Great Lakes Dairy Sheep Symposium - the Department of Animal ...
Great Lakes Dairy Sheep Symposium - the Department of Animal ... Great Lakes Dairy Sheep Symposium - the Department of Animal ...
tendency is more marked the higher the number of botanical species in the pasture, on condition that botanical heterogeneity is accompanied by a corresponding spatial differentiation of the various essences. In fact, structural differentiation of pasture facilitates identification and prehension of the most appetising essences or parts of the plant. Species which are erect in their growth, such as most forage grasses, offer a better chance of the sheep selecting morphological parts of the single plant than do creeping species, such as some spontaneous or cultivated legumes (Table 7) (Avondo and Lutri, 2004). Table 7 – Protein and digestible organic matter content of the whole plant and of selected parts of various pasture essences (Avondo and Lutri, 2004). Digestible organic Crude Protein matter % DM % DM Whole Selected Whole Selected Botanical essence Grasses Growth habit Plant part plant part Barley Erect 18.2 25.4 68.1 72.0 Bromus spp Erect 15.8 23.7 69.1 73.7 Mixed pasture Erect 10.1 16.8 62.6 69.8 Mixed pasture Erect 8.8 11.3 73.4 76.6 Mixed pasture Legumes Erect 10.8 16.7 - - Vetch Creeping 20.9 22.2 66.4 69.1 Clover Creeping 21.4 26.5 75.1 75.6 Mixed pasture Creeping 16.4 18.0 68.4 68.2 Mixed pasture Creeping 16.8 20.7 - - In general, as biomass increases so does intake capacity. However, even with high herbage availability, as pasture quality gets worse, due to the biological evolution toward maturity, searching for less lignified plant parts or essences results in a reduction in time spent eating. If biomass levels are high and the grass is over 30 cm high, then sheep may trample the grass to reach the tips. As a result some of the biomass is wasted and the sheep spend less time grazing. If, however, the biomass levels are high but the pasture is short and dense, then intake increases because bite prehension is facilitated. Ideally, in order to avoid the negative effects of excessive selective activity by the sheep, the pasture should be dense and not more than 8 -10 cm in height. This can be done by choosing the right type of pasture and maintaining an adequate stocking rate. Herbage quality can be expressed in terms of digestibility, filling value, crude protein (CP) or structural carbohydrate content. It affects pasture intake. Good quality herbage can enable the "potential" intake to be reached. This is an intake that allows the animal to satisfy its requirements without physical limitations. Generally the levels of intake observed in highly digestible pastures are higher than in lignified pastures. It is also true that if the pasture is highly digestible then the 98
animals can reduce intake, as they reach the state of satiety earlier, due to the effects of metabolic control. The correlation between chemical components and intake for the different types of pasture is rather weak. It is difficult to clearly assess the pasture conditions of natural pasture with the type of undifferentiated data on its chemical composition which are available. Mean data does not, indeed, explain all the qualitative differences between essences and certainly does not consider the spatial distribution of these essences, even though they may be of significant importance in selective behaviour. Most attempts to predict pasture intake by sheep based on feed chemical and nutritional composition have emphasised the important role played by NDF content (Lanari et al., 1993), as this influences rumen wall distension. Table 8 reports some intake prediction equations based on NDF content. The correlation is always negative. Table 8 - Regression equations between dry matter intake (g/kg metabolic weight) and pasture NDF content. Category of forage Regression equation All forages Alfalfa hays Miscellaneous hays All forages Grass forages Polyfytes hays Miscellaneous hays I = 107.4 – 0.644 NDF I = 104.6 – 0.488 NDF I = 117.4 – 0.760 NDF I = 134.5 –1.10 NDF I = 95.3 + 6.70 NDF – 0.0668 NDF 2 I = 136.5 – 0.12 NDF I = 96.5 – 0.38 NDF – 0.0000004 NDF 4 Effect of supplementary feeding on herbage intake 99 Macchioni et al., 1990 Macchioni et al., 1990 Macchioni et al., 1990 Reid et al., 1988 Rohweder et al., 1978 Dulphy et al., 1990 Lanari et al., 1993 When feed supplements are used there is nearly always a reduction in pasture intake, because of the substitution effect (S). S is the variation in herbage intake per unit of supplement provided and it can vary greatly, from 0 to 1. In certain conditions it may, indeed, be even lower than 0 or greater than 1. Obviously, the lower the S value, the higher the total feed intake. Intake response to a supplement is strongly influenced by the quantity and quality of the available herbage. When there is little biomass available or its quality is poor, the supplement causes an increase in total dry matter intake and an improvement in animal performance; in these cases S is very low or null. In conditions where there is a large amount of available biomass or its quality is high, the efficacy of supplement is almost nullified due to the high S level. Figure 3 shows the S values found by Molle et al. (1997), when a corn grain supplement was given to dairy sheep grazing for 5 hours/d on good quality ryegrass sward of different height and biomass. One can see S values higher than 1. When biomass reaches DM yields higher than 2 t/ha, all the herbage was substituted by concentrates.
