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Recognizing Deadly Venomous Snakes from Harmless Snakes of Sri Lanka

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Declaration <strong>of</strong> Our Core<br />

Commitment to Sustainability<br />

Dilmah owes its success to the quality <strong>of</strong> Ceylon Tea. Our business was founded therefore on an enduring<br />

connection to the land and the communities in which we operate. We have pioneered a comprehensive<br />

commitment to minimising our impact on the planet, fostering respect for the environment and ensuring<br />

its protection by encouraging a harmonious coexistence <strong>of</strong> man and nature. We believe that conservation<br />

is ultimately about people and the future <strong>of</strong> the human race, that efforts in conservation have associated<br />

human well-being and poverty reduction outcomes. These core values allow us to meet and exceed our<br />

customers’ expectations <strong>of</strong> sustainability.<br />

Our Commitment<br />

We reinforce our commitment to the principle <strong>of</strong> making business a matter <strong>of</strong> human service and to<br />

the core values <strong>of</strong> Dilmah, which are embodied in the Six Pillars <strong>of</strong> Dilmah.<br />

We will strive to conduct our activities in accordance with the highest standards <strong>of</strong> corporate best<br />

practice and in compliance with all applicable local and international regulatory requirements and<br />

conventions.<br />

We recognise that conservation <strong>of</strong> the environment is an extension <strong>of</strong> our founding commitment to<br />

human service.<br />

Front Cover:<br />

Chrysopelea ornata (Ornate flying snake (E); Pol mal karawala,<br />

Malsara, Panina karawala (S))<br />

One <strong>of</strong> the most vibrantly coloured snakes in <strong>Sri</strong> <strong>Lanka</strong> that<br />

has been aptly named Malsara (u,aird) which in Sinhalese<br />

means ‘cupid’. Unfortunately this harmless snake has become an<br />

innocent victim owing to its colouration (red, black, and yellow),<br />

which symbolise ‘danger’ in the animal world. Commonly found<br />

in the canopy <strong>of</strong> tall trees <strong>of</strong> the wet zone, this is a rare species<br />

in the drier parts <strong>of</strong> the island. Humans living on the periphery<br />

<strong>of</strong> forests are likely to encounter them more <strong>of</strong>ten, as they will<br />

come to feed on geckoes commonly found in human settlements,<br />

and unfortunately when they do, they are <strong>of</strong>ten killed due to<br />

misplaced fears.<br />

Back Cover:<br />

Left to Right Python molurus (Rock python), Echis carinatus<br />

(Saw scaled viper), Eryx conicus (Sand boa)<br />

We will assess and monitor the quality and environmental impact <strong>of</strong> its operations, services and<br />

products whilst striving to include its supply chain partners and customers, where relevant and to<br />

the extent possible.<br />

We are committed to transparency and open communication about our environmental and social<br />

practices.<br />

We promote the same transparency and open communication <strong>from</strong> our partners and customers.<br />

We strive to be an employer <strong>of</strong> choice by providing a safe, secure and non-discriminatory working<br />

environment for its employees whose rights are fully safeguarded and who can have equal<br />

opportunity to realise their full potential.<br />

We promote good relationships with all communities <strong>of</strong> which we are a part and we commit to<br />

enhance their quality <strong>of</strong> life and opportunities whilst respecting their culture, way <strong>of</strong> life and<br />

heritage.


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Citation<br />

Wickramasinghe, L. J. Mendis (2014). Recognising <strong>Deadly</strong> <strong>Venomous</strong> <strong>Snakes</strong> <strong>from</strong><br />

<strong>Harmless</strong> <strong>Snakes</strong> <strong>of</strong> <strong>Sri</strong> <strong>Lanka</strong>. Colombo, <strong>Sri</strong> <strong>Lanka</strong>: Ceylon Tea Services PLC.<br />

Text & Photographs by<br />

L.J. Mendis Wickramasinghe<br />

Assisted by<br />

Nethu Wickramasinghe, Dulan Ranga Vidanapathirana<br />

and Gayan Chathuranga<br />

Design and Layout by<br />

Kasun Pradeepa. Wild Studio<br />

ISBN: 978-955-0081-12-7<br />

Ceylon Tea Services PLC<br />

MJF Group<br />

111, Negombo Road<br />

Peliyagoda<br />

<strong>Sri</strong> <strong>Lanka</strong><br />

Contact<br />

info@dilmahconservation.org<br />

January 2014.<br />

Recognising<br />

<strong>Deadly</strong> <strong>Venomous</strong><br />

SNAKES<br />

<strong>from</strong> <strong>Harmless</strong> <strong>Snakes</strong><br />

<strong>of</strong> <strong>Sri</strong> <strong>Lanka</strong><br />

Printed and bound<br />

Karunaratne & Sons (Pvt)Ltd.<br />

Authored by<br />

L. J. Mendis Wickramasinghe<br />

Advised by Channa Bambaradeniya, Ph.D.<br />

& Gernot Vogel, Ph.D.<br />

Edited by Devaka Weerakoon Ph.D.


Message <strong>from</strong> the Founder<br />

An unfortunate fear and loathing <strong>of</strong> snakes permeates through our society, and<br />

since childhood, I too have been no exception to this disappointing lapse. It is sadly<br />

commonplace for those who encounter snakes to attempt to kill them, owing to<br />

suspicions they harbour that all snakes are poisonous or deadly.<br />

I have learned that these misconceptions which cause panic and revulsion stems<br />

<strong>from</strong> a poor understanding <strong>of</strong> snakes and their habits, coupled with the inability and<br />

unwillingness to distinguish between different species <strong>of</strong> snakes. This results in the<br />

thoughtless killing <strong>of</strong> snakes, even though many types found in <strong>Sri</strong> <strong>Lanka</strong> are largely<br />

harmless and do not attack unless threatened. I have also observed that this has proved<br />

to be a particular challenge within the plantation sector, and tea estates must be no<br />

exception for efforts in biodiversity conservation.<br />

Mr. L. J. Mendis Wickramasinghe has compiled a comprehensive illustrated field guide<br />

on behalf <strong>of</strong> Dilmah Conservation, which not only seeks to help readers easily identify<br />

snakes so that they will be able to deal with these creatures safely and considerately,<br />

but cultivate an interest and appreciation for the vital ecological role they play in<br />

pest control. It is my wish that this field guide will spark the curiosity <strong>of</strong> students in<br />

particular, and motivates them to pursue further learning and contribute towards the<br />

study and conservation <strong>of</strong> reptiles in <strong>Sri</strong> <strong>Lanka</strong>.<br />

