Sphaerium levinodis. 

Henslowiana ostroumovi (Pirogov et Starobogatov, 1974) F 

Euglesa (Henslowiana) ostroumovi Pirogov, Starobogatov, 1974: 332-333, fig. 2 12. 

● Type locality: Kamyzjak channel [the Volga delta], offshore, near Kirovskoj settlement, Astrakhan' Region, 

depth 1.5 m. 


● Distribution: rivers and lakes of Europe and south of western Siberia (Stadnichenko, 1984; Starobogatov et 

al., 2004). 

Henslowiana polonica (Anistratenko et Starobogatov, 1990 [1991]) F 

Euglesa (Hensloviana) polonica Anistratenko, Starobogatov, 1990 [1991]: 18, fig. 4. 

● Type locality: Poland, surroundings of Poznan, powiat Sztem-Mastovo, spring. 


● Distribution: Europe and Siberia to Kamchatka, rivers and lakes (Starobogatov et al., 2004). 

Henslowiana ruut (Timm, 1975) F 

Euglesa ruut Timm, 1975: 225, 227, fig. 9. 

● Type locality: Lake Vörtsjärv. 

● Holotype: Vörtsjärv Limnological Station. 

● Distribution: northern Europe, Ukraine, Ural River (Stadnichenko, 1984; Pirogov et al., 1994; Starobogatov 

et al., 2004). 

Henslowiana securidens (Krivosheina, 1978) F 

Euglesa (Henslowiana) securidens Krivosheina, 1978: 1495, textfig. (10). 

● Type locality: Bukhtarma River mouth, mountain-valley part of Bukhtarma Water Reservoir (upper Irtysh 

basin, East Kazakhstan). 


● Distribution: lakes and small rivers of eastern Kazakhstan (Irtysh basin) (Starobogatov et al., 2004). 

Henslowiana semenkevitschi (Lindholm, 1909) F 

Pisidium nordenskioeldi var. semenkevitschi Lindholm, 1909: 86, Taf. 2, fig. 49-50. 

● Type locality: Baikal, Angarskyi sor. 



Henslowiana sibirica (Clessin in Westerlund, 1876) F 

Pisidium sibiricum Clessin – Westerlund, 1876a: 69, fig. 21. 

● Type locality: "Jenissei, Worogowa Selo (lat. 60º50')". 


● Distribution: lakes of north of western Siberia (Starobogatov et al., 2004). 


Henslowiana subhenslowana (Krivosheina, 1978) F 

Euglesa (Henslowiana) subhenslowana Krivosheina, 1978: 1495-1496, textfig. (11). 

● Type locality: lake part (Kurchum area) of Bukhtarma Water Reservoir (upper Irtysh basin, East Kazakhstan). 


● Distribution: lakes and small rivers of eastern Kazakhstan (Irtysh basin) (Starobogatov et al., 2004). 

Henslowiana suecica (Clessin in Westerlund, 1873) F 

Pisidium henslowanum β suecicum Clessin – Westerlund, 1873: 536-537.

● Type locality: “vid Göteborg I Qvilleebäcken”. 


● Distribution: lakes and rivers of Europe and Siberia, excluding Kamchatka (Stadnichenko, 1984; Starobogatov 

et al., 2004). 

Henslowiana supina (A. Schmidt, 1850) F 

Pisidium supinum Schmidt, 1850: 119. 

● Type locality: "Naumburg a.d.S., Panke bei Berlin". 


● Distribution: rivers of Europe, Ural River (Stadnichenko, 1984; Pirogov et al., 1994; Starobogatov et al., 

2004). 

Henslowiana tenuicostulata (Krivosheina, 1978) F 

Euglesa (Cymatocyclas) tenuicostulata Krivosheina, 1978: 1498, textfig. (14). 

● Type locality: Volchii Cape, lake part of Bukhtarma Water Reservoir (upper Irtysh basin, East Kazakhstan). 


● Distribution: lakes of Europe and Altai (Starobogatov et al., 2004). 

Henslowiana trigonoides (W. Dybowski, 1902) F 

[= Pisidium korotnevi part. Kozhov, 1936] 

Pisidium trigonoides W. Dybowski, 1902: 95. 




