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il ' ii - Northern Research Station - USDA Forest Service

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causing visible deformation, a habit apparently much Baldratiaa---two of the largest genera of Lasiopterinimore<br />

widespread in the Lasiopterini than previously were described, among other taxa, in Mrhn's revision.<br />

thought. Thepresence of the midges in such cases is During my research, I reared 26 species belonging to<br />

often discovered only after exuviae are found protruding these genera, some of them from hosts recorded by<br />

from normal looking leaves or stems (Dorchin 1997; Mrhn. A study of Mrhn's collection was therefore<br />

Fedotova 1992a, b, 1993). Larvae may be solitary or essential. Unfortunately, the present condition of this A<br />

gregarious, and when gregarious, may cause considerable collection, which is deposited in the Staatliches Museum •<br />

damage to shoots (Dorchin 1997) or to fruits and seeds fOrNaturkunde in Stuttgart, renders it impossible to<br />

(Fedotova 1991, 1992b). locate and study most of the type material, as the<br />

microscope slides are st<strong>il</strong>l temporary, with glycerol as h<br />

The great majority of the chenopod infesting Lasiopterini mounting medium, and the types are actually not labeled:<br />

complete their life cycle inside their galls, but some The problem is not limited to Baldratia and Stefaniola<br />

exceptions are known for which the mature larvae fall to alone, but also concerns the genera Ozirhincus and<br />

the ground and pupate in it (e.g., Stefanio!a ustjurensis Lasioptera which were revised in the same work, as well<br />

Fedotova (1991) and Baldratiajaxartica Fedotova as the Palaearctic species of the tribe Trotter<strong>ii</strong>ni. Collect-<br />

(1992b)). It was previously thought that most species ing the relevant galls all over again and obtaining from<br />

, have only one generation per year (MOhn 1971). How- them larvae as well as associated adults seems to be the<br />

ever, this impression might be the result of insufficient best solution to this problem. Dr. Fedotova, now at the<br />

biologicaland ecological data, as many species recently Agricultural Academy in Ust Kinelskij, Russia, has<br />

studied by Fedotova (1991, 1992b, 1993, 1993a, 1995) recently described the adults of few of Mrhn's species<br />

proved to have two or even three generations per year, from Kazakhstan (Fedotova 1989a, b, c, 1992) and I have<br />

and thisis also implied by partial data I have obtained on undertaken this task in Israel. However, further surveys<br />

the Israeli fauna (Dorchin 1997). throughout north Africa, the Middle East, and central<br />

Asia are necessary, although at this time seem somewhat<br />

An interesting ecological adaptation found in my study unrealistic.<br />

reveals that despite their delicacy and frag<strong>il</strong>ity, some<br />

Israeli desert species are active throughout the hottest Another difficulty is created by the inaccessib<strong>il</strong>ity of<br />

months 0fthe year, when temperatures may reach 45°C many relevant types of species described by Russian<br />

(--113°F). Thus, Cecidomy<strong>ii</strong>dae are an exception to most workers. Conditions presently preva<strong>il</strong>ing in Russia,<br />

Nematocera, which prefer cool and moist biotopes, and especially communication problems, form a great<br />

are probably t-helargest and most diverse nematoceran hindrance to any study relying on the ava<strong>il</strong>ab<strong>il</strong>ity of these<br />

fam<strong>il</strong>y to be found in desert areas, types.<br />

Taxonomy and Taxonomic Problems LITERATURE CITED<br />

in view of the potential in the Lasiopterini for many new Dorchin, N. 1997. Gall midges infesting Chenopodiaceae<br />

and interesting taxonomic discoveries, it is important to (Diptera: Cecidomy<strong>ii</strong>dae) in Israel. Tel Aviv, Israel: Tel<br />

•mention that the tribe exhibits several complicated Aviv University. 131 p. M.Sc. thesis. (In Hebrew)<br />

taxonomic problems and is in urgent need of comprehen-<br />

• sive taxonomic revision and phylogenetic analysis. Fedotova, Z.A. 1989a. Gall midges of the genus<br />

However, these tasks confront some serious, technical Stefaniola in Kazakhstan. communication 1.<br />

difficulties. The first difficulty is related to the revision Zoologicheskiy Zhurnal. 68(5): 59-71. (In Russian)<br />

of the Palaearctic Lasiopterini, published by Dr. Edwin !<br />

. Mrhn between 1966 and 1971 in the series Die Fliegen Fedotova, Z.A. 1989b. Gall midges of the genus<br />

der Palaearktischen Region. This revision, including the Stefaniola in Kazakhstan USSR. communication 2. w<br />

. keys and description of 72 new species, is entirely based Zoologicheskiy Zhurnal. 68(6): 78-86. (In Russian)<br />

on larvae recovered from galls found in dried plants,<br />

most of which were chenopods collected years earlier in Fedotova, Z.A. 1989c. Gall midges of the genus<br />

north Africa, central Asia, and the Mediterranean basin. Stefaniola in Kazakhstan USSR. communication 3.<br />

The identification of cecidomy<strong>ii</strong>d larvae is extremely Zoologicheskiy Zhumal. 68(8): 57-66. (In Russian)<br />

difficult; and the usefulness of any revision based on<br />

larval Characters alone is therefore rather limited. Fedotova, Z.A. 1990. New species of gall midges<br />

(Diptera, Cecidomy<strong>ii</strong>dae) found on pastures with<br />

Another complication concerns Mrhn's types. Thirty- Anabasis spp. and Nanophyton erinaceum<br />

five new species of Stefaniola and 19 new species of (Chenopodiaceae) in Kazakhstan. Entomologicheskoye<br />

Obozreniye. 69: 223-232. (In Russian)<br />

,<br />

20

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