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Water Framework Directive Marine Plants Task Team Tools Paper ...

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fauna and flora and exposed shores are characterised by their limited algal abundance<br />

and wide lichen zone on the upper littoral. However despite exposure appearing to<br />

contribute to the abundance and zonation patterns of algae in the intertidal there is no<br />

significant impact on the levels of species richness. Exposed shores did result in<br />

slightly lower average species richness (but not significantly different to shores of<br />

other exposure ratings), this is more likely to be due to their limited abundance and<br />

therefore harder to locate. There was also little difference in species composition<br />

between shores of varying exposure level.<br />

Shore type refers to the most dominant type of substrate present on the shore, such as<br />

rock platforms, boulders and pebbles. This can contribute quite significantly to the<br />

levels of species richness as certain substrates are more habitable than others and<br />

provide more favourable conditions for attachment. Rock ridges, outcrops and<br />

platforms were shown to have significantly higher species richness than shores<br />

consisting predominantly of boulders, pebbles and vertical rock. This is probably due<br />

to the levels of stability offered by large fixed areas of hard substrate compared with<br />

pebbles and boulders which are less stable and unable to support climax communities<br />

as effectively. The following shore types are given in order of their contribution to the<br />

level of species richness: Rock ridges/outcrops/platforms > Irregular rock and<br />

boulders > steep/vertical rock > pebbles, stones and small rocks > shingle and gravel.<br />

Subhabitat type and number have a similar effect to shore type with the presence of<br />

some subhabitats resulting in higher levels of species richness. Large, wide rockpools<br />

provide very favourable habitats by limiting the effects of desiccation providing a<br />

more tolerable environment than is experienced on open rock. The following<br />

subhabitat types are given in order of their contribution to the level of species<br />

richness: wide shallow/large/deep rockpools > basic rockpools and crevices ><br />

overhangs > caves. Equally with increasing number of subhabitat types there is a<br />

significant increase in the levels of algal species richness recorded as higher<br />

subhabitat diversity results in higher species diversity.

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