Henslowiana uenoi (Mori, 1938) F 

Pisidium uenoi Mori, 1938b: 258, 265, pl. 8, fig. 8. 

● Type locality: Etorohu-Zima (Iturup Island). 


● Distribution: South Kurile Islands (Iturup) (Prozorova, 1996 [1997]a; Starobogatov et al., 2004). 

Henslowiana waldeni (Kuiper, 1975) F 

Pisidium waldeni Kuiper, 1975: 31-36, textfigs. 13, 15-17, 19, 21-26, 30. 

● Type locality: "Buetjörni (= Davggeluobal), 147 m NN, Flachsee (tiefste Stelle 9 m), Varangerhalbiusel im 

NO Norwegens". 


● Distribution: north of Europe and of Siberia, Ukraine, in lakes (Stadnichenko, 1984; Starobogatov et al., 

2004). 

Hiberneuglesa Starobogatov in Dolgin, 1983 

Type species: Pisidium hibernicum Westerlund, 1894 (OD) 

Hiberneuglesa anjuica (Starobogatov et Korniushin, 1989) F 

Euglesa (Hiberneuglesa) anjuica Starobogatov, Korniushin, 1989: 16. fig. 2,3, 3, u [e] 

● Type locality: Quaternary deposits of the basin of the river Malyj Anjuj, 28 km from Anjujsk settlement 

[Chukchi Peninsula]. 


● Distribution: lakes and rivers of Europe (Starobogatov et al., 2004). 

Hiberneuglesa bodamica (Starobogatov et Korniushin, 1989) F 

Euglesa (Hiberneuglesa) bodamica Starobogatov, Korniushin, 1989: 16, fig. 1, 6, 3 d. 

● Type locality: Bodensee [Lake of Constance] near Honnenborn, Bavaria, Germany.


● Distribution: northern Europe, western Siberia, lakes (Starobogatov et al., 2004). 

Hiberneuglesa hibernica (Westerlund, 1894) F 

Pisidium (Fossarina) hibernicum Westerlund, 1894: 205. 

● Type locality: "Irland bei Gleengariff in Cork". 

● Types: not traced. Possibly, in NMG. 

● Distribution: central part of Dnieper basin, lakes (Stadnichenko, 1984; Starobogatov et al., 2004). However, 

Korniushin (1996) stated that H. hibernica occurs in Scandinavia only. 

Hiberneuglesa kolhymensis (Starobogatov et Streletzkaja, 1967) F 

Odhneripisidium (?) kolhymense Starobogatov, Streletzkaja, 1967: 256-257, fig. 70. 

● Type locality: Quaternary deposits of Malyi Anyui River, 28 km upstream of Anyuisk settlement. 


● Distribution: Kolyma basin (Starobogatov et al., 2004). 

Hiberneuglesa kuiperi (Starobogatov et Streletzkaja, 1967) F 

Galileja kuiperi Starobogatov, Streletzkaja, 1967: 251, 253, fig. 61 

● Type locality: lake 8 km northwestward from Achajvajam (continental part of the Kamchatka Region). 


● Distribution: lakes of Koryak upland (Starobogatov et al., 2004). 

Hiberneuglesa lindholmi (Starobogatov et Streletzkaja, 1967) F 

Odhneripisidium lindholmi Starobogatov, Streletzkaja, 1967: 256, fig. 69. 

● Type locality: Quaternary deposits on left bank of the Allaikha River (Indigirka River basin). 


● Distribution: Kolyma basin (Starobogatov et al., 2004). 

Hiberneuglesa normalis (Stelfox, 1929) F 

Pisidium hibernicum var. normalis Stelfox, 1929: 9. 

● Type locality: "Windley Creek, Sutton Coldfield, Warwickshire". 


● Distribution: basin of the Baltic Sea, central Dnieper basin, rivers and permanent waterbodies of the rivers 

plains (Starobogatov et al., 2004). 

Hiberneuglesa parvula (Clessin in Westerlund, 1873) F 

Pisidium parvulum Clessin – Westerlund, 1873: 553. 

● Type locality: “I Sverige, I Blekinge vid Roneby ymnig i Langasjö”. 


● Distribution: Europe and western Siberia, rivers and permanent waterbodies of the river plains (Starobogatov 

et al., 2004). 

