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A field guide to mesozoic birds and other winged dinosaurs

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Thorn-nosed Dapingfang Bird Dapingfangornis sentisorhinus<br />

Time: 120 Ma ago Location: Liaoning, China Habitat: Jiufotang Formation, temperatesubtropical<br />

swamps dominated by ginkgo <strong>and</strong> conifer trees, set among shallow lakes <strong>and</strong><br />

stagnant waterways. Size: WS 22cm (8.6in); BL 12cm (4.7in); TL 23cm (9in) Features:<br />

Head large w/ short, thin snout, feathered <strong>to</strong> near the tip. Teeth sharp <strong>and</strong> recurved. Feet<br />

w/ strong talons. Single pair of very long <strong>and</strong> thin wire-like rectrices tipped w/ broad vaned<br />

plumes. Biology: Probably carnivorous, feeding on fish, lizards & <strong>other</strong> small vertebrates.<br />

A supposed horn high on the snout may be an artifact of preservation, though it may<br />

reflect a pair of actual, laterally-projecting lachrymal bones (Mortimer 2010). However,<br />

these were shorter than the overlying feathers & probably would not have been visible in<br />

life.<br />

Chaoyang Long-wing Longipteryx chaoyangensis<br />

Time: 120 Ma ago Location: Liaoning, China Habitat: Jiufotang Formation. Temperatesubtropical<br />

swamps dominated by ginkgo <strong>and</strong> conifer trees, set among shallow lakes <strong>and</strong><br />

stagnant waterways. Size: WS 34cm (1.1ft); BL 16cm (6in); TL ~18cm (7in) Features:<br />

Snout very long w/ large, curved teeth restricted <strong>to</strong> the jaw tips. Wings large & broad (medial<br />

primary 9cm) w/ large alular & major claws. Minor claw vestigial. Legs short. Stub-tail<br />

lacked any long rectrices. Biology: Teeth large & conical but slightly flattened <strong>and</strong> curved;<br />

this, combined with their presence only in the jaw tips, has led <strong>to</strong> the suggestion that these<br />

were fishers. However, short leg <strong>and</strong> perching feet may indicate an arboreal, insectivorous<br />

lifestyle instead. Broad wings <strong>and</strong> wishbone ana<strong>to</strong>my suggest that these <strong>birds</strong> may have<br />

employed continuous flapping flight.<br />

Zheng’s Boluochi Bird Boluochia zhengi<br />

Time: 120 Ma ago Location: Liaoning, China Habitat: Jiufotang Formation (see above)<br />

Size: WS ~40cm (1.3ft); BL ~20cm (8in); TL unknown Features: Very similar <strong>to</strong> “C.” yangi,<br />

also w/ a long, narrow snout <strong>and</strong> very large, strongly hooked teeth. Differed primarily in<br />

smaller size & foot ana<strong>to</strong>my, where the outer <strong>to</strong>e diverged significantly from the <strong>other</strong>s.<br />

Biology: Possibly the same species as “C.” yangi. B. zhengi was initially reported <strong>to</strong> have a<br />

hooked, rap<strong>to</strong>rial beak, but this was a misinterpretation due <strong>to</strong> poor preservation <strong>and</strong> the<br />

unusually large, hooked teeth.<br />

Yang’s Bent-<strong>to</strong>oth “Camp<strong>to</strong>dontus” yangi<br />

Time: 120 Ma ago Location: Liaoning, China Habitat: Jiufotang Formation (see above)<br />

Size: WS ~50cm (1.6ft); BL 25cm (10in); TL unknown Features: Head long w/ very long<br />

snout. Teeth extremely large & strongly curved w/ a distinctive bend at the tip, & restricted<br />

<strong>to</strong> the jaw tips. Wings long w/ alular & major claws. Legs relatively short. Biology: Similar<br />

in many respects <strong>to</strong> Longipteryx, but notable for the much larger teeth, which would have<br />

protruded conspicuously from the slender jaws in life. Larger than Longipteryx, these may<br />

also have fed on fish &/or arboreal prey, but possibly specializing in larger prey than L.<br />

chaoyangensis. The genus name Camp<strong>to</strong>dontus is preoccupied <strong>and</strong> will need <strong>to</strong> be replaced<br />

if it is not a synonym of Boluochia.<br />

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