A field guide to mesozoic birds and other winged dinosaurs
Barrett’s Seabird Enaliornis barretti Time: 100 Ma ago Location: England, UK Habitat: Cambridge Greensand Formation. Shores and lagoons surrounding semi-tropical islands in a shallow sea. Size: WS unknown; BL ~55cm (1.8ft); TL unknown Features: Marine hesperornitheans known from a partial skeleton & a number of fragments. Head small, but most of the skull & snout unknown. Legs very long, w/ long, robust tibia, short & stocky femur (probably less than half tibia length), & short but robust tarsus shorter than the femur. Distinguished by their small size, primitive characteristics, & internal characteristics of the hind limbs & pelvis (lack of femoral neck, mid-sized antitrochantor, keel beneath the synsacrum). Biology: These small seabirds combined some features in common with more advanced hesperornitheans, while retaining some primitive features found in ichthyornitheans & other carinatans. They were therefore probably coastal divers or dippers which may have retained some flight ability, though the wing is not known. Tanke’s Pasquia Bird Pasquiaornis tankei Time: 95 Ma ago Location: Saskatchewan, Canada Habitat: Ashville Formation, nearshore marine enviornment in the northern Western Interior Seaway Size: WS >36cm (1.2ft); BL ~70cm (2.3ft); TL unknown Features: Jaws long & toothed. Wings relatively long. Legs long & slender. Biology: The legs were positioned more directly under the body than in later hesperornitheans, possibly allowing a waddling gait on land. The toes could not twist during proportion as in lobed-toe birds, indicating normally webbed feet. The relatively long & robust wing bones suggest possible retention of flight ability, though the sternum is unknown. Probably surface swimmers (Sanchez 2012). Varner’s Brodkorb Bird Brodavis varneri Time: 78 Ma ago Location: Kansas, USA Habitat: Sharon Springs Member, Pierre Shale Formation. Relatively shallow river deltas and estuaries opening into the Western Interior Seaway. Size: WS unknown; BL ~90cm (3ft); TL unknown Features: Marine hesperornitheans. Body long & barrel-shaped, w/ short torso & long hips. Legs very long, w/ tibia over half the length of the torso + hips. Tarsus short & stocky, less than ¼ tibia length. Neck long, probably equal in length to torso + hips. Wings unknown. Biology: The hip bones of these birds were not as advanced & highly fused as in related species. This, compared w/ the robust foot bones, may suggest a slightly different method of underwater propulsion compared w/ similar species inhabiting deeper saltwater environments off the coasts. The skeleton was overall more lightly built & not as solid as other hesperornitheans. This implies a more limited diving ability, restricting these birds to near-surface feeding. The lighter skeleton may also suggest some flight ability was retained. 166
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- Page 183 and 184: Glossary • Alula: Vaned, pennaceo
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Barrett’s Seabird Enaliornis barretti<br />
Time: 100 Ma ago Location: Engl<strong>and</strong>, UK Habitat: Cambridge Greens<strong>and</strong> Formation.<br />
Shores <strong>and</strong> lagoons surrounding semi-tropical isl<strong>and</strong>s in a shallow sea. Size: WS unknown;<br />
BL ~55cm (1.8ft); TL unknown Features: Marine hesperornitheans known from a partial<br />
skele<strong>to</strong>n & a number of fragments. Head small, but most of the skull & snout unknown.<br />
Legs very long, w/ long, robust tibia, short & s<strong>to</strong>cky femur (probably less than half tibia<br />
length), & short but robust tarsus shorter than the femur. Distinguished by their small<br />
size, primitive characteristics, & internal characteristics of the hind limbs & pelvis (lack<br />
of femoral neck, mid-sized antitrochan<strong>to</strong>r, keel beneath the synsacrum). Biology: These<br />
small sea<strong>birds</strong> combined some features in common with more advanced hesperornitheans,<br />
while retaining some primitive features found in ichthyornitheans & <strong>other</strong> carinatans.<br />
They were therefore probably coastal divers or dippers which may have retained some<br />
flight ability, though the wing is not known.<br />
Tanke’s Pasquia Bird Pasquiaornis tankei<br />
Time: 95 Ma ago Location: Saskatchewan, Canada Habitat: Ashville Formation, nearshore<br />
marine enviornment in the northern Western Interior Seaway Size: WS >36cm<br />
(1.2ft); BL ~70cm (2.3ft); TL unknown Features: Jaws long & <strong>to</strong>othed. Wings relatively<br />
long. Legs long & slender. Biology: The legs were positioned more directly under the body<br />
than in later hesperornitheans, possibly allowing a waddling gait on l<strong>and</strong>. The <strong>to</strong>es could<br />
not twist during proportion as in lobed-<strong>to</strong>e <strong>birds</strong>, indicating normally webbed feet. The<br />
relatively long & robust wing bones suggest possible retention of flight ability, though the<br />
sternum is unknown. Probably surface swimmers (Sanchez 2012).<br />
Varner’s Brodkorb Bird Brodavis varneri<br />
Time: 78 Ma ago Location: Kansas, USA Habitat: Sharon Springs Member, Pierre Shale<br />
Formation. Relatively shallow river deltas <strong>and</strong> estuaries opening in<strong>to</strong> the Western Interior<br />
Seaway. Size: WS unknown; BL ~90cm (3ft); TL unknown Features: Marine hesperornitheans.<br />
Body long & barrel-shaped, w/ short <strong>to</strong>rso & long hips. Legs very long, w/ tibia<br />
over half the length of the <strong>to</strong>rso + hips. Tarsus short & s<strong>to</strong>cky, less than ¼ tibia length. Neck<br />
long, probably equal in length <strong>to</strong> <strong>to</strong>rso + hips. Wings unknown. Biology: The hip bones<br />
of these <strong>birds</strong> were not as advanced & highly fused as in related species. This, compared<br />
w/ the robust foot bones, may suggest a slightly different method of underwater propulsion<br />
compared w/ similar species inhabiting deeper saltwater environments off the coasts.<br />
The skele<strong>to</strong>n was overall more lightly built & not as solid as <strong>other</strong> hesperornitheans. This<br />
implies a more limited diving ability, restricting these <strong>birds</strong> <strong>to</strong> near-surface feeding. The<br />
lighter skele<strong>to</strong>n may also suggest some flight ability was retained.<br />
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