Hiberneuglesa portentosa (Stelfox in Ellis, 1940) F 

Pisidium hibernicum var. portentosa Stelfox – Ellis, 1940: 77. 

● Type locality: "Co. Down". 

● Holotype: BMNH. 

● Distribution: Europe and western Siberia, rivers and permanent waterbodies of the river plains (Starobogatov 

et al., 2004). 

Hiberneuglesa subhibernica (Starobogatov et Korniushin, 1989) F 

Euglesa (Hibemeuglesa) subhibernica Starobogatov, Korniushin: 1989: 15, fig. 1 4, 3, 6. 

● Type locality: Sweden, Ronneby. 

● Holotype: ZIN.

● Distribution: northern Europe, western Siberia, rivers and permanent waterbodies of the river plains (Starobogatov 

et al., 2004). 

Pseudeupera German, 1913 

Type species: Pisidium landeroini Germain, 1909 (OD) 

Pseudeupera alta (Mori, 1938) F 

Pisidium subtruncatum altum Mori, 1938b: 255, 259, 269, pl. 9, fig. 12. 

● Type locality: Akan-Ko, Hokkaido. 


● Distribution: South Kurile Islands (Prozorova, 1996 [1997]a; Starobogatov et al., 2004). 

Pseudeupera altaica (Krivosheina, 1979) F 

Neopisidium altaicum Krivosheina, 1979: 603-604, textfig. (A). 

● Type locality: Pennoe bog near Leninogorsk City (East Kazakhstan), 2 m. 


● Distribution: lakes and rivers of southern Altai (Starobogatov et al., 2004). 

Pseudeupera arcidens (Krivosheina, 1978) F 

Euglesa (Pseudeupera) arcidens Krivosheina, 1978: 1492-1493, textfig. (6). 

● Type locality: dam of Bobrovka River at Bobrovka village, Glubokovsky District (upper Irtysh basin, East 

Kazakhstan). 


● Distribution: lakes and rivers of Europe, south of western Siberia, Altai (Starobogatov et al., 2004). 

Pseudeupera humerosa (Pirogov et Starobogatov in Timm, 1975) F 

Euglesa humerosa Pirogov et Starobogatov in Timm, 1975: 221, 223. fig. 7. 

● Type locality: delta of the River Volga, near the village of Baranovka. 


● Distribution: rivers and lakes of Europe (Stadnichenko, 1984; Starobogatov et al., 2004). 

Pseudeupera humiliumbo (Krivosheina, 1978) F 

Euglesa (Pseudeupera) humiliumbo Krivosheina, 1978: 1491-1492, textfig. (4). 


Kazakhstan). 


● Distribution: lakes and rivers of Europe, south of western Siberia, Altai (Stadnichenko, 1984; Starobogatov 

et al., 2004). 

Pseudeupera mucronata (Clessin in Westerlund, 1876) F 

Pisidium mucronatum Clessin in Westerlund, 1876a: 71. 

● Type locality: "Jenissei, Dudino (lat. 69º15') samt Nischnij Inbatsk (63º50')". 


● Distribution: rivers and lakes of western and eastern Siberia, Baikal Lake, Kamchatka, Kolyma, 0-5 m 



Pseudeupera pallida (Gassies, 1855) F 

Pisidium pallidum Gassies, 1855: 116, pl. 1, fig. 10. 

● Type locality: "а Pourret et Saint-Marcel près d'Agen, au Sud" (France). 

● Syntypes: MNHN, MCZ No. 19730. 

● Distribution: lakes and rivers of Europe, south of western Siberia, Altai, central Asia (Stadnichenko, 1984; 

Izzatullaev, 2002; Starobogatov et al., 2004).

Pseudeupera parallelodon (Krivosheina, 1978) F 

Euglesa (Pseudeupera) parallelodon Krivosheina, 1978: 1494, textfig. (8). 


Kazakhstan). 


● Distribution: Europe, basin of Irtysh, Altai, lakes (Starobogatov et al., 2004). 