- Page 57 and 58: show that milk production increases
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tendency is more marked <strong>the</strong> higher <strong>the</strong> number <strong>of</strong> botanical species in <strong>the</strong> pasture, on condition<br />
that botanical heterogeneity is accompanied by a corresponding spatial differentiation <strong>of</strong> <strong>the</strong><br />
various essences. In fact, structural differentiation <strong>of</strong> pasture facilitates identification and<br />
prehension <strong>of</strong> <strong>the</strong> most appetising essences or parts <strong>of</strong> <strong>the</strong> plant.<br />
Species which are erect in <strong>the</strong>ir growth, such as most forage grasses, <strong>of</strong>fer a better chance<br />
<strong>of</strong> <strong>the</strong> sheep selecting morphological parts <strong>of</strong> <strong>the</strong> single plant than do creeping species, such as<br />
some spontaneous or cultivated legumes (Table 7) (Avondo and Lutri, 2004).<br />
Table 7 – Protein and digestible organic matter content <strong>of</strong> <strong>the</strong> whole plant and <strong>of</strong> selected<br />
parts <strong>of</strong> various pasture essences (Avondo and Lutri, 2004).<br />
Digestible organic<br />
Crude Protein<br />
matter<br />
% DM<br />
% DM<br />
Whole Selected Whole Selected<br />
Botanical essence<br />
Grasses<br />
Growth habit Plant part plant part<br />
Barley<br />
Erect 18.2 25.4 68.1 72.0<br />
Bromus spp<br />
Erect 15.8 23.7 69.1 73.7<br />
Mixed pasture Erect 10.1 16.8 62.6 69.8<br />
Mixed pasture Erect 8.8 11.3 73.4 76.6<br />
Mixed pasture<br />
Legumes<br />
Erect 10.8 16.7 - -<br />
Vetch<br />
Creeping 20.9 22.2 66.4 69.1<br />
Clover<br />
Creeping 21.4 26.5 75.1 75.6<br />
Mixed pasture Creeping 16.4 18.0 68.4 68.2<br />
Mixed pasture Creeping 16.8 20.7 - -<br />
In general, as biomass increases so does intake capacity. However, even with high herbage<br />
availability, as pasture quality gets worse, due to <strong>the</strong> biological evolution toward maturity,<br />
searching for less lignified plant parts or essences results in a reduction in time spent eating.<br />
If biomass levels are high and <strong>the</strong> grass is over 30 cm high, <strong>the</strong>n sheep may trample <strong>the</strong><br />
grass to reach <strong>the</strong> tips. As a result some <strong>of</strong> <strong>the</strong> biomass is wasted and <strong>the</strong> sheep spend less time<br />
grazing. If, however, <strong>the</strong> biomass levels are high but <strong>the</strong> pasture is short and dense, <strong>the</strong>n intake<br />
increases because bite prehension is facilitated. Ideally, in order to avoid <strong>the</strong> negative effects <strong>of</strong><br />
excessive selective activity by <strong>the</strong> sheep, <strong>the</strong> pasture should be dense and not more than 8 -10 cm<br />
in height. This can be done by choosing <strong>the</strong> right type <strong>of</strong> pasture and maintaining an adequate<br />
stocking rate.<br />
Herbage quality can be expressed in terms <strong>of</strong> digestibility, filling value, crude protein (CP)<br />
or structural carbohydrate content. It affects pasture intake. Good quality herbage can enable <strong>the</strong><br />
"potential" intake to be reached. This is an intake that allows <strong>the</strong> animal to satisfy its requirements<br />
without physical limitations. Generally <strong>the</strong> levels <strong>of</strong> intake observed in highly digestible pastures<br />
are higher than in lignified pastures. It is also true that if <strong>the</strong> pasture is highly digestible <strong>the</strong>n <strong>the</strong><br />
98