I firmly believe that it is the duty <strong>of</strong> Dilmah Conservation to strengthen its commitment<br />

to promoting socially and environmentally conscious educational initiatives towards<br />

fostering awareness, appreciation and respect for <strong>Sri</strong> <strong>Lanka</strong>’s natural wealth. Thus, I<br />

am confident that this publication ‘Recognising <strong>Deadly</strong> <strong>Venomous</strong> <strong>Snakes</strong> <strong>from</strong> <strong>Harmless</strong><br />

<strong>Snakes</strong> <strong>of</strong> <strong>Sri</strong> <strong>Lanka</strong>’ will mark a small but noteworthy contribution by Dilmah<br />

Conservation towards bridging the gaps in knowledge about snakes, minimising<br />

damaging misconceptions and thereby enabling their conservation by helping a broad<br />

audience improve their understanding.<br />

Merrill J. Fernando<br />

Founder – Dilmah Conservation


Recognising<br />

<strong>Deadly</strong> <strong>Venomous</strong><br />

SNAKES<br />

<strong>from</strong> <strong>Harmless</strong> <strong>Snakes</strong><br />

<strong>of</strong> <strong>Sri</strong> <strong>Lanka</strong><br />

Authored by<br />

L. J. Mendis Wickramasinghe<br />

Advised by Channa Bambaradeniya, Ph.D.<br />

& Gernot Vogel, Ph.D.<br />

Edited by Devaka Weerakoon Ph.D.


Foreword<br />

Preface<br />

08<br />

<strong>Snakes</strong> are <strong>of</strong>ten considered deadly and dangerous and therefore persecuted<br />

by humans like no other group <strong>of</strong> animals. Yet snakes are an important part<br />

<strong>of</strong> natural and manmade ecosystems as they prey on many species that we<br />

consider as pests and help to keep their populations under control. One <strong>of</strong><br />

the biggest threats to snakes today is inadvertent killing by humans. Much <strong>of</strong><br />

these killings result due to our widespread fear and loathing towards snakes,<br />

as well as great deal <strong>of</strong> misinformation and misconceptions regarding them.<br />

In <strong>Sri</strong> <strong>Lanka</strong>, as this book clearly indicates, only 21 out <strong>of</strong> the 102 recorded<br />

species <strong>of</strong> snakes are deadly poisonous. Out <strong>of</strong> these only 5 species are<br />

known to cause fatalities in humans. Therefore, most <strong>of</strong> the snakes that we<br />

encounter in life are likely to be harmless. Further, snakes do not ordinarily<br />

attack humans unless provoked, startled or injured. Therefore, we can avoid<br />

the inadvertent killing <strong>of</strong> these useful creatures by becoming more aware<br />

about them. One important aspect <strong>of</strong> awareness is to develop an ability to<br />

identify poisonous snakes. This will not only help save snakes but will also<br />

be useful to save the life <strong>of</strong> a person bitten by a snake, as it is important to<br />

establish the identity <strong>of</strong> the snake before providing treatment to the snake<br />

bite victim. Identification <strong>of</strong> snakes is not a difficult task as most poisonous<br />

snakes show characteristic colour and scale patterns that will enable correct<br />

identification. However, many non poisonous snakes tend to mimic the<br />

colour patterns shown by poisonous species as a form <strong>of</strong> defense <strong>from</strong> their<br />

predators. As this is a trait that has evolved to confuse their predators it is not<br />

surprising that we too would be confused by this, which in this case is not<br />

working towards their favor. This book attempts to educate the reader about<br />

the value <strong>of</strong> snakes as well as how to differentiate poisonous snakes <strong>from</strong> non<br />

poisonous species. I am sure this book will help the reader to lessen their fear<br />

and hatred towards snakes while at the same time develop an appreciation<br />

and respect for their role in our ecosystems.<br />

Pr<strong>of</strong>. Devaka Weerakoon<br />

Department <strong>of</strong> Zoology,<br />

University <strong>of</strong> Colombo<br />

<strong>Snakes</strong> are a group <strong>of</strong> animals which have been especially revered in the <strong>Sri</strong><br />

<strong>Lanka</strong>n context. According to ‘Deepawamsa’, an ancient chronicle <strong>of</strong> <strong>Sri</strong> <strong>Lanka</strong>,<br />

the ‘Naga (Snake)’ people, one <strong>of</strong> the four powerful tribes in <strong>Sri</strong> <strong>Lanka</strong> who<br />

ruled the Northern and Western parts <strong>of</strong> the Island during the 6th century BC to<br />

3rd century BC, were snake worshippers. King Buddhadasa, the only monarch<br />

known to not only possess the skills <strong>of</strong> a physician and a surgeon, but also those<br />

<strong>of</strong> a veterinarian, was fabled to have treated a sick cobra as far back as 340-<br />

368 AD, revealing how compassionate ancient people were towards snakes. The<br />

ruins <strong>of</strong> the ancient cities also speak <strong>of</strong> the harmony between snakes and the<br />

people <strong>of</strong> <strong>Sri</strong> <strong>Lanka</strong>. These include guard-stones with a cobra-king providing<br />

protection to the premises, and seven hooded cobra statues built to protect the<br />

tanks constructed by the ancient Kings. However, changing times and four<br />

centuries <strong>of</strong> colonization have resulted in these ancient traditions being replaced<br />

by a lack <strong>of</strong> tolerance towards snakes. Like in the rest <strong>of</strong> the world, snakes have<br />

now become a feared group <strong>of</strong> animals in <strong>Sri</strong> <strong>Lanka</strong>.<br />