Pseudeupera pirogovi (Starobogatov in Stadnichenko, 1984) F 

[= Euglesa difficilis Pirogov et Starobogatov in Timm, 1975, non Euglesa difficilis Popova et Starobogatov in 

Popova, Devjatkin et Starobogatov, 1970; Euglesa astrahanica Pirogov et Starobogatov in Timm, 1984, nom. 

nov. pro E. difficilis Pirogov et Starobogatov in Timm, 1975, non Popova et Starobogatov in Popova, Devjatkin 

et Starobogatov, 1970] 

Euglesa (Henslowiana) pirogovi Starobogatov – Stadnochenko, 1984: 302, 305, fig. (nom. nov. pro Euglesa 

difficilis Pirogov et Starobogatov in Timm, 1975, non Euglesa difficilis Popova et Starobogatov in Popova, 

Devjatkin et Starobogatov, 1970). 

● Type locality: flood-plains of the delta of the River Volga in the Astrakhan Nature Conservation Area. 

● Holotype: ZIN??. 

● Distribution: lakes and rivers of Europe and southern part of western Siberia (Starobogatov et al., 2004). 

● Remark: two substituting names for Euglesa difficilis Pirogov et Starobogatov in Timm, 1975, non Euglesa 

difficilis Popova et Starobogatov in Popova, Devjatkin et Starobogatov, 1970 were published in 1984. Both 

names were published in books with only year at the title page as the date. Nevertheless, the book of Stadnichenko 

(signed for the printing on the 22.12.1983) seems to appear earlier than that of Timm (signed for the 

printing on the 10.09.1984). Thus Euglesa (Henslowiana) pirogovi is senior objective synonym of Euglesa 

astrahanica. 

Pseudeupera poltavica (Korniushin, 1991) F 

Cingulipisidium (Potamopisidium) poltavicum Korniushin, 1991: 17-19, fig. 1 (5-7). 

● Type locality: Vorskla River, vicinities of Poltava. 


● Distribution: basin of Dnieper, rivers (Starobogatov et al., 2004). 

Pseudeupera rotundotrigona (Krivosheina, 1978) F 

Euglesa (Pseudeupera) rotundotrigona Krivosheina, 1978: 1492, textfig. (5). 


Kazakhstan). 


● Distribution: lakes and rivers of Europe and southern part of western Siberia, Ukraine (Stadnichenko, 1984; 


Pseudeupera starobogatovi (Krivosheina, 1978) F 

Euglesa (Pseudeupera) starobogatovi Krivosheina, 1978: 1491, textfig. (3). 

● Type locality: debris of Ulba River, Ulba Perevalochnaya railway station (East Kazakhstan). 


● Distribution: north of Europe (Starobogatov et al., 2004), south of Western Siberia (Krivosheina, 1978). 

Pseudeupera subalta (Starobogatov et Budnikova, 1985 [1986]) F 

Euglesa (Pseudeupera) subalta Starobogatov, Budnikova, 1985 [1986]: 105-107, fig. 10 



● Distribution: Kunashir Island (Prozorova, 1996 [1997]a; Starobogatov et al., 2004).

Pseudeupera subcuneata (Krivosheina, 1978) F 

Euglesa (Pseudeupera) subcuneata Krivosheina, 1978: 1494-1495, textfig. (9). 


Kazakhstan). 


● Distribution: lakes of Europe, south of western Siberia, Altai (Stadnichenko, 1984; Starobogatov et al., 

2004). 

Pseudeupera subtruncata (Malm, 1855) F 

Pisidium subtruncatum Malm, 1855: 92, fig. 

● Type locality: Sweden: near Qvillebäcken 


● Distribution: Europe and western Siberia, Central Asia, lakes and rivers (Stadnichenko, 1984; Pirogov et 


Pseudeupera supiniformis (Pirogov et Starobogatov, 1974) F 

Euglesa (Cymatocyclas) supiniformis Pirogov et Starobogatov, 1974: 333-334, fig. 3 1. 

● Type locality: Bolshoj Karabulak bayou [the Volga delta], depth 2.5 m. 


● Distribution: the Volga delta (Starobogatov et al., 2004). 

Pseudeupera talievi (Starobogatov et Streletzkaja, 1967) F 

Galileja talievi Starobogatov, Streletzkaja, 1967: 243, fig. 65. 

● Type locality: Malyi Posolskyi Sor, Baikal Lake. 


● Distribution: Baikal and vicinities (Slugina et al., 1994; Starobogatov et al., 2004; Slugina, Starobogatov, 

2004). 

