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Mind, Body, World- Foundations of Cognitive Science, 2013a

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salesman’s tour increases linearly, the computational effort for finding the shortest<br />

route increases exponentially.<br />

Because <strong>of</strong> its importance and difficulty, a number <strong>of</strong> different approaches<br />

to solving the travelling salesman problem have been explored. These include a<br />

variety <strong>of</strong> numerical optimization algorithms (Bellmore & Nemhauser, 1968). Some<br />

other algorithms, such as simulated annealing, are derived from physical metaphors<br />

(Kirkpatrick, Gelatt, & Vecchi, 1983). Still other approaches are biologically<br />

inspired and include neural networks (Hopfield & Tank, 1985; Siqueira, Steiner,<br />

& Scheer, 2007), genetic algorithms (Braun, 1991; Fogel, 1988), and molecular computers<br />

built using DNA molecules (Lee et al., 2004).<br />

Given the difficulty <strong>of</strong> the travelling salesman problem, it might seem foolish<br />

to suppose that cognitively simple agents are capable <strong>of</strong> solving it. However, evidence<br />

shows that a colony <strong>of</strong> ants is capable <strong>of</strong> solving a version <strong>of</strong> this problem,<br />

which has inspired new algorithms for solving the travelling salesman problem<br />

(Dorigo & Gambardella, 1997)!<br />

One study <strong>of</strong> the Argentine ant Iridomyrmex humilis used a system <strong>of</strong> bridges<br />

to link the colony’s nest to a food supply (Goss et al., 1989). The ants had to choose<br />

between two different routes at two different locations in the network <strong>of</strong> bridges;<br />

some <strong>of</strong> these routes were shorter than others. When food was initially discovered,<br />

ants traversed all <strong>of</strong> the routes with equal likelihood. However, shortly afterwards, a<br />

strong preference emerged: almost all <strong>of</strong> the ants chose the path that produced the<br />

shortest journey between the nest and the food.<br />

The ants’ solution to the travelling salesmen problem involved an interaction<br />

between the world and a basic behaviour: as Iridomyrmex humilis moves, it deposits<br />

a pheromone trail; the potency <strong>of</strong> this trail fades over time. An ant that by chance<br />

chooses the shortest path will add to the pheromone trail at the decision points<br />

sooner than will an ant that has taken a longer route. This means that as other ants<br />

arrive at a decision point they will find a stronger pheromone trail in the shorter<br />

direction, they will be more likely to choose this direction, and they will also add to<br />

the pheromone signal.<br />

Each ant that passes the choice point modifies the following ant’s probability <strong>of</strong><br />

choosing left or right by adding to the pheromone on the chosen path. This positive<br />

feedback system, after initial fluctuation, rapidly leads to one branch being<br />

‘selected.’ (Goss et al., 1989, p. 581)<br />

The ability <strong>of</strong> ants to choose shortest routes does not require a great deal <strong>of</strong> individual<br />

computational power. The solution to the travelling salesman problem emerges<br />

from the actions <strong>of</strong> the ant colony as a whole.<br />

The selection <strong>of</strong> the shortest branch is not the result <strong>of</strong> individual ants comparing<br />

the different lengths <strong>of</strong> each branch, but is instead a collective and self-organizing<br />

Elements <strong>of</strong> Embodied <strong>Cognitive</strong> <strong>Science</strong> 211

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