The feeling <strong>of</strong> fear evoked by snakes coupled with an inherent reaction <strong>of</strong><br />

disgust <strong>of</strong>ten results in attempts to kill snakes. This fear could be justified when<br />

considering the number <strong>of</strong> deaths caused annually by poisonous snakes. Further,<br />

superstitions and misconceptions also play a significant role in vilifying these<br />

creatures.<br />

In reality, only few species <strong>of</strong> snakes can actually<br />

cause harm to people. Further, many snakes play<br />

a beneficial role in both natural and manmade<br />

ecosystems, as the natural enemies <strong>of</strong> many<br />

species who can otherwise undergo a massive<br />

population growth and become pests to man.<br />

Therefore, humans must find a way to co-exist<br />

with snakes on this limited land we have. As such,<br />

we must become aware <strong>of</strong> how to co-exist. This<br />

book intends to provide the reader with a broad<br />

overview about snakes in <strong>Sri</strong> <strong>Lanka</strong>, with special<br />

emphasis on deadly poisonous snakes, and their<br />

identification.<br />

Ruins <strong>of</strong> a seven hooded cobra, protecting the water near<br />

Urusita Wewa, Embilipitiya


Contents<br />

10<br />

Acknowledgements<br />

This book was made possible thanks to Dilmah Conservation, who invited<br />

me to write a simple guide for the general public to identify venomous snakes<br />

in <strong>Sri</strong> <strong>Lanka</strong>. The fieldwork was made possible thanks to the Biodiversity<br />

Secretariat <strong>of</strong> the Ministry <strong>of</strong> Environment who provided funds and the<br />

Department <strong>of</strong> Wildlife Conservation who provided the permit to conduct<br />

the research work. My heartfelt appreciation goes to Mr. Dulan Ranga<br />

Vidanapathirana and Mr. Gayan Chathuranga for assisting me while<br />

photographing and handling the snakes in the field. I would also like to thank<br />

Dr. Channa Bambaradeniya for his valuable comments and my colleagues at<br />

the Herpetological Foundation <strong>of</strong> <strong>Sri</strong> <strong>Lanka</strong> (HFS), for various courtesies. I<br />

wish to thank Dr. Gernot Vogel whose invaluable comments undoubtedly<br />

improved the quality <strong>of</strong> this book. I would also wish to extend my gratitude<br />

to Pr<strong>of</strong>. Devaka Weerakoon for the valuable time and effort he took to edit<br />

the text to its final form in which it is presented here. Last but not least to my<br />

dear wife Nethu, for her commitment to making this a success!<br />

Foreword .................................................................................................... 8<br />

Preface ........................................................................................................ 9<br />

Acknowledgements ..................................................................................... 10<br />

1. <strong>Snakes</strong> <strong>of</strong> <strong>Sri</strong> <strong>Lanka</strong> ..................................................................................... 12<br />

1.1 Snake bites in <strong>Sri</strong> <strong>Lanka</strong> ....................................................................... 16<br />

2. What is a Snake? ........................................................................................ 17<br />

3. Identification <strong>of</strong> <strong>Venomous</strong> snakes ............................................................... 20<br />

3.1 Mimicry ........................................................................................... 20<br />

4. Kraits .......................................................................................................... 24<br />

4.1 Bungarus caeruleus (Thel karawala/ Indian krait) ................................. 26<br />

4.2 Non venomous species that mimic the Indian Krait ............................... 28<br />

4.2.1 Lycodon aulicus (Alu radanakaya/ Wolf snake, House snake) .............. 28<br />

4.2.2 Lycodon osmanhilli (Mal radanakaya/ Flowery wolf snake) ................. 30<br />

4.2.3 Lycodon striatus (Kabara radanakaya/ Shaw’s wolf snake) ................... 31<br />

4.3 Bungarus ceylonicus (Mudu karawala/ <strong>Sri</strong> <strong>Lanka</strong>n krait) ...................... 33<br />

4.4 Non venomous species that mimic the <strong>Sri</strong> <strong>Lanka</strong>n Krait ...................... 35<br />

4.4.1 Cercaspis carinata (Dhara radanakaya/ <strong>Sri</strong> <strong>Lanka</strong> wolf snake) ............. 35<br />

4.4.2 Dryocalamus nympha (Geta radanakaya, Geta karawala/ Bridal snake) 37<br />

4.4.3 Dryocalamus gracilis (Meegata radanakaya/ Scarce bridal snake) ......... 38<br />

5. Vipers ........................................................................................................ 39<br />

5.1 Daboia russelii (Thith polanga/ Russell’s viper) ..................................... 40<br />

5.2 Non venomous species that mimic the Russell’s viper ............................ 42<br />

5.2.1 Eryx conicus (Wali pimbura, Kota pimbura / Sand boa) ..................... 42<br />

5.2.2 Python molurus (Dara pimbura, Ran pimbura/ Rock python) ........... 44<br />

5.3. Echis carinatus (Wali polanga/ Saw scaled viper) .................................. 46<br />

5.4. Non venomous species that mimic the Saw scaled viper ......................... 48<br />

5.4.1 Boiga triginata (Garandi mapila, Ran mapila, .................................. 48<br />

Kaeta mapila/ Gamma cat snake)<br />

6. Cobra ......................................................................................................... 50<br />

6.1 Naja naja (Naya or Nagaya/ Indian cobra, Spectacled cobra) ................ 50<br />

Literature cited ................................................................................................. 53


1. <strong>Snakes</strong> <strong>of</strong> <strong>Sri</strong> <strong>Lanka</strong><br />

<strong>Sri</strong> <strong>Lanka</strong>n snake fauna comprise <strong>of</strong> 102 species belonging to 10 families 1-4 .<br />

Out <strong>of</strong> these 102 species, 87 live on land, 14 live in the ocean, and the<br />

remaining one inhabits brackish water. Nearly 49% (50 species) <strong>of</strong> the<br />

snake species found in <strong>Sri</strong> <strong>Lanka</strong> are endemic to the island, or do not occur<br />

naturally anywhere else in the world 5 .<br />

Trimeresurus trigonocephalus (Green pit viper (E); Pala polanga (S)).<br />

Left: An Enhydrina schistosa (Hook nose sea snake (E); Valakkadiya (S)) killed by fishermen.<br />

Right: A close up image showing the typical flat tail <strong>of</strong> sea snakes.<br />

12<br />

The snakes <strong>of</strong> <strong>Sri</strong> <strong>Lanka</strong> can be categorized into four groups, depending on<br />

the lethality <strong>of</strong> their venom. Accordingly, 21 species can be considered as<br />

deadly venomous; five species as moderately venomous and 12 species as<br />

mildly venomous. The remaining 64 species are non-venomous 1-4, 6-10 . This<br />

demonstrates that the majority <strong>of</strong> snake species (63%) are in fact, harmless.<br />

Members <strong>of</strong> the genus Boiga (Cat snakes (E); Mapila (S)), one <strong>of</strong> the most<br />

feared snakes, belong to the mildly venomous group, and its bite causes much<br />

less pain than one inflicted by a hypodermic needle. Moderately venomous<br />

snakes include four species <strong>of</strong> the genus Hypnale (Hump-nosed vipers (E);<br />

Kunakatuwa (S)), and Trimeresurus trigonocephalus (Green pit viper (E); Pala<br />

polanga (S)). Their bites will result in harmful effects such as gangrene,<br />

necrosis, tissue damage, kidney failure, blisters etc. Hypnale hypnale, which<br />

was previously known to be a moderately venomous snake 11 has now been<br />

classified as a deadly venomous snake 12-16 .<br />

Left: Hypnale hypnale (Hump nosed viper (E); Kunakatuwa/ Mukalang thelissa (S).<br />

Right: Close-up <strong>of</strong> the head showing the typical posture <strong>of</strong> hump nosed vipers,<br />

keeps their head slightly angled approximately at 45º.