Pseudeupera tenuicardo (Krivosheina, 1978) F 

Euglesa (Pseudeupera) tenuicardo Krivosheina, 1978: 1493-1494, textfig. (7). 


Kazakhstan). 


● Distribution: rivers and lakes of southern Altai (Starobogatov et al., 2004). 

Pseudeupera tenuisculpta (Pirogov et Starobogatov in Timm, 1975) F 

Euglesa tenuisculpta Pirogov et Starobogatov in Timm, 1975: 235, 237, fig. 15. 

● Type locality: delta of the River Volga in a flood-plain in Bolshoi Karabulak. 


● Distribution: eastern Europe, lakes and rivers (Stadnichenko, 1984; Starobogatov et al., 2004). 

Pseudeupera turanica (Clessin in Martens, 1874) F 

Pisidium turanicum Clessin – Martens, 1874: 38-39, pl. 3, fig. 34. 

● Type locality: "in lacu Durmankul vallis Sarafschan". 



● Remark: Kuiper (1989, quoted after Korniushin, 1996) considered this species as a synonym of Pisidium 

obtusale. Korniushin (1996) placed P. turanica in the genus Cyclocalyx. 

Pseudeupera turgida (Clessin in Westerlund, 1873) F 

[= Euglesa incrassata Ellis, 1940] 

Pisidium subtruncatum β turgidum Clessin – Westerlund, 1873: 542.

● Type locality: “Dalarne I Husbysjön vid Säther”. 


● Distribution: rivers and lakes of Europe (Stadnichenko, 1984; Starobogatov et al., 2004). 

Pseudeupera volgensis (Pirogov et Starobogatov in Timm, 1975) F 

Euglesa volgensis Pirogov et Starobogatov in Timm, 1975: 233-235, fig. 13-14. 

● Type locality: delta of the River Volga near the village of Kirovskij. 


● Distrtibution: lakes and rivers of Europe (Stadnichenko, 1984; Starobogatov et al., 2004), Western Siberia 

(Korniushin, 1996). 

Рseudosphaerium Korniushin, 1990 

Type species: Pisidium pseudosphaerium Favre, 1927 (OD) 

Рseudosphaerium favrei (Kuiper, 1947) F 

Pisidium favrei Kuiper, 1947: 35-37. 

● Type locality: "Natuurmonument Naardermeer, Spookgat" (the Netherlands). 

● Holotype: Zoological Museum, Amsterdam, No. 2710a. 

● Distribution: Europe, possibly in the basin of the Baltic Sea, lakes (Starobogatov et al., 2004). 

Рseudosphaerium pseudosphaerium (Favre, 1927) F 

Pisidium milium var. pseudosphaerium Favre, 1927: 298. 

● Type locality: "Schweiz, Bassin de Genève, marais de la Touvière, blättriger Torf". 

● Lectotype (Adler, 1993): MHNG. 

● Distribution: basin of the Baltic Sea and central basin of Dnieper, in small rivers and permanent waterbodies 

on the river plains (Stadnichenko, 1984; Starobogatov et al., 2004). 

Pulchelleuglesa Starobogatov in Dolgin, 1983 

Type species: Pisidium pulchellum Jenyns, 1832 (OD) 

Pulchelleuglesa acuticostata (Starobogatov et Korniushin, 1989) F 

Euglesa (Hiberneuglesa) acuticostata Starobogatov, Korniushin, 1989: 18, fig. 2, 1. 

● Type locality: surroundings of Berlin, Ebersvalde. 


● Distribution: northern and central Europe, western Siberia, lakes (Starobogatov et al., 2004). 

Pulchelleuglesa pulchella (Jenyns, 1832) F 

Pisidium pulchellum Jenyns, 1832: 306, pl. 21, fig. 1. 

● Type locality: “throughout Cambridgeshire and likewise in other part of the country” [England]. 


● Distribution: northern Eurasia, eastward to Kolyma River (Stadnichenko, 1984; Starobogatov et al., 2004). 

Roseana Fagot, 1892 

Type species: Pisidium roseum Scholtz, 1843 (SD Boettger, 1961) 

Roseana alexandri (Stadnichenko, 1980) F 

Euglesa (Roseana) alexandri Stadnichenko, 1980: 33. 

