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A field guide to mesozoic birds and other winged dinosaurs

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Holy Confucius Bird Confuciusornis sanctus<br />

Time: 124.6-122 Ma ago Location: Liaoning, China Habitat: Yixian Formation Size: WS<br />

63cm (2ft); BL 25cm (10in); TL 50cm (1.6ft) Features: Head round w/ long, sharply pointed<br />

bill. Body black <strong>to</strong> dark grey, mottled w/ chestnut or reddish brown. Wings extremely<br />

long & pointed in adults, slightly shorter in immature specimens. Wing claws very large,<br />

strongly curved & unfeathered. Primaries white, secondaries & coverts grey <strong>to</strong> black. Legs<br />

very short w/ small feet <strong>and</strong> medial hallux. Males ribbon-tailed w/ single pair of very long<br />

rectrices. Females lack rectrices, tail consisting of tiny coverts (also present in males). Biology:<br />

Young chicks have longer, bony tails which fuse & shorten as they mature. Males<br />

develop rectrices at a young age at or near hatching; rectrices are proportionally longer in<br />

smaller individuals. Growth slow during 1st year or two, & rapid from mid- <strong>to</strong> full-size.<br />

Wing ana<strong>to</strong>my radically different from avians; breast small & undeveloped, breastbone<br />

bears only a small, cartilaginous keel. Large opening in humerus indicates unique musculature<br />

allowing some degree of flapping, but angle at which wings could be raised above<br />

back limited by shoulder ana<strong>to</strong>my. Probably scaled trees using large wing claws, may have<br />

been able <strong>to</strong> navigate canopy using medial hallux. May have glided between trees using<br />

weak flapping, perhaps swooping in flocks over lakes where many have been preserved. By<br />

far the most common bird species in the Yixian, they were possibly colonial, forming huge<br />

flocks, some of which died en masse during volcanic eruptions & were buried simultaneously<br />

on the lake bot<strong>to</strong>ms. Young juveniles absent from large fossil assemblages, indicating<br />

that <strong>birds</strong> below mid-size lived in isolation or in more inl<strong>and</strong> environments before joining<br />

flocks. Diet is unknown in this species.<br />

Du’s Confucius Bird Confuciusornis dui<br />

Time: 124.6 Ma ago Location: Liaoning, China Habitat: Yixian Formation Size: WS 52cm<br />

(1.7ft); BL 17cm (7in); TL 40cm (1.3ft) Features: Very similar in overall ana<strong>to</strong>my <strong>to</strong> C.<br />

sanctus. Differed in smaller adult size, an upturned bill tip, & a much smaller alular claw.<br />

Biology: The differing bill shape is a strong indica<strong>to</strong>r of a unique diet, though no s<strong>to</strong>mach<br />

contents are known. Later confuciusornithids are known <strong>to</strong> have fed on small fish, & this<br />

seems like a good possibility for the bill function of C. dui.<br />

Zheng’s Dawn Confucius Bird Eoconfuciusornis zhengi<br />

Time: 131 Ma ago Location: Hebei, China Habitat: Sichakou Member, Huajiying Formation<br />

Size: WS 38cm (1.2ft); BL 17cm (7in); TL 40cm (1.3ft) Features: Beak short & pointed,<br />

head triangular. Wings pointed but broad & relatively s<strong>to</strong>ut, w/ long secondaries. Legs<br />

short with partially reversed hallux. Ribbon-tail consisting of a single pair of very long<br />

rectrices. Biology: Dark preservation of melanin in the only known specimen suggests a<br />

very dark, possibly black uniform coloration. Similar in most respects <strong>to</strong> the Confuciusornis,<br />

but much earlier chronologiacally. Also differed in small shoulder bones (coracoids) &<br />

relatively solid vertebrae.<br />

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