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Production Practices and Quality Assessment of Food Crops. Vol. 1

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120 S. Pereira-Lorenzo <strong>and</strong> A. M. Ramos-Cabrer<br />

This means there is no close correlation between male sterility <strong>and</strong> quality <strong>of</strong><br />

the fruit but perhaps between male sterility <strong>and</strong> productivity as suggested by Liu<br />

<strong>and</strong> Zhou (1999).<br />

Meiosis in pollen mother cells occurs in the first week <strong>of</strong> June in Italian cultivars,<br />

10 to 15 days before anthesis (Botta et al., 1995). Pollen viability varied<br />

from 81.3 ± 6.1% based on fluorochromatic reaction to 58.2 ± 7.0% on hanging<br />

drops <strong>and</strong> 50.1 ± 4.5% germination on agar media. Pistillate flowers have six to<br />

eight styles whose tips present a hollow at full bloom. The ovary presents seven<br />

(rarely six or eight) carpels. Each flower has 10 to 16 anatropous ovules. Bearing<br />

mono-embryonic seeds (marron type) has been related to the high occurrence <strong>of</strong><br />

anomalies, such as delayed <strong>of</strong> embryo sac differentiation <strong>and</strong> the presence <strong>of</strong> supernumerary<br />

nuclei in the embryo sac (Botta et al., 1995).<br />

3.3. Phenology<br />

Chestnut is a deciduous tree as corresponding to the temperate area where it is<br />

cultivated. Castanea species are generally cold hardy. Castanea dentata <strong>and</strong> C.<br />

mollissima have been reported to survive down to –30 °C (Andersen, 1994).<br />

After chilling requirement, bud break begins at the end <strong>of</strong> March for Asiatic<br />

cultivars, <strong>and</strong> interspecific hybrids (Table 6), being the most precocious, followed<br />

by the European cultivars. Interspecific hybrids maintain their precocity contribution<br />

origin from the Asiatic. However, some hybrid cultivars <strong>of</strong> C. sativa are as<br />

precocious as C. crenata (Table 6) such as Garone Negro. New shoots are very<br />

sensitive to spring frost, <strong>and</strong> this limits the use <strong>of</strong> Asiatic cultivars to orchards located<br />

at altitudes lower than 500 m. However, European cultivars can avoid spring frost<br />

because <strong>of</strong> their late bud break.<br />

Flowering time is highly variable with the local climatic condition (Breisch, 1995).<br />

Asiatic cultivars <strong>and</strong> hybrids produce the male flowering between the end <strong>of</strong> May<br />

<strong>and</strong> the end June, while European cultivars are between mid-June to mid-July.<br />

Table 7 show that the time <strong>of</strong> flowering for Euro-Asiatic hybrids is more precocious<br />

than European cultivars.<br />

Precocity is also maintained at harvest time, on the 1st September for C. crenata<br />

<strong>and</strong> C. mollissima. Most <strong>of</strong> the cultivars <strong>of</strong> C. sativa produces in October <strong>and</strong> 1st<br />

November, but some cultivars are as precocious as Asiatic species such as Temprana<br />

<strong>and</strong> Pilonga which when cultivated in South Spain (Pereira-Lorenzo <strong>and</strong> Ramos-<br />

Cabrer, 2002).<br />

4.1. Origin <strong>of</strong> the cultivars<br />

4. CULTIVARS<br />

As Vavilov (1951) described; growers realised a first domestication selecting the<br />

best nut from wild populations. With the knowledge <strong>of</strong> vegetative propagation as<br />

grafting, a second domestication began which resulted in fixing the best genotypes.

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