● Type locality: Salgir river, near village Perevalnoe, Crimea. 

● Holotype: Zoological Institute of Ukranian Academy of Sciences. 

● Distribution: Crimea (Stadnichenko, 1984). 


Pisidioidea and its reality should be proven.

Roseana borealis (Clessin in Westerlund, 1876) F 

Pisidium boreale Clessin – Westerlund, 1876a: 70-71, fig. 23. 

● Type locality: "Jenissei, norr om Lusino (lat. 68º40')". 

● Lectotype ( Starobogatov, Streletzkaya, 1967): ZIN, No. 1 in systematic catalog. Korniushin (1996) stated 

that types are kept in SMF. 

● Distribution: lakes of Europe and Siberia, central Asia (Izzatullaev, 2002; Starobogatov et al., 2004). 


Roseana globularis (Clessin in Westerlund, 1873) F 

Pisidium globulare Clessin – Westerlund, 1873: 532-533. 

● Type locality: “I Sverige… I Norge…”. 

● Syntypes: SMF. 

● Distribution: Europe, western Siberia (Stadnichenko, 1984; Pirogov et al., 1994; Starobogatov et al., 2004). 

Roseana rosea (Scholtz, 1843) F 

Pisidium roseum Scholtz, 1843: 140. 

● Type locality: Silesia (fide Westerlund, 1890). 


● Distribution: basin of the Baltic Sea, central Dnieper basin, permanent and semipermanent waterbodies 

(Stadnichenko, 1984; Starobogatov et al., 2004). 

Roseana silesiaca (Starobogatov et Korniushin in Korniushin, 1991) F 

[= ?Pisidium fossarinum Clessin in Westerlund, 1873 sensu Korniushin, 1990; Pisidium acuminatum Clessin 

in Westerlund, 1873 sensu Korniushin, 1990] 

Euglesa (Roseana) silesiaca Starobogatov et Korniushin – Korniushin, 1991: 21, fig. 1 (14-16). 

● Type locality: Krkonoše Mountains, Czechoslovakia. 


● Distribution: small waterbodies of Europe (Starobogatov et al., 2004). 

Tetragonocyclas Pirogov et Starobogatov, 1974 

Type species: Pisidium milium Held, 1836 (OD) 

Tetragonocyclas baudoniana (de Cessac, 1855) F 

Pisidium baudonianum de Cessac, 1855: 74-75. 

● Type locality:"au Mouchetard, dans la pêcherie de la grand prairie du Château". 


● Distribution: lakes of Europe and Siberia (Starobogatov et al., 2004). 

● Remark: Korniushin (1996) spelled the species name as Tetragonocyclas baudonianum. 

Tetragonocyclas milium (Held, 1836) F 

Pisidium milium Held, 1836: 281. 

● Type locality: Bavaria (from the author paper’s title). 


● Distribution: Palearctic (Starobogatov et al., 2004), including Eurasia and North Africa, and North America 

(Piechocki, 1989). 

Tetragonocyclas quadrangula (Starobogatov et Budnikova, 1985 [1986]) F 

Euglesa (Tetragonocyclas) quadrangula Starobogatov, Budnikova, 1985 [1986101, fig. 5. 



● Distribution: Kurile Islands (Prozorova, 1996 [1997]a).

Tetragonocyclas subtetragona (Starobogatov et Budnikova, 1985 [1986]) F 

Euglesa (Tetragonocyclas) subtetragona Starobogatov, Budnikova, 1985 [1986]: 99-100, fig. 4. 



● Distribution: southern Kurile Islands (Prozorova, 1996 [1997]a; Starobogatov et al., 2004). 

Tetragonocyclas tetragona (Normand, 1854) F 

Pisidium tetragonum Normand, 1854: 5. 

● Type locality: "Valenciennes, ... dans la fôret de Raismes" (France).. 


● Distribution: Europe, western Siberia, lakes and river creeks (Starobogatov et al., 2004). 

● Remark: Piechocki (1989) considered this species to be a synonym of T. milium (Held). Korniushin (1996) 

spelled the species name as Tetragonocyclas tetragonum. 