Likewise, Echis carinatus (Saw-scaled viper (E); Vali polonga (S)) is restricted to<br />

the dry and arid zones <strong>of</strong> the island. Moreover, as the snake is very small in size<br />

(30 cm SVL), the amount <strong>of</strong> venom excreted in a single bite is very small 17 .<br />

Left: Hypnale zara (Zara’s hump nosed viper (E); Pahatharata thelissa (S)).<br />

Right: Close-up <strong>of</strong> the head showing its pointed snout/ rostral appendage, which is<br />

the most relible identification character for all hump nosed vipers in common.<br />

Calliophis melanurus (<strong>Sri</strong> <strong>Lanka</strong> coral snake (E); Depath kaluwa (S)).<br />

A very small snake similar in size to an ink tube <strong>of</strong> a carbon pen.<br />

14<br />

Left: Hypnale nepa (Merrem’s hump-nosed viper (E); Mukalan thelissa/<br />

Mukalang kunakatuwa(S)) Right: Close-up <strong>of</strong> the head.<br />

Out <strong>of</strong> the 21 species <strong>of</strong> snakes considered to be deadly poisonous, fourteen<br />

are sea snakes. These sea snakes are non-aggressive in nature, and since they<br />

are found only in deep waters they hardly interact with humans. Out <strong>of</strong> the<br />

remaining seven deadly poisonous species, two are coral snakes (Calliophis<br />

melanurus and Calliophis haematoetron) incapable <strong>of</strong> inflicting damage to<br />

humans owing to their small size (SVL, 30 cm). Out <strong>of</strong> the five species<br />

remaining, Bungarus ceylonicus (Ceylon krait (E); Mudu karawala (S)), is<br />

non-aggressive in nature and is to some extent an uncommon species.<br />

Therefore in <strong>Sri</strong> <strong>Lanka</strong>, the majority <strong>of</strong> human deaths occur as a result <strong>of</strong> lethal<br />

snake bites caused by the 3 remaining species <strong>of</strong> deadly poisonous snakes:<br />

Bungarus caeruleus, (Indian krait (E); Thel karawala (S)), Naja naja, (Indian<br />

cobra (E); Nagaya (S)), and Daboia russelii (Russell’s viper (E); Thith polanga<br />

(S)) 14, 16, 18-19 .<br />

Due to the lack <strong>of</strong> understanding <strong>of</strong> snakes in <strong>Sri</strong> <strong>Lanka</strong>, they are frequently<br />

killed regardless <strong>of</strong> their identity. Therefore, the ability to identify at least these<br />

three species will save the lives <strong>of</strong> human beings and a vast number <strong>of</strong> innocent<br />

and beneficial snakes. Therefore, the primary aim <strong>of</strong> this book is to help the<br />

reader to correctly identify life-threatening species, and thereby prevent the<br />

futile killing <strong>of</strong> a large proportion <strong>of</strong> harmless snakes.


1.1 Snake Bites in <strong>Sri</strong> <strong>Lanka</strong><br />

16<br />

In <strong>Sri</strong> <strong>Lanka</strong>, close to 37,000 snakebite cases are reported annually to<br />

hospitals 16 , <strong>of</strong> which about 100 cases will result in the death <strong>of</strong> the victim.<br />

Most <strong>of</strong> these deaths occur in rural areas where the patients are brought to<br />

the hospitals at a very late stage due to the lack <strong>of</strong> transportation facilities.<br />

If the patients are given medical attention without delay these deaths can be<br />

avoided.<br />

We are attacked by snakes mostly due to our lack <strong>of</strong> understanding <strong>of</strong> what<br />

their preferred habitats might be. Apart <strong>from</strong> their naturally occurring<br />

habitats like termite mounds and bandicoot tunnels, venomous snakes like<br />

Cobras, Russell’s vipers and Kraits are attracted to places such as piles <strong>of</strong> rock<br />

used for construction work where there are plenty <strong>of</strong> empty spaces for them<br />

to hide, garbage mounds, stacked bricks, piled up coconut leaves, coconut<br />

roots, wood piles etc. In other words, while destroying their natural habitats<br />

we too are unintentionally recreating ideal habitats for them. Additionally,<br />

we also create ideal conditions for their prey, such as rats. Therefore, it is not<br />

a real surprise when snakes, like all other living beings, are attracted to places<br />

where there is plenty <strong>of</strong> food and shelter.<br />

The human-snake conflict is an issue that will continue to escalate due to<br />

the ever increasing human population and the consequent loss <strong>of</strong> natural<br />

habitat. However, the threat <strong>from</strong> snakes to humans remains comparatively<br />

low (approximately 100 deaths per year compared to other hazards such as<br />

accidents involving vehicles which claimed 2,721 lives and injured another<br />

26,847 persons in 2010 alone). Despite this, we are not afraid to travel<br />

on the roads, in a vehicle as we are aware <strong>of</strong> the different risks associated<br />

with modes <strong>of</strong> transportation. However, our fear <strong>of</strong> the unknown (such as<br />

our inability to differentiate a commonly encountered harmless snake <strong>from</strong><br />

a deadly venomous snake or our lack <strong>of</strong> understanding about their ways)<br />

drives us to kill them on sight.<br />

2. What is a Snake ?<br />

Many people fear snakes because they believe that:<br />

• All snakes are venomous and are seeking to kill humans.<br />

• <strong>Snakes</strong> have an unpleasant skin which is either sticky and slimy or scaly;<br />

• <strong>Snakes</strong> are dirty and unclean.<br />

However, it should be noted that many people reach the aforementioned<br />

conclusions without even having touched a snake or closely examining them. In<br />

reality, when a person is given the opportunity to closely inspect a snake, they<br />

become pleasantly surprised when they learn that these assumptions are baseless.<br />

