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T H E R O O T S OF RELIGION 197

'memeplexes'. As usual with memes, we gain understanding by

going back to the genetic origin of the analogy.

For didactic purposes, I treated genes as though they were

isolated units, acting independently. But of course they are not independent

of one another, and this fact shows itself in two ways.

First, genes are linearly strung along chromosomes, and so tend to

travel through generations in the company of particular other genes

that occupy neighbouring chromosomal loci. We doctors call that

kind of linkage linkage, and I shall say no more about it because

memes don't have chromosomes, alleles or sexual recombination.

The other respect in which genes are not independent is very

different from genetic linkage, and here there is a good memetic

analogy. It concerns embryology which - the fact is often misunderstood

- is completely distinct from genetics. Bodies are not

jigsawed together as mosaics of phenotypic pieces, each one contributed

by a different gene. There is no one-to-one mapping

between genes and units of anatomy or behaviour. Genes

'collaborate' with hundreds of other genes in programming the

developmental processes that culminate in a body, in the same kind

of way as the words of a recipe collaborate in a cookery process

that culminates in a dish. It is not the case that each word of the

recipe corresponds to a different morsel of the dish.

Genes, then, co-operate in cartels to build bodies, and that is one

of the important principles of embryology. It is tempting to say that

natural selection favours cartels of genes in a kind of group

selection between alternative cartels. That is confusion. What really

happens is that the other genes of the gene pool constitute a major

part of the environment in which each gene is selected versus its

alleles. Because each is selected to be successful in the presence of

the others - which are also being selected in a similar way - cartels

of co-operating genes emerge. We have here something more like a

free market than a planned economy. There is a butcher and

a baker, but perhaps a gap in the market for a candlestick maker.

The invisible hand of natural selection fills the gap. That is different

from having a central planner who favours the troika of butcher

+ baker + candlestick maker. The idea of co-operating cartels

assembled by the invisible hand will turn out to be central to our

understanding of religious memes and how they work.

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