LACUSTRININAE Korniushin, 1989 

Lacustrina Sterki, 1916 

Type species: Pisidium idahoense Roper, 1890 (OD) 

Lacustrina chukchensis (Prozorova, 1988) F 

Pisidium chukchensis Prozorova, 1988a: 1576, fig 1 а-в (a, b, v). 




● Distribution: Chukotka, Kamchatka (Starobogatov et al., 2004). 

Lacustrina dilatata (Westerlund, 1897) F 

[= Pisidium maculatum W. Dybowski, 1902; Pisidium subtilestriatum Lindholm, 1909; Pisidium carelicum 

Odhner, 1926] 

Pisidium (Fossarina) dilatatum Westerlund, 1897b: 131. 

● Type locality: "Sibirien. Olenek, oberhalb der Alahis-Mündung". 


● Distribution: rivers and lakes of northern Eurasia (except south of West Siberia), Baikal Lake, 2-50 m 


Lacustrina etorohuense (Mori, 1935) F 

Pisidium amnicum etorohuense Mori, 1935: 194-195, fig. 1. 

● Type locality: "Toro-Numa (Etorohu Island, South Kurile Group; type locality); Syana-Numa (Etorohu), 

Sibetori-Ko (Etorohu)". 


● Distribution: Iturup (Kurile Islands) (Zatrawkin, 1985a; Prozorova, 1996 [1997]a). 

● Remark: the types were not designated or mentioned in the original publication. 

Order MYOIDA 

CORBULIDAE Lamarck, 1818 

Potamocorbula Habe, 1955 

Type species: Corbula amurensis Schrenck, 1862 (OD)

Potamocorbula amurensis (Schrenck, 1862) B 

[= Corbula amplexa Adams, 1862; Corbula frequens Yokoyama, 1922; Corbula pustulosa Yokoyama, 1922; 

Corbula sematensis Yokoyama, 1922; Corbula vladivostokensis Bartsch, 1922] 

Corbula amurensis Schrenck, 1862: 412. 

● Type locality: Chikhacheva Bay [Tatar Strait, northern Japan Sea]. 


● Distribution: river estuaries of Japan and southern Okhotsk seas (Peter the Great Bay, Chikhacheva Bay, 

Amur estuary, Tugur Bay), brackish water lake Lebyazhye on southern Sakhalin; 1-13 m (valves down to 

1600 m) (Scarlato, 1981; Zatrawkin, Bogatov, 1987).

APPENDIX 

Species described from the considered territory but never mentioned in recent literature: 

VIVIPARIDAE 

Paludina decollata Siemaschko, 1847 

Siemaschko, 1847: 100-101, pl. 1, fig. 1. 

● Type locality: "Im Gouvern. Petersburg unweit Jamburg auf dem Gute Zabalkanskaja". 



● Remark: this species is most probably a synonym of one of the numerous viviparids from the Northeast 

Europe, but it has not been mentioned in recent literature, even as a synonym. 

ACROLOXIDAE 

Ancylastrum ovatum Clessin, 1907 

Clessin, 1907: 7. 


● Types: lost as the Clessin’s collection was destroyed during World War II (P. Glöer, pers. communication). 

● Distribution: type locality (Clessin, 1907). 

Ancylastrum issykulense Clessin, 1907 

Clessin, 1907: 7. 




Ancylastrum turkestanicum Clessin, 1907 

Clessin, 1907: 7. 




Ancylastrum dextrorsum Clessin, 1907 

Clessin, 1907: 7. 




PLANORBIDAE 

Anisus starobogatovi Frolova, 1984 

Frolova, 1984: 47, textfig. 

● Type locality: “the floodplain of the Ishim River in 18 kilometers from Petropavlovsk” 



● Remark. Vinarski et al. (2006) suggested that the species Choanomphalus rossmaessleri (Auerswald in A. 

Schmidt, 1851) was described under the name Anisus starobogatovi, however, there is no possibility to check 

this opinion due to loss of the type materials.

Gyraulus acutus Clessin, 1907 

Clessin, 1907: 7-8. 




● Remark. Shadin (1952) considered Gyraulus acutus as a junior synonym of Gyraulus gredleri (Bielz) [= 

Anisus acronicus, partim]. 