• Majority <strong>of</strong> snakes are non-venomous and attack only if they feel threatened,<br />

in order to catch their prey, and not because they enjoy it. It needs to be<br />

understood that snakes play an important role in maintaining the ecological<br />

balance by controlling the population size <strong>of</strong> many species.<br />

• <strong>Snakes</strong> in fact possess a clean, dry skin.<br />

• Some snakes are in fact very colourful and beautiful.<br />

However, these animals have gained an unpopular reputation due to various<br />

reasons, and as a result they are killed by people who are unaware <strong>of</strong> their ecological<br />

importance. They are an integral part <strong>of</strong> a balanced ecosystem and play a significant<br />

ecological role as predators, thereby controlling populations <strong>of</strong> many species<br />

including pests such as rodents. <strong>Snakes</strong> are also good indicators <strong>of</strong> environmental<br />

pollution. An additional benefit <strong>of</strong> these creatures is that their venom,


which contains proteins, can be extracted<br />

for medicinal purposes, hence possessing<br />

an economic value.<br />

<strong>Snakes</strong> come in a diverse range <strong>of</strong> sizes,<br />

and are entirely carnivorous typically<br />

swallowing their prey whole. Depending<br />

on their size, their prey also varies <strong>from</strong><br />

small insects consumed by small earth<br />

snakes like Typhlops to heavy animals<br />

consumed whole by large pythons.<br />

Left: The scale arrangement <strong>of</strong> Cylindrophis maculate<br />

(<strong>Sri</strong> <strong>Lanka</strong> pipe snake (E); Depath naya (S)), a<br />

harmless burrowing snake. The image shows its tail<br />

rolled up, mimicking a hood <strong>of</strong> a cobra a typical<br />

defense mechanism against predators.<br />

Above: A Green pit viper, with wide open mouth<br />

showing its fangs.<br />

18<br />

All snakes have razor sharp teeth, which are used to prevent their prey <strong>from</strong><br />

escaping., They do not use their teeth to break their prey into smaller parts or<br />

for chewing since they swallow their prey whole. Fangs are specialized teeth<br />

that are only found in venomous snakes. These fangs are tube-like and makes<br />

it possible for the venom to be introduced efficiently into a wound once the<br />

snake strikes, thus immobilizing the prey in the shortest possible time.<br />

The venom and its delivery system are special tools they have developed<br />

through years <strong>of</strong> evolution, and it must be emphasized that their main<br />

purposes are to aid the capture <strong>of</strong> prey and self-defense (because snakes are<br />

both predators and prey), and are not intended specifically to hurt humans as<br />

many seem to believe. <strong>Snakes</strong> that lack venom resort to constriction in order<br />

to immobilize their prey.<br />

Left: Coeloganthus helena (Trinket snake (E);<br />

Katakaluwa (S)), a slightly venomous snake, with<br />

its mouth wide open showing its teeth. The orange<br />

arrow shows its fangs positioned towards the rear<br />

<strong>of</strong> its mouth, such fangs are known as back fangs.<br />

As mentioned before, not all snakes<br />

are venomous. The composition <strong>of</strong> the<br />

venom and its quantities vary <strong>from</strong><br />

species to species. In general it is a<br />

complex mixture <strong>of</strong> proteins which can<br />

be categorized according to the organs<br />

they attack i.e. neurotoxic (attacks<br />

the nervous system) or haemototoxic<br />

(attacks the circulatory system).<br />

Depending on their ability to kill their<br />

prey and the composition <strong>of</strong> venom they<br />

possess, snakes have been grouped in to<br />

four categories i.e. deadly venomous,<br />

moderately venomous, mildly venomous<br />

and non-venomous.


Can you identify them?<br />

Saw scaled viper and Gamma cat snake<br />

3. Identification <strong>of</strong><br />

<strong>Venomous</strong> <strong>Snakes</strong><br />

Which one is deadly?<br />

Russell’s viper and Pythons<br />

20<br />

In the animal world, different species use various techniques to survive. These<br />

include their need to find food and defense mechanisms geared to combat<br />

predation. <strong>Snakes</strong> have the ability to do this quite successfully, even to the<br />

extent <strong>of</strong> deceiving us human beings.<br />

3.1 Mimicry<br />

The most common mechanism used by snakes is called mimicry. Where<br />

non-venomous species mimic or superficially look like (casually resemble) a<br />

venomous snake, it serves as a warning to others, especially their predators,<br />

that they are venomous and must be avoided.<br />

Only one <strong>from</strong> these three are deadly venomous, which one is it?


Kraits and<br />

Wolf snakes<br />

15<br />

Although these species look<br />

quite similar, not all <strong>of</strong> them<br />

are venomous because the<br />

non-venomous ones mimic the<br />

venomous ones i.e. harmless<br />

species mimic<br />

the colour<br />

patterns <strong>of</strong><br />

deadly<br />

venomous<br />

snakes.


4. Kraits<br />

In <strong>Sri</strong> <strong>Lanka</strong>, there are two species <strong>of</strong> Kraits, the Indian krait and the Ceylon<br />

krait. They are both deadly venomous. Interestingly, there are six harmless<br />

species that mimic these kraits, including wolf snakes and bridal snakes.<br />

The best and the easiest character that can be used to differentiate the venomous<br />

Kraits <strong>from</strong> the non-venomous mimics is the dorsal scale row shown in the<br />

figure below.<br />

<strong>Venomous</strong> Kraits<br />

Large hexagonal scales running<br />

down the spine<br />

Non - venomous mimics<br />

All scales on back identical in size<br />

24<br />

Both snakes shown above display a common form <strong>of</strong> behavior seen among snakes, whereby<br />

they conceal their heads under their bodies as a defense mechanism. Above: Indian krait that is<br />

deadly venomous. Below: <strong>Sri</strong> <strong>Lanka</strong> wolf snake which is a non venomous species.


