Horticulture Principles and Practices

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(b) F 1 Rr Rr round round F 2 RR Rr Rr rr Phenotype: 3 round 1 wrinkled Punnett square analysis R r P 1 P 2 RR round rr wrinkled R RR round Rr round Rr round F 1 r Rr round rr wrinkled (a) (a) Homozygous loci (RR,rr) interact to produce heterozygous locus (Rr); (b) heterozygous loci interact to reveal homozy- FIGURE 6–1 gous locus (rr). 6.6 COMPLEX INHERITANCE Mendel’s work focused on traits that were controlled by one or a few genes. Such traits are called simply inherited or Mendelian traits. The genetics are straightforward and easy to breed. Unfortunately, many traits are controlled by many genes and hence have complex inheritance that is not readily amenable to Mendelian treatment. At the simpler end of the complex inheritance spectrum is incomplete dominance (or partial dominance), whereby the effect of the dominant allele over the recessive allele is not decisive, resulting in an average or blending of the two effects (Figure 6–2). In another situation, the two contrasting alleles are equally expressed (codominance). There are situations in nature whereby one gene has more than two alternative forms (multiple alleles), one being the ABO blood groups in humans. On the other end of the inheritance spectrum are traits that are governed by numerous genes whose individual effects are difficult to distinguish (polygenes, quantitative genes). The trait they control is called quantitative traits. Each gene contributes little effect to the overall phenotype. Such traits are best measured (metrical traits) rather than counted as in Mendelian traits because they are difficult to breed. Mendelian traits can be neatly categorized into distinct classes (ratios) by counting. Sometimes the expected ratios do not manifest because of various interferences. The phenomenon of epistasis, whereby alleles at different loci (nonallelic genes) interact, can modify genetic outcomes and consequently the expected ratios. For example, instead of the Quantitative Traits Traits that are controlled by many genes, each contributing a small effect to the overall phenotypic expressions of the trait. 6.6 Complex Inheritance 197
FIGURE 6–2 Incomplete dominance gene action produces pink flowers instead of red in the case of complete dominance. P P 1 RR (red) P 2 rr (white) F 1 Rr Rr Rr (red) F 2 Phenotypic ratio: RR Rr Rr rr (red) (pink) (pink) (white) 1 red : 2 pink : 1 white 9:3:3:1 ratio expected when two loci of interest are considered (i.e., a dihybrid cross), alleles at another locus can contribute to the expression of the phenotype, resulting in modifications of the ratio. Examples of modifications are 9:7 (complementary genes), 13:1 (suppressor genes), and 15:1 (duplicate genes). Such modifications can complicate breeding efforts. Genes that are located on the same chromosome are said to be linked (genetic linkage) and are expected to be inherited from one generation to the next as a block. However, the event of crossing over can interfere with such smooth, intact transmission of gene blocks. Some genes are more tightly linked than others. The former are difficult to interrupt or break. It is a blessing to plant breeding when good gene blocks occur. Unfortunately, there are occasions in which good and desirable genes are linked with undesirable genes! Plant breeders used techniques like repeated crossing to provide opportunities for genetic recombination to occur to break undesirable genetic associations. 198 Chapter 6 Breeding Horticultural Plants 6.7 THE IMPORTANCE OF REPRODUCTIVE SYSTEMS Plant breeders must be familiar with the reproductive systems of the plants they seek to manipulate. The genetic structure of plants depends on their mode of reproduction. Plants may be sexually reproducing or asexually reproducing, the latter not involving the sexual process or crossing. Asexually reproducing species remain “pure” or genetically identical from one generation to the next (clones) because the sexual process involves meiosis, the process that creates natural genetic variability. To breed sexually reproducing species, the breeder must understand the floral biology and other factors associated with their flowering. To cross plants, you must know when the flowers open, how long the pollen stays viable, when the stigma is receptive, and other factors. Some sexually reproducing species are called self-pollinating because they receive pollen grains from their own flowers or from like plants. This habit makes them have a narrow genetic base. They are homozygous at most loci and produce relatively uniform products. Other species are cross-pollinating, receiving pollen from an available source. Cross-pollinated species are heterozygous at most loci and have a broad genetic base. These two basic reproductive behaviors dictate how the species are genetically manipulated. If you treat a cross-pollinated species as self-pollinated, once you remove the artificial restriction, it would revert back to its old ways! In addition to the state genetic differences, there is a critical factor that plant breeders exploit in breeding plants. It is known that the farther apart genetically parents in a cross are, the more vigorous the product of their mating. Because cross-pollinated species are heterozygous, they can maintain a high genetic load without consequences. Lethal recessive alleles can enjoy a heterozygous advantage, remain innocuous while being suppressed by a dominant allele. However, when selfed (crossed with itself,