Planorbis moellendorffii W. Dybowski, 1904 

W. Dybowski, 1904: 46-47, fig. 3a-c. 

● Type locality: "Natschiki" (Kamchatka). 


● Distribution: type locality (Dybowski, 1904). 

SUBULITIDAE 

Stenogyra retteri Rosen in Kobelt, 1904 T 

Kobelt, 1904: 87. 

● Type locality: "Aulié-ata im Syr-Darja Gebiet". 



LYMNAEIDAE 

Lynmaea karpinskii (Siemaschko, 1847) F 

Lynmaeus karpinskii Siemaschko, 1847: 103. 

● Type locality: "bei Jamburg, beim Gute Zabalkanskaja". 



● Remark: this species is most probably a synonym of one of the numerous lynmaeids from the Northeast 

Europe, but it has not been mentioned in recent literature, even as a synonym. 

PHYSIDAE 

Physa sartlandinensis Mozley, 1934 F 

Mozley, 1934: 7, pl. 1, fig. 6. 

● Type locality: Lake Sartlan, Brabinsk Steppe, Siberia. 

● Holotype: USNM No. 469613. 

● Distribution: type locality (Mozley, 1934). 

● Remark: Taylor (2003) considers this species to be synonymous with Physa dalmatina Küster, 1844. 

VALVATIDAE 

Valvata antiquilina Mozley, 1934 F 

Mozley, 1934: 1-2, pl. 1, fig. 4. 

● Type locality: Lake Khomotenoye, Aj-Bulat drainage basin, Siberia, approximately 370 km southeast of 

Omsk (= Khomutinoe Lake, Burla Lake system, Altay). 


● Distribution: type locality (Mozley, 1934). 

● Remark: the holotype was designated in the original publication as “type”. The lot in the type collection of 

USNM with the catalogue number 469212 contains single specimen, which matches the original illustration 

and measurements. Nevertheless the specimen is labeled as syntype on the later printed pabel.

UNIONIDAE 

Unio annulatus Kobelt, 1912 

Unio annulatus Kobelt, 1912: 28, Taf. XDII, fig. 2628. 

[= Unio kalmykorum Bogatschev, 1924] 

● Type locality: Ilek River, a tributary of the Ural River (Orenburg Region). 

● Types: Not traced, possibly in SMF. 

● Distribution: European Russia. 

● Remark: This species (under the name Unio kalmykorum) was mentioned by Zatravkin (1983: 41) in his list of 

unionid mussels of USSR (see also Zatravkin, 1983a). Later, this species has disappeared from catalogues (see 

Starobogatov et al., 2004; Kantor, Sysoev, 2005). Zhadin (1938: 78) and Haas (1940: 122) regarded it as a 

varitety of U. pictorum (L.), however, its true identity is still not established. 

Unio stepanoffi Drouët, 1881 

Drouët, 1881: 15. 

● Type locality: "la Crimée". 


● Distribution: Crimea (Drouët, 1881). 

Anodonta middendorffii Siemaschko, 1849 

Siemaschko, 1849: 237, figs. 2-3. 

● Type locality: l'Oka près d'Orol (Siem. – after Drouët, 1881). 


● Distribution: Oka, Bug, Orel, Dnieper rivers (Drouët, 1881). 

Anodonta parmata Drouët, 1881 

Drouët, 1881: 26-27. 

● Type locality: "la Russie méridionale". 


● Distribution: type locality (Drouët, 1881). 

Anodonta georgiana Drouët, 1881 

Drouët, 1881: 28-29. 

● Type locality: "le Caucase". 



Anodonta lenkoranensis Drouët, 1881 

Drouët, 1881: 30. 

● Type locality: "le Lenkoran, а Lenkoran ...; le Kour". 



? DREISSENIDAE 

Mytilus albus Siemaschko, 1847 

Siemaschko, 1847: 124. 

● Type locality: "In Bug". 



● Remark: this is most probably a synonym of a species of Dreissena, possibly a forgotten senior synonym of 

D. bugensis.

PISIDIIDAE s.l. 

Pisidium miliolum Clessin, 1907 

Clessin, 1907: 8. 




Pisidium schmidti Clessin, 1907 

Clessin, 1907: 8. 



● Distribution: type locality (Clessin, 1907).

catalogue of the continental mollusks of russia and - Ruthenica ...

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