Prominent white cross bars in a juvenile.<br />

Simple identification character: Large hexagonal shaped scales running down<br />

its spine. Scales on its back are smooth, with the mid row much larger than all<br />

the other surrounding scales. This feature is the most important characteristic<br />

when identifying the species.<br />

Adult specimen, where the cross bars or white lines have reduced and scattered.<br />

4.1 Bungarus caeruleus<br />

Indian krait (E);Thel karawala (S)<br />

26<br />

Toxicity: <strong>Deadly</strong> venomous<br />

Size: 25 cm- 140 cm<br />

Colouration: Back bluish black to a pale faded bluish grey, with white<br />

cross bars occurring in pairs which become less distinct at the anterior end.<br />

These cross bars are prominent in juveniles or young animals and the lines<br />

gradually disappear or become reduced to scattered cross bars in adults. The<br />

ventral side, or the underside is <strong>of</strong>f-white.<br />

Left: Clear pairs <strong>of</strong> cross bars <strong>of</strong> a juvenile. Right: The cross bars gradually diminishing in adults.<br />

But both show enlarged hexagonal scale rows along the spine.<br />

Distribution: Non-endemic. Distributed in the dry, arid, and intermediate<br />

zones.<br />

Behavior: : It is a nocturnal species that is aggressive at night. Commonly found<br />

in and around human settlements. May attack if threatened but generally nonaggressive<br />

during day time. Will roll into a ball with its head well-concealed<br />

when agitated, and might spring out <strong>from</strong> this position upon further agitation.<br />

Feeds on: Other snakes, geckoes, lizards, rodents etc.<br />

Reproduction: They lay 10- 16 eggs per clutch.


4.2 Non venomous species that<br />

mimic the Indian Krait<br />

Three harmless Wolf snakes mimic the Indian Krait.<br />

Juvenile specimen.<br />

Scales on the back are all identical, and smooth. The subcaudal scales on the<br />

underside on its tail are all divided.<br />

28<br />

4.2.1 Lycodon aulicus<br />

Toxicity: Non venomous<br />

Size: 18 cm- 80 cm<br />

Wolf snake, House snake (E); Alu radanakaya (S)<br />

Adult specimen.<br />

Colouration: Back dark brown to blackish brown, with white cross bars that<br />

divide laterally which are prominent in the anterior end. These cross bars<br />

are prominent in juveniles and young, and the lines gradually disappear or<br />

become reduced to scattered cross bars in adults. Lips are prominently white,<br />

while the ventral or underside is <strong>of</strong>f-white.<br />

Distribution: Non-endemic. Distributed in all parts <strong>of</strong> the island up to 2000 m<br />

asl. Commonly found in rural areas close to forests, in houses (especially among<br />

piles <strong>of</strong> wood, piles <strong>of</strong> stone, foundations and walls where there are plenty <strong>of</strong><br />

cracks/crevices), stacked bricks, piled up coconut leaves, and coconut roots etc.<br />

Behavior: : It is a nocturnal species that is aggressive at night and attacks fiercely.<br />

Will roll in to a ball with its head well concealed when agitated, and empty their<br />

bowels with a smell similar to that <strong>of</strong> rotten dead mice (hence the Sinhala name<br />

“Kunu mee karawala”) as a defense mechanism.<br />

Feeds on: Geckoes, lizards, small rodents etc.<br />

Reproduction: They lay about 20 eggs.


Distribution: This species is endemic to <strong>Sri</strong> <strong>Lanka</strong>. Distributed in all parts <strong>of</strong><br />

the island up to 2000 m asl. It is commonly found in urbanized and semiurbanized<br />

areas, in houses (especially under flower pots, piles <strong>of</strong> stone, loose<br />

soil, in foundations and walls where there are plenty <strong>of</strong> cracks/crevices), stacked<br />

bricks, piled up coconut leaves, coconut roots, piles <strong>of</strong> wood and in piled up<br />

goods such as clothes, books, boxes etc.<br />

Behavior: It is a nocturnal species that is aggressive at night and attacks- and<br />

bites fiercely. Will roll in to a ball with its head well concealed when agitated,<br />

and empty their bowels with a smell similar to rotten dead mice. Hence, in<br />

Sinhala, it is called “Kunu mee karawala”, owing to the particularity <strong>of</strong> its<br />

defense mechanism.<br />

Feeds on: geckoes, lizards, small rodents etc.<br />

Reproduction: They lay 8-10 eggs per clutch.<br />

4.2.3 Lycodon striatus<br />

Shaw’s wolf snake (E); Kabara radanakaya (S)<br />

4.2.2 Lycodon osmanhilli<br />

Flowery wolf snake (E); Mal radanakaya (S)<br />

Toxicity: Non venomous<br />

30<br />

Size: 15 cm- 60 cm<br />

Colouration: They show two variations in colour. One morph has a light<br />

brown to a yellowish tinge, with divided yellow cross bars throughout its<br />

back. The other has a uniform light brown coloured body with no white<br />

markings on the back. In both cases the head colouration is light brown with<br />

a yellowish tinge. Eyes black and prominently seen because <strong>of</strong> the lighter skin<br />

colour compared to others, with white lips upon which every lip scale has a<br />

brown mark in the center.<br />

Scales on the back are all identical, and smooth. The subcaudal scales on<br />

the underside on its tail are all divided.


Toxicity: Non venomous<br />

Size: 9 cm- 28 cm<br />

Colouration: The back is dark brown to blackish brown in colour, with white<br />

cross bars which are prominent at the anterior end that divide laterally and<br />

are broken. In some, these cross bars are yellow. Its lips are white, however<br />

not as distinctly as L. aulicus.The ventral or the underside is <strong>of</strong>f-white.<br />

Scales on the back are all identical, and smooth. The subcaudal scales on<br />

the underside on its tail are all divided.<br />

Distribution: Non-endemic. Distributed in all parts <strong>of</strong> the island up to 2000<br />

m asl. Commonly found in houses (especially in piled up goods, in home<br />

gardens, mounds <strong>of</strong> stone, foundations and walls where there are plenty <strong>of</strong><br />

cracks/crevices), stacked bricks, piled up coconut leaves, coconut roots, and<br />

piles <strong>of</strong> timber etc.<br />

Behavior: It is a nocturnal species that is non-aggressive and does not attack.<br />

Will roll in to a ball with its head well concealed when agitated.<br />

Feeds on: Small skink and lizards.<br />

Reproduction: They lay 5-6 eggs per clutch.<br />

An adult specimen showing the typical banding pattern and enlarged scale row on the spine.<br />

4.3 Bungarus ceylonicus<br />

<strong>Sri</strong> <strong>Lanka</strong>n krait (E); Mudu karawala (S)<br />

Toxicity: <strong>Deadly</strong> venomous<br />

Size: 100- 110 cm<br />

32<br />

Colouration: Back jet black, with white cross bars. These cross bars are prominent<br />

in juveniles/young and the lines gradually disappear or become reduced to<br />

scattered cross bars in adults. Ventral or the underside is black or alternately black<br />