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HORTICULTURE Principles and Practic
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HORTICULTURE Principles and Practic
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With love to Theresa, quarterback;
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Brief Contents Preface xxi PART 1 T
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Contents Preface xxi PART 1 THE UND
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5.3 PLANT GROWTH PROCESSES 160 5.4
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8.20 COMMON GREENHOUSE DISEASES 276
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12.3 INTERNAL ENVIRONMENTAL CONTROL
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PART 6 Summary 541 References and S
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22.18 INDOOR COMPOSTING SYSTEMS 668
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Preface Horticulture is the area of
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ACKNOWLEDGMENTS I am very grateful
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PART 1 THE UNDERLYING SCIENCE CHAPT
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1 What Is Horticulture? PURPOSE AND
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(a) (c) (b) (d) FIGURE 1-1 The many
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FIGURE 1 Bridge. The plaza view of
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CYCADS Many people mistake these pr
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FIGURE 2 The world's largest unbran
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FIGURE 2 Sold flowers are loaded on
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FIGURE 1-4 Formal landscaping featu
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1.4 ROLEOFTHENURSERY AND SEED INDUS
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1.5 HORTICULTURE AND SOCIETY Hortic
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TABLE 1-3 U.S. Horticultural Export
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Turfgrass Operation 1. Landscape te
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What Is Horticulture? This site pro
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Examples of botanical gardens http:
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2 Classifying and Naming Horticultu
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Eight major taxa are commonly used
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TABLE 2-3 The Divisions of the King
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HISTORY OF PLANT TAXONOMY PAUL R. F
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AGE OF HERBALISTS Two major events
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possible system of nomenclature. Ho
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TABLE 1 Type Categories for Plant N
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2.3 OTHER CLASSIFICATION SYSTEMS (O
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2. Shrubs. A shrub has no main trun
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Simple Fruits Fleshy Fruits Drupe B
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FIGURE 2-14 A pome, represented by
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2.3.5 CLASSIFICATION OF VEGETABLES
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(a) (b) FIGURE 2-22 (Source: George
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FIGURE 2-25 A narrowleaf plant. (So
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FIGURE 2-29 Parts of a typical gras
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such as rosemary, sage, thyme, marj
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c. Leaves d. Bulbs 2. Cut across (t
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Whole plant Organs FIGURE 3-1 Level
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ibonucleic acid (RNA), proteins, an
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called cristae; this extreme foldin
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By virtue of its position, the prim
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Phloem Tissue Structurally, phloem
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(a) Stalk (b) Culm FIGURE 3-5 Cross
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Scale Compressed stem (a) Whole bul
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Upper epidermis Palisade layer FIGU
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usually occur in xerophytes. In cer
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FIGURE 3-22 Selected common leaf ma
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FIGURE 3-25 Selected common leaf ti
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absorption of water and minerals fr
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Outer bark Inner bark FIGURE 3-37 T
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Anther Filament Stamen FIGURE 3-41
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Exocarp Parts of a typi- FIGURE 3-4
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PRACTICAL EXPERIENCE LABORATORY 1.
