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6 Wood Discoloration

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204 8 Habitat of <strong>Wood</strong> Fungi<br />

Strands: only rarely on timber in laboratory culture, cream-ochre-dark<br />

brown; fibers to dark brown; no vessels.<br />

Gloeophyllum sepiarium, Yellow-Red Gill Polypore<br />

Fruit body: (Fig. 8.16d, e) annual to perennial, pileate, broadly sessile, dimidiate,<br />

rosette shaped, often imbricate in clusters from a common base or fused<br />

laterally, to 7 cm wide, 12 cm long and 6–8 mm thick, margin slightly wavy;<br />

upper surface when young yellowish brown, then reddish brown and grey to<br />

black when old; scrupose, warted to hispid, finally zonate often differently<br />

colored; hymenophore with straight lamellae (15–20/cm, behind the margin),<br />

edges of lamellae golden brown in active growth, later umber brown, side surface<br />

of lamellae ochre-brown; usually mixed with daedaleoid to sinuous pore<br />

areas (1–2/mm); monstrous fruit bodies in the dark; trimitic; bipolar;<br />

Strands: only rarely on timber in laboratory culture, white-cream; fibers<br />

yellow to brown, no vessels.<br />

Gloeophyllum trabeum, Timber Gill Polypore<br />

Fruit body (Fig. 8.16f, g): annual to perennial, pileate, sessile, imbricate with<br />

several basidiomes from a common base or elongated and fused along wood<br />

cracks, to 3 cm wide, 8 cm long, 8 mm thick; upper surface soft and smooth,<br />

hazelnut to umber brown to grayish when old, weakly zonate to almost azonate,<br />

lighter margin; hymenophore semi-lamellate or labyrinthine to partly poroid<br />

(2–4/mm), rarely lamellate specimens with up to four lamellae/mm along the<br />

margin, ochre to umber brown; monstrous fruit bodies in the dark; dimitic;<br />

bipolar;<br />

Strands: only on timber in laboratory culture, white-beige to yellow-orangegrey<br />

brown, below 1 mm thick; fibers yellow to brown; no vessels.<br />

Significance: predominantly saprobic, G. sepiarium and G. trabeum exceptionally<br />

on living trees; belonging to the strongest brown-rot fungi of coniferous<br />

structural timber; often on stumps; broad moisture optimum (about 40 to<br />

200% u; Table 8.7), on stored timber and on finished timber that is again<br />

moistened, like poles, posts, fences, sleepers and mining timber. The fungi are<br />

the most important destroyers of conifers windows (cf. Fig. 8.17) that had accumulated<br />

moisture due to inappropriate window construction and handling<br />

faults by the user (e.g., injuring of the lacquer layer by nails). For example,<br />

3.5 million (7%) of wooden windows were partly or completely destroyed by<br />

fungi, predominantly by G. abietinum, in Germany between 1955 and 1965<br />

(Seifert 1974). Fungi survive in the sun-warmed and dry window timber due<br />

to their heat and dryness resistance [G. abietinum: 5–7yearssurvivalindry<br />

timber: Theden (1972)]. Fungi cause (by means of substrate mycelium) decay<br />

first only in the wood interior (“interior rot”). The serious brown rot under<br />

the varnish layer is often only recognized if fruit bodies develop. Except on<br />

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