Juvenile specimen with<br />

prominent white cross<br />

bars. The posterior half<br />

<strong>of</strong> the head is white<br />

coloured.


and white. Posterior half <strong>of</strong> the head in juveniles is white. However the white<br />

colour diminishes gradually with age.<br />

Simple identification: Large hexagonal shaped scales running down its spine.<br />

Scales smooth on back. They are much larger than all the other surrounding<br />

scales.<br />

4.4 Non venomous species that<br />

mimic the <strong>Sri</strong> <strong>Lanka</strong>n Krait<br />

There are three harmless snakes (<strong>Sri</strong> <strong>Lanka</strong>n wolf snake and two bridal snakes)<br />

that mimic the <strong>Sri</strong> <strong>Lanka</strong>n krait.<br />

Left: Faded white cross bars in an adult. Right: Prominent white cross bars in a juvenile.<br />

34<br />

Distribution: Wet, intermediate and rarely seen in the dry zone.<br />

Behavior: It is a nocturnal species that is aggressive at night. Commonly<br />

found in and around human settlements. Potentially non-aggressive in<br />

nature, but may attack if provoked or threatened. Will roll in to a ball with its<br />

head well concealed when agitated, and will remain still until the perceived<br />

danger passes. They display this behavior much better than the Indian Krait.<br />

Feeds on: Other snakes, geckoes, lizards and rodents etc.<br />

Reproduction: They lay 6-10 eggs per clutch.<br />

4.4.1 Cercaspis carinata<br />

<strong>Sri</strong> <strong>Lanka</strong> wolf snake (E); Dhara radankaya (S)<br />

Toxicity: Non venomous<br />

Size: 65 cm<br />

An adult specimen.<br />

Colouration: Head jet black and shiny, the body is black with white cross bars.<br />

These cross bars are prominent in juveniles/young and as they become adults the<br />

lines gradually disappear or become reduced to scattered cross bars. Ventral or<br />

the underside is black or alternately black and white. Posterior half <strong>of</strong> the head<br />

in juveniles is white and gradually diminishes with age.


A Juvenile specimen showing prominent cross bars and the<br />

posterior part <strong>of</strong> the head is white coloured.<br />

36<br />

Simple identification: Scales on the back are deeply ridged, keeled or rough,<br />

a character evident only in this species out <strong>of</strong> those that mimic kraits.<br />

Scales on its back are all identical, and deeply ridged. scales on the underside<br />

on its tail or subcaudal scales are all single or undivided.<br />

Distribution: Found only in the wet zone.<br />

Behavior: It is a nocturnal species that is aggressive at night. Commonly<br />

found under leaf litter <strong>of</strong> the forest floor and in home gardens close to forests.<br />

May attack if provoked or threatened. Will roll in to a ball with its head well<br />

concealed when agitated, and will remain the same till the perceived danger<br />

passes.<br />

Feeds on: Geckoes, lizards, skinks, small rodents and rarely on other snakes<br />

etc.<br />

Reproduction: They lay 8-12 eggs per clutch.<br />

4.4.2 Dryocalamus nympha<br />

Bridal snake (E); Geta radankaya, Geta karawala (S)<br />

Toxicity: Non venomous<br />

Size: 70 cm<br />

Colouration: The back <strong>of</strong> the animal is black with white cross bars which may<br />

be yellowish in juveniles. These cross bars are prominent in juveniles/young and<br />

the lines gradually disappear or become reduced to scattered cross bars in adults.<br />

Posterior cross bands are connected laterally. Ventral or the underside is white.<br />

Simple identification: Possesses a comparatively larger head with a slimmer<br />

neck area. Its body is adapted to climb trees, with a long tail and prominent<br />

ventral ridges on either side.<br />

Scales on the back are all identical and smooth. Underside scales on tail or the<br />

subcaudal scales are all divided.<br />

Distribution: Found in the dry and intermediate zones.<br />

Behavior: It is a nocturnal non-aggressive species. Commonly found in large tree<br />

trunks and under debris, in and around home gardens or human settlements.<br />

Will roll in to a ball with its head well concealed when agitated.


Feeds on: Skinks and eggs <strong>of</strong> geckoes etc.<br />

Reproduction: They lay eggs. But the number is not known, but a gravid<br />

female specimen with 5 eggs (upon external observation) has once been<br />

recorded.<br />

4.4.3 Dryocalamus gracilis<br />

Scarce bridal snake (E); Meegata radankaya (S)<br />

5. Vipers<br />

<strong>Sri</strong> <strong>Lanka</strong> has two species <strong>of</strong> true vipers (Russell’s viper and Saw scaled viper).<br />

They are both deadly venomous, but very <strong>of</strong>ten people get these confused with<br />

non-venomous pythons.<br />

The best and the easiest characters to discriminate the venomous Russell’s viper<br />

<strong>from</strong> the two non-venomous pythons (Rock python/ Indian python and Sand<br />

Boa) are listed in the table below.<br />

Distinguishing characters Russell’s viper Rock Python Sand Boa<br />

Markings on back<br />

Regular, oval<br />

Irregular, cloud shaped<br />

Irregular, cloud shaped<br />

Distribution <strong>of</strong> scales on head<br />

Large number <strong>of</strong><br />

small sized scales<br />

Both large and<br />

small sized<br />

Small sized, large<br />

number<br />

Head scales<br />

Rough<br />

Smooth<br />

Rough<br />

A preserved museum specimen <strong>of</strong> Scarce bridal snake which was used to describe the species<br />

scientifically, deposited at the Natural History Museum London.<br />

Toxicity: Non venomous<br />

Snout shape <strong>from</strong> top<br />

Labial pits<br />

Comparatively pointed<br />

Absent<br />

Blunt<br />

Present<br />

Blunt<br />

Absent<br />

38<br />

Size: 50 cm<br />

Remarks: A very rare species, only known <strong>from</strong> a single record.<br />

