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A field guide to mesozoic birds and other winged dinosaurs

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Martin’s Yan Bird Yanornis martini<br />

Time: 120 Ma ago Location: Liaoning, China<br />

Habitat: Jiufotang Formation Size: WS ~80cm (2.6ft); BL 32cm (1ft); TL unknown Features:<br />

Head small & triangular, w/ small beak at jaw tips. Teeth located behind beak. Neck<br />

long. Wings long (manus, ulna & humerus ~equal in length, ¾ the length of <strong>to</strong>rso), broad,<br />

<strong>and</strong> powerful. Alular digit clawed. Small claw on major digit. Minor digit fused <strong>to</strong> major.<br />

Legs short, w/ femur & tibia equal in length, tarsus ¾ length of tibia. Toes equal <strong>to</strong> tarsus<br />

length w/ robust but weakly curved claws. Hallux small. Biology: Evidence shows that<br />

these fed on small fish, but also ingested large amounts of gizzard s<strong>to</strong>nes like herbivorous<br />

<strong>birds</strong>, so were probably omnivorous generalists. May have employed seasonal diet switching.<br />

Feet characteristic of ground foragers. Robust body, small head & long neck may have<br />

given these an appearance similar <strong>to</strong> modern game <strong>birds</strong> like pheasants.<br />

Grabou’s Yixian Bird Yixianornis graboui<br />

Time: 120 Ma ago Location: Liaoning, China Habitat: Jiufotang Formation Size: WS<br />

60cm (2ft); BL 27cm (11in); TL 35cm (1.1ft) Features: Head small & triangular, w/ small<br />

beak at jaw tips. Teeth short & peg-like. Neck long. Wings short (humerus & ulna just<br />

over half <strong>to</strong>rso length, manus shorter than ulna) but broad (primary feather length up <strong>to</strong><br />

15cm). Alular & major digits w/ small weakly curved claws. Minor digit fused <strong>to</strong> major.<br />

Legs long w/ long tibia but short tarsus (less than half tibia length). Toes very long & thin<br />

(nearly twice the length of the tarsus). Toes w/ small & very weakly curved claws. Tail w/ at<br />

least eight pennaceous rectrices up <strong>to</strong> 8cm long arranged in<strong>to</strong> exp<strong>and</strong>able fan, somewhat<br />

rounded in shape when exp<strong>and</strong>ed. Biology: Similar <strong>to</strong> Y. martini, but w/ much longer legs,<br />

longer, more slender <strong>to</strong>es, & somewhat shorter wings. Comparison of leg & wing proportions<br />

<strong>to</strong> modern <strong>birds</strong> indicate these may have been semi-aquatic, foraging both from the<br />

water surface & on the ground like modern ducks & geese. May have employed some degree<br />

of foot-propelled diving (Bell & Chiappe 2010). While there is currently no evidence<br />

of webbed feet, the long, slender <strong>to</strong>es & small, nearly straight claws would be consistent w/<br />

this ecology.<br />

Linghe Songling Bird Songlingornis linghensis<br />

Time: 120 Ma ago Location: Liaoning, China Habitat: Jiufotang Formation Size: WS unknown;<br />

BL 10cm (4in); TL unknown Features: Head slender w/ thin, pointed jaws & numerous,<br />

close-packed teeth. Jaw tips beaked. Wings known from fragments, but indicate<br />

well-developed shoulder girdle musculature. Biology: Similar in many respects <strong>to</strong> <strong>other</strong><br />

yanornithiformes, except for small size & better-developed wings. Probably omnivorous<br />

shore<strong>birds</strong> wading or swimming after fish & <strong>other</strong> forage.<br />

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