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4.1 CLIMATE, WEATHER, AND HORTICULT
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concentration in the atmosphere.A c
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TABLE 4-1 Climatic Adaptation of Se
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and upward. Another important gener
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Rate of photosynthesis mg/sq. dm/hr
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and plants that flower under only c
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times of the year. Growers start th
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content. This section is sometimes
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TABLE 4-7 Soil Mineral Nutrients Es
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Micronutrients (Trace Elements) Mic
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Neutral FIGURE 4-11 A representatio
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4.4 FERTILIZERS Fertilizer sources
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One of the most commonly used contr
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Chlorosis (the yellowing of green l
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Fertilizers may be applied before p
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It is neither practical nor safe to
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Solution: How much of ammonium nitr
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1°C (34°F), the optimum temperatu
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Cellulose sponge Perched water tabl
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Overhead Sprinkler Irrigation Water
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FIGURE 4-19 Furrow irrigation of le
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can self-install an underground irr
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1. Surface drainage. Surface draina
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Secondary Tillage Primary tillage i
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(a) (b) (c) (d) FIGURE 4-20 (Source
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texture. The most commonly used gra
Page 172 and 173: TABLE 4-11 Selected Standard Mixes
Page 174 and 175: Steam Pasteurization Steam pasteuri
Page 176 and 177: Maracher, H. 1986. Mineral nutritio
Page 178 and 179: 5 Plant Physiology PURPOSE AND EXPE
Page 180 and 181: Growth in an organism follows a cer
Page 182 and 183: 5.1.2 THE ROLE OF SIGNALS IN GROWTH
Page 184 and 185: waxes are embedded. Waxes consist o
Page 186 and 187: 5.3.1 PHOTOSYNTHESIS Photosynthesis
Page 188 and 189: CO 2 FIGURE 5-6 The C 4 pathway of
Page 190 and 191: Growth and Development The general
Page 192 and 193: TABLE 5-2 Energy Produced from Aero
Page 194 and 195: Certain plants are adapted to dry e
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Page 198 and 199: Shoot Elongation In certain plants,
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Page 202 and 203: FIGURE 5-13 Ripening of plantain sh
Page 204 and 205: Terminal bud removed Unbranched pla
Page 206 and 207: conditions—pertaining to light, m
Page 208 and 209: them to maturity. The major process
Page 210 and 211: 6 Breeding Horticultural Plants PUR
Page 212 and 213: Similarly, there can be no plant br
Page 214 and 215: APPLICATION, CHALLENGES, AND PROSPE
Page 216 and 217: hit with target DNA. Therefore, it
Page 218 and 219: Generally, within ten days of exper
Page 220 and 221: Aziz A.N., Sauve R.J., Zhou S., 200
Page 224 and 225: e.g., Aa × Aa), the lethal allele
Page 226 and 227: eeder’s equation. Simply stated,
Page 228 and 229: Before the seed or product becomes
Page 230 and 231: 6.18.2 THE GENERAL STEPS OF RDNA TE
Page 232 and 233: 2. Political disagreement. There ar
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Page 236 and 237: PART 2 PROTECTING HORTICULTURAL PLA
Page 238 and 239: 7 Biological Enemies of Horticultur
Page 240 and 241: 8. Weeds may clog drains, waterways
Page 242 and 243: is also a root parasite that obtain
Page 244 and 245: LEGISLATIVE Both state and federal
Page 246 and 247: Example Integrated cultural, physic
Page 248 and 249: 7.4.2 IMPORTANT INSECT ORDERS Insec
Page 250 and 251: Egg FIGURE 7-3 Life cycle of an ins
Page 252 and 253: sucking insects (also found with so
Page 254 and 255: FIGURE 7-12 Corn earworm damage. (S
Page 256 and 257: TABLE 7-1 Selected Fungal Diseases
Page 258 and 259: 7.6.1 SMALL ANIMALS Rabbits, mice,
Page 260 and 261: FIGURE 7-16 The disease triangle. P
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Page 264 and 265: SUMMARY Insects are a major class o
Page 266 and 267: For the home growers or those who c
Page 268 and 269: for consumers and the environment).
