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6 Wood Discoloration

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126 6 <strong>Wood</strong> <strong>Discoloration</strong><br />

Fig.6.3. Blue stain in wood. A Artificial bluing of pine boards by Phoma exigua. B Detail.<br />

C Thick, brown hyphae of P. exigua. D Chlamydospores (photo G. Koch). E Perithecia (A,<br />

B, C, E from Schmidt and Huckfeldt 2005), — 5 cm, --- 5 mm<br />

pits (thin hyphae through the margo) and grow there from cell to cell through<br />

the pits. Because fungi colonize the sapwood tracheids and fibers, components<br />

of the capillary liquid also might be used as nutrients. Although there are<br />

special microhyphae, transpressoria (Fig. 2.5), which can break through the<br />

wood cell wall, probably by physical pressure and/or enzymatic action (Schmid<br />

and Liese 1966; Liese 1970), in most cases the strength properties of wood are<br />

hardly affected. Thus, the occasionally used term “blue rot” is wrong. Some<br />

species however caused some strength loss. Toughness was the property most<br />

seriously affected (Seifert 1999; Schirp et al. 2003b). In most cases, however,<br />

the damage to wood is mainly cosmetic. The damage however affects domestic<br />

and export earnings for the forest industries. For example, Pinus radiata in<br />

New Zealand is highly susceptible to blue stain with an estimated annual loss<br />

in revenue of NZ$ 100 million per year (Thwaites et al. 2004).<br />

Temperature minimum depends on the species, and is between 0 and −3 ◦ C;<br />

the optimum is between 18 and 29 ◦ C and the maximum is between 28 and<br />

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