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A field guide to mesozoic birds and other winged dinosaurs

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Strong Primitive Archaeopter Protarchaeopteryx robusta<br />

Time: 124.5 Ma ago Location: Liaoning, China Habitat: Lower Yixian Formation (see<br />

above) Size: WS ~65cm (2ft); BL 55cm (1.8ft); TL 65cm (2ft) Features: Head short & high<br />

w/ rounded snout. Teeth numerous (up <strong>to</strong> 18 in both the upper & lower jaws). Upper front<br />

two teeth unusually large & chisel-shaped, giving a “buck <strong>to</strong>othed” appearance. Wings<br />

w/ large claws, but remiges unknown. Legs long. Tail frond small & square. Biology: The<br />

large, chisel shaped teeth indicate that these were probably specialized herbivores. The<br />

wings were relatively small, & the remix length unknown, but they may have at least allowed<br />

some limited parachuting ability, slowing descents on the rare occasions that they<br />

climbed or rested in trees (Chatterjee & Templin 2004). The contemporary species Incisivosaurus<br />

gauthieri, known from a complete skull, is probably a synonym.<br />

Zou’s Tail Feather Caudipteryx zoui<br />

Time: 124.5 Ma ago Location: Liaoning, China Habitat: Lower Yixian Formation (see<br />

above) Size: WS 70cm (2.2ft); BL 90cm (3ft); TL 1m (3.3ft) Features: Head triangular w/<br />

narrow, pointed snout. Teeth restricted <strong>to</strong> tip of upper jaw; lower jaw <strong>to</strong>othless. Upper<br />

front two teeth large. Wings very small & lacking secondary remiges. Minor digit highly<br />

reduced, lacking a claw & probably fused <strong>to</strong> major digit via soft tissue. Alular claw slightly<br />

smaller than major claw. Legs long. Tail very short, w/ rectrical frond limited <strong>to</strong> the final<br />

third of the tail length. All rectrices nearly equal in length, creating a square- or diamondshaped<br />

frond when seen from above. Biology: Presence of numerous small gizzard s<strong>to</strong>nes<br />

in some fossils indicates probable herbivorous diet. Tail frond light in color with a dense,<br />

dark b<strong>and</strong>ed pattern, suitable both for display <strong>and</strong> camouflage. Highly reduced wing feathers<br />

able <strong>to</strong> fold tightly against the body & may have also been used in display. Fossils of this<br />

species are common only in a relatively small region of the Yixian Formation, indicating<br />

very specific habitat preferences.<br />

Dong’s Tail Feather Caudipteryx dongi<br />

Time: 124.5 Ma ago Location: Liaoning, China Habitat: Lower Yixian Formation (see<br />

above) Size: WS 110cm (3.6ft); BL 75cm (2.5ft); TL 90cm (3ft) Features: Head long &<br />

relatively low & narrow, w/ teeth restricted <strong>to</strong> tip of upper jaw; lower jaw <strong>to</strong>othless. Upper<br />

front two teeth large. Neck long & slender w/ short feathers. Wings relatively large<br />

compared <strong>to</strong> <strong>other</strong> caudipterids. Secondary remiges long but restricted <strong>to</strong> the outer wing.<br />

Minor digit highly reduced, lacking a claw & probably fused <strong>to</strong> major digit via soft tissue.<br />

Biology: Similar <strong>to</strong> C. zoui, often considered synonymous. However, larger wings & presence<br />

of secondary remiges in a smaller specimen is opposite the growth pattern seen in<br />

Similicaudipteryx, so C. dongi may be a distinct species.<br />

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