Page 270 and 271: TABLE 8-1 Strategy 4: Strategies an
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gibberellic acid spray overcomes st
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In a competitive industry, a variet
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Chemicals gain access to humans thr
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2. Pesticide management. Controllin
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Every organism has its natural enem
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TABLE 8-3 Selected Examples of Biol
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1 2 YEAR 3 4 FIGURE 8-5 cycle. A cr
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6. Heat treatment. In the greenhous
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Organic Compounds (Organics) Organi
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under enclosed conditions (e.g., wa
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FIGURE 8-9 A tractor-mounted spraye
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8.11.9 LANDSCAPE PESTS AND THEIR CO
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application, a particular herbicide
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Further, they do not provide unifor
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SUMMARY Herbicides are chemicals us
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Sulfur may be applied for both prev
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8.23 PREVENTING GREENHOUSE DISEASES
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PART 3 PROPAGATING HORTICULTURAL PL
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9 Sexual Propagation PURPOSE AND EX
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Anther Microspore Megaspore mother
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Lettuce seeds Red light Darkness Fa
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FEDERAL AND STATE SEED LAWS Federal
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Germination Test In laboratory prac
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FIGURE 15 The essential structures
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processing into flour or meal). How
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physiologically immature seeds must
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seeds may be treated in this way be
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The two basic modes of seedling eme
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locations in the field. Home garden
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FIGURE 9-9 A plastic flat. (Source:
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(a) (b) FIGURE 9-12 (a) Sowing seed
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y the gardener or grower. Whatever
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REFERENCES AND SUGGESTED READING Co
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species enables vegetative propagat
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for rapid rooting. There are two ba
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Cutting involving one node (e.g., s
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This practice is especially importa
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10.6.4 STICKING THE CUTTING Cutting
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(a) Indexing by budding Diseased pl
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10.11 M ETHODS OF GRAFTING Grafting
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Scion Wax FIGURE 10-17 Steps in bar
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MODULE 3 BUDDING 10.12 TYPES OF BUD
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MODULE 4 LAYERING 10.13 TYPES OF LA
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Buried part of shoot is nicked FIGU
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variety of ways. In air layering, a
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FIGURE 10-34 by using cormels. Prop
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Psuedobulbs In the Dendrobium orchi
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The technique is used widely in cro
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PART 4 GROWING PLANTS INDOORS CHAPT
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11 Growing Houseplants PURPOSE AND
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TABLE 11-1 Common houseplants Commo
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Saddle leaf Philodendron selloum To
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Window Displays Plants in windows e
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CONTAINER GARDENS DR. TERRI W. STAR
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annuals and hardy perennial species
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of the large container filled with
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perfection about four to six weeks
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FIGURE 11-6 Flowers displayed on th
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TABLE 11-5 Plant Selected Plants fo
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The lighting condition near these w
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Fluorescent Lights Fluorescent ligh
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may be used for one pot or a group
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garden rooms, atriums, or a large c
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The photoperiod affects when the ho
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patted firm to keep the plant erect
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Other Materials Apart from clay and
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(a) ( FIGURE 11-25 Support for plan
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TABLE 11-7 Common Problems of House
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• Keep soil moist all the time
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• Prefers high temperatures • P
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amount and quality of light. If sup
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12 Controlled-Environment Horticult
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6. Curvilinear 7. Curved eave 8. Do
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Detached greenhouses have several a
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12.2.3 FRAME DESIGN There are two b
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horticultural business a less-expen
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Texas, Hawaii, and California. The
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source of heat for times when the t
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FIGURE 12-17 Greenhouse production
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FIGURE 12-21 Moving tables allowing
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Research program on greenhouse engi
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greenhouses equipped with a variety
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FIGURE 1 Annual energy required per
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This system was demonstrated in a 5
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FIGURE 6 Amounts of waste energy ut
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Ekholt, B.A., D.R. Mears, M.S. Gini
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or object to be warmed. Failure to
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objects in its path (e.g., the floo
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FIGURE 12-27 Motorized ventilation
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FIGURE 12-30 Movable internal shade
