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A field guide to mesozoic birds and other winged dinosaurs

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Appendix C: Evolutionary<br />

Linnaean Classification<br />

Several attempts have been made in recent years <strong>to</strong> construct traditional<br />

or Linnaean classifications of Mesozoic <strong>birds</strong>. However, most of<br />

these attempts have been flawed in some way. For example, Livezey & Zusi<br />

(2007) presented a classification of all <strong>birds</strong> which included only a few Mesozoic<br />

taxa. These were united in several unorthodox, often paraphyletic<br />

groups which contradicted most contemporary phylogenies (e.g. uniting<br />

Archaeopteryx <strong>and</strong> Confuciusornis in a single group <strong>to</strong> the exclusion of<br />

<strong>other</strong> <strong>birds</strong>, including Rahonavis as an euenantiornithean etc.). While Linnaean<br />

classifications are clearly not as useful as phylogenies for elucidating<br />

the interrelationships of taxa, there are situations where they may be useful<br />

for communication. For that reason they should be made as comprehensive<br />

<strong>and</strong> as phylogenetically rigorous as possible within the confines of the<br />

Linnaean structure, <strong>and</strong> for these reasons Livezey & Zusi’s classification is<br />

considered inadequate.<br />

An<strong>other</strong> Linnaean classification was presented by Ben<strong>to</strong>n (2004).<br />

While more phylogenetically rigorous than that of Livezey & Zusi (2007), it<br />

suffered from oversimplification due <strong>to</strong> few included taxa. Presented here is<br />

a more comprehensive <strong>and</strong> rigorous Linnaean taxonomy of all known Mesozoic<br />

bird taxa (Subfamily rank <strong>and</strong> above). The goal of this taxonomy is<br />

not <strong>to</strong> be used in preference <strong>to</strong> phylogenetic nomenclature, but <strong>to</strong> be available<br />

as an alternative <strong>to</strong> published Linnaean classifications on the occasion<br />

that the use of Linnaean ranks is necessary or preferred by the author. The<br />

names <strong>and</strong> placement of taxa listed here are informed by their phylogenetic<br />

definitions in Appendix B.<br />

Class Aves Linnaeus 1758<br />

Order Caenagnathiformes Sternberg 1940<br />

Family Caudipteridae Zhou & Wang 2000<br />

Family Avimimidae Kurzanov, 1981<br />

Superfamily Caenagnathoidea Sternberg 1940<br />

Family Caenagnathidae Sternberg 1940<br />

Subfamily Caenagnathinae Sternberg 1940<br />

Subfamily Elmisaurinae Osmolska 1981<br />

Family Ovirap<strong>to</strong>ridae Osborn 1924<br />

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