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FIGURE 12-33 A high pressure sodium
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Source of Water The quality of loca
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FIGURE 12-37 Overhead sprinkler irr
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Intermittent Feed Greenhouse plants
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However, in winter, greenhouse vent
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OUTCOMES ASSESSMENT 1. Explain the
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. Foliage or green plants. Foliage
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2. Labor. The size of the labor for
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FIGURE 13-1 Greenhouse production o
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FIGURE 13-2 Lettuce plug is inserte
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13.8.4 AGGREGATE HYDROPONIC SYSTEMS
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(a) (b) (c) FIGURE 13-6 Plug produc
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14 Growing Succulents PURPOSE AND E
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FIGURE 14-3 Leaf succulent represen
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frost-hardy. Their rosettes are usu
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TABLE 14-1 Plant Selected Popular S
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(a) (b) FIGURE 14-12 Typical bromel
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14.7.1 WHAT ARE CACTI? 14.7 CACTI C
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FIGURE 14-16 Opuntia. (Source: Crai
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FIGURE 14-23 Mammillaria. (Source:
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FIGURE 14-28 Both desert and jungle
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Growing mix Gravel Cacti (a) (b) FI
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PART5 GROWING PLANTS OUTDOORS: ORNA
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15 Principles of Landscaping PURPOS
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8. Create recreational grounds. Suc
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knowledge, with concern for resourc
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(a) (b) (c) FIGURE 15-2 The occurre
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GUIDELINES FOR LANDSCAPE DESIGN DAV
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the landscape. Some very successful
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Rhythm and Line Panoramic view of a
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FIGURE 15-10 A formal garden. The e
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How frequently do they entertain? A
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the patio should be located on the
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15.7.1 SELECTING PLANTS A homeowner
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Plant Arrangement in the Landscape
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Shadow FIGURE 15-15 Planting a tree
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SUMMARY Landscaping enhances the su
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3. Supply materials on a timely bas
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such as preparation rooms (for mixi
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of environmental fluctuations. Furt
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FIGURE 16-4 A bare-root tree seedli
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17 Installation of the Landscape PU
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for walks, driveways, and patios (F
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Planting may be limited to accentin
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17.3.3 PREPARING THE BED The soil s
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FIGURE 17-4 Bedding plants raised i
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SUMMARY Bedding plants are largely
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TABLE 17-6 Selected Ground Covers T
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TABLE 17-7 Selected Ornamental Gras
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they determine the success and surv
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12. Wildlife attraction. Trees in t
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pennsylvanica), hackberry (Celtis s
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y winds. A stake, which is often a
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TABLE 17-8 Selected Narrowleaf Ever
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TABLE 17-11 Selected Deciduous Shru
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Blooming bushes 1. Blue mist shrub
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Planting Bulblets and Bulbils Speci
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may be divided such that each secti
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FIGURE 18-1 (Source: George Acquaah
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Cool-Season (Temperate) Grasses In
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Growth Habit Turfgrasses are the mo
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Heavy Use Lawns on playgrounds and
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The seed should be free from weeds
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Source of Sod As with seed, sod sup
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A plug of sod FIGURE 18-7 Plugging
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way, plants are able to adapt to th
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form of a can placed on the lawn wi
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TABLE 18-3 Some Common Lawn and Tur
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even surface soil surface for layin
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MacCaskey, M. 1987. All about lawns
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Pruning is sometimes done in conjun
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4. Pruning may be done to reduce th
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19.4.2 SAWS A saw may be designed t
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defeat the purpose of pruning. The
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Bud withers as cut end dries back d
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19.6 STRATEGIES FOR PRUNING ABOVEGR
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Rejuvenation Pruning Cut canes to a
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3. In the third and subsequent year
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(a) Cut Prune (b) FIGURE 19-16 Step
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Eucalyptus and Paulownia are amenab
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TRAINING & PRUNING DECIDUOUS FRUIT
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Summer pruning eliminates an energy
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a) b) FIGURE 2 Newly planted apple
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FIGURE 6 Wooden limb spreaders can
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FIGURE 9. An apple tree trained to
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years to promote continued lateral
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Horizontal Espalier The horizontal
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19.16.1 CANE FRUITS Cane fruits are
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FIGURE 19-26 Shearing of Christmas
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pyramid-like form that is wider at
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After selecting the appropriate spe
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PART 6 GROWING PLANTS OUTDOORS: VEG
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20 Growing Vegetables Outdoors PURP
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The National Agricultural Statistic
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(This item omitted from WebBook edi
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growers should take to determine an
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pests and reduce/ eliminate hail da
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square yard (10 to 68 grams per squ
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High tunnels help increase the prof
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(This item omitted from WebBook edi
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20.4 VEGETABLE MARKET TYPES Fresh V
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Establishing the Crop Planting into
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home water supply from the tap. Thi
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Cole crop Cabbage Root Potato Bean
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6. Adequate nutrition. While overfe
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variable, ranging from creamy yello
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There are two general production pr
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This toxin is heat resistant and no
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large, or jumbo. The bulb may be sw
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REFERENCES Growing selected vegetab
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TABLE 21-1 Popular Herbs and Their
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(a) (b) (c) (d) (e) (f) FIGURE 21-1
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22 Organic Farming PURPOSE AND EXPE
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22.3 PRINCIPLES OF ORGANIC FARMING
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and the specific materials to be us
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22.8 MANAGING SOIL PHYSICAL QUALITY
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preemergent or early postemergent o
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Composting is a deliberate activity
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22.14.5 THE CARBON-TO-NITROGEN RATI
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Moisture Supply Water is required b
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Compost materials FIGURE 22-4 a wir
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As microbial decomposition proceeds
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Establishment and Management of an
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night, freezing can occur in spring
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accomplished by stratification. It
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transmitted by the dagger nematode
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PART 7 SPECIAL TECHNIQUES AND HANDL
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24 Cut Flowers and Floral Design PU
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to more than four-fold in standard
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Temperature and Humidity Wilting re
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FLORAL DESIGN: AN OVERVIEW BY WM. J
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Principle Definition Types (or Uses
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pH value-a measure of the acidity o
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FIGURE 6 Parallel Design-Parallel d
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24.3.2 TOOLS AND MATERIALS The tool
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3. Establish the focal point. 4. Ad
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Natural Drying To dry naturally, fl
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24.4.3 DRIED FLOWER ARRANGEMENTS Dr
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25 Terrarium Culture PURPOSE AND EX
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FIGURE 25-3 Terrarium containers ar
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FIGURE 25-5 Assortment of tools use
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25.6.7 ENHANCING THE DISPLAY Certai
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(a) (b) FIGURE 26-1 Bonsai can be c
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TABLE 26-3 Plant A Selection of Pop
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26.3.1 COLLECTING BONSAI PLANTS FRO
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Strip bark Bare branch FIGURE 26-9
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26.5.2 SANITATION It is critical to
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27 Postharvest Handling and Marketi
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whereas oranges are picked (they ha
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(b) (a) (c) (d) (e1) (e2) (f) FIGUR
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To reduce packaging injury, contain
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is replaced by the by-product of re
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Stored produce may lose some color,
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with pricing. When selling by volum
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(a) (b) FIGURE 27-5 Horticultural p
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APPENDIX A Temperature: Converting
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APPENDIX B Metric Conversion Chart
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APPENDIX D Common and Scientific Na
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Pecan (Carya illinoensis) Peony (Pa
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GLOSSARY A Abaxial Turned away from
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Cellulose A complex carbohydrate th
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Floriculture The science and practi
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M Macronutrient An essential elemen
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Root cap A mass of hard cells cover
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INDEX A-frame, 395 A-horizon, 108 A
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defined, 390 fertilization, 432-434
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Radiant heaters, 378 Radicle, 90 Re
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color plate 1 (a) (b) (c) (d) (e) M
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color plate 3 (b) (a) (c) (d) (e) (
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color plate 5 (a) (b) (d) (c) (e) (
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color plate 7 (b) (c) (d) (a) (e) (
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color plate 9 (a) (b) (c) (d) (e) (
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color plate 11 (a) (c) (b) (d) Grow
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color plate 13 (a) (b) (c) (d) (e)
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color plate 21 (a) (b) (c) (e) (d)
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color plate 23 (c) (b) (a) (d) (e)
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color plate 25 (c) (a) (b) (d) (e)
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color plate 27 (a1) (a2) (b2) (b1)
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color plate 31 (a) (b) (c) Floral d
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Horticulture Principles and Practices

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