Colouration: Back, black with white cross bars. Similar to Dryocalamus<br />

nympha.<br />

Simple identification: Can only be separated by Dryocalamus nympha, by<br />

the number <strong>of</strong> costal scales. Where D. nympha, has 13 scales and D. gracilis<br />

has 15 scales.<br />

Distribution: Single record <strong>from</strong> Colombo, wet zone.<br />

Mid body scales on back<br />

Tail scales on back<br />

Sub-caudal/under side <strong>of</strong><br />

tail scales<br />

Vestigial hind limbs<br />

Distribution<br />

Rough/keeled<br />

Rough/keeled<br />

Divided<br />

Absent<br />

< 2000 m asl<br />

Smooth<br />

Smooth<br />

Single<br />

Present<br />

< 2000 m asl<br />

Smooth<br />

Rough/keeled<br />

Single<br />

Present<br />

Arid zone and coastal<br />

dry zone<br />

Behavior: Unknown Feeds on: Unknown Reproduction: Unknown


The row <strong>of</strong> spots on back <strong>of</strong> a Russell’s viper which are comparatively large and oval<br />

shaped outlined with a black line, with its outer border whitish.<br />

Simple identification: From the large oval spots that run down the body, and<br />

the keeled costal scales. The pair <strong>of</strong> scales over the eye or the supra ocular, and the<br />

supra nasal scales are large. This species can be identified <strong>from</strong> its loud hissing<br />

noise.<br />

5.1 Daboia russelii<br />

Russell’s viper (E); Thith polanga (S)<br />

Toxicity: <strong>Deadly</strong> venomous<br />

40<br />

Size: 120-130 cm<br />

Colouration: Earthy brown with three rows <strong>of</strong> dark brown spots on the back,<br />

one running down the spine and the other two on the sides <strong>of</strong> body. The row<br />

<strong>of</strong> spots on its back are comparatively large and oval shaped, outlined with<br />

a black line, with its outer border whitish, while those found on the lateral<br />

body are small and somewhat oval. On its head a pair <strong>of</strong> dark brown patches<br />

forms a light brown ‘V’ pointing toward the snout. Ventral or the underside<br />

is <strong>of</strong>f white with black or dark brown blotches.<br />

Head close up <strong>of</strong> Russell’s viper showing a pair <strong>of</strong> dark brown patches, which forms<br />

a light brown ‘V’ pointing toward the snout.<br />

Distribution: Found both in the wet and the dry zones < 2000 m asl.<br />

Behavior: It is a nocturnal species that is aggressive at night. Commonly found<br />

in forests, in and around human settlements, paddy fields, under piled-up goods,<br />

and coconut husks etc. If provoked or threatened, it will coil and raise a third<br />

<strong>of</strong> its body and produce a loud hissing noise which is the loudest <strong>of</strong> all snakes.


Feeds on: Adults feed on rodents and birds, while juveniles may feed on<br />

geckoes, lizards, frogs etc.<br />

Reproduction: They give birth to 30-35 live young.<br />

5.2 Non venomous species that<br />

mimic the Russell’s viper<br />

Toxicity: Non venomous<br />

Size: Female-100 cm, males-60 cm<br />

Colouration: Earthy brown with large dark brown blotches on back which are<br />

connected but may be separated. Much smaller blotches are found on the side<br />

<strong>of</strong> its body, which are randomly distributed. Ventral or the underside is pinkish<br />

in colour.<br />

The two non-venomous pythons (rock python and sand boa) superficially<br />

look a lot like a Russell’s viper. Out <strong>of</strong> which the sand boa is more similar but<br />

is less commonly found.<br />

5.2.1 Eryx conicus<br />

Sand boa (E); Wali pimbura, Kota pimbura (S)<br />

Mid body colour pattern <strong>of</strong> a Sand Boa.<br />

Simple identification: Smooth mid body scales, and the blotches on its back.<br />

Scales on the back <strong>of</strong> its tail are deeply ridged (keeled/rough) The subcaudal<br />

scales on the underside <strong>of</strong> its tail are single or undivided.<br />

43<br />

A close up view <strong>of</strong> the head <strong>of</strong> a Sand boa showing its<br />

rough and small even sized scales.


Distribution: Found only in the coastal dry zone and arid zone.<br />

Behavior: It is a nocturnal species that moves slowly and is lethargic. It is<br />

rare but can be found burrowed within soil in dry and arid parts <strong>of</strong> the<br />

island. They constrict their prey to immobilize them.<br />

Feeds on: small mammals, geckoes, lizards, skinks, frogs etc.<br />

Reproduction: They give birth to live young.<br />

5.2.2 Python molurus<br />

Rock python (E); Dara pimbura, Ran pimbura (S)<br />

Large sized blotches on mid body.<br />

Simple identification: The pattern <strong>of</strong> blotches on its body. The subcaudal scales<br />

on the underside <strong>of</strong> its tail are single or undivided.<br />

44<br />

Close up <strong>of</strong> head showing the markings <strong>of</strong> an arrow pointing towards head, and<br />

scales <strong>of</strong> different sizes.<br />

Toxicity: Non venomous<br />

Size: 600 cm<br />

Colouration: Whitish or yellowish with three rows <strong>of</strong> large sized blotches<br />

on back. Colours vary <strong>from</strong> a light brownish tinge to a dark brown. The row<br />

<strong>of</strong> blotches on the back is much larger than the rows on the side <strong>of</strong> body.<br />

Ventral or the underside is cream with small dark brown or black spots.<br />

Distribution: < 2000 m asl<br />

Behavior: A slow moving nocturnal species, but if threatened will move fast to<br />

hide. If threatened they will also coil and warn by hissing several times before<br />

attempting to bite. It is a highly camouflaged species that lives amongst leaf<br />

litter, and well-adapted as an ambush predator. Uncommon but can be found in<br />

forests, and in semi-aquatic environments. It is also well-adapted to climb trees.<br />

They constrict their prey to immobilize them.


Toxicity: <strong>Deadly</strong> venomous<br />

Size: 30-40 cm<br />

Colouration: Light brown or pale earthy brown with a series <strong>of</strong> dark brown<br />

spots outlined with whitish lines on back. On its head there is a distinct <strong>of</strong>f-white<br />

marking which may look like an arrow, or the foot print <strong>of</strong> a bird, or a crucifix.<br />

Ventral or the underside is <strong>of</strong>f white to light brown, with small dark brown spots.<br />

The vestigial<br />

hind limbs <strong>of</strong><br />

Python mollurus<br />

(Rock Python).<br />

Feeds on: Large to small mammals (except those that are carnivorous and<br />

very large species), flying mammals, small sized crocodiles, land monitors,<br />

birds etc. They constrict their prey to immobilize them.<br />

Reproduction: They lay 12- 40 eggs, and are the only snakes in <strong>Sri</strong> <strong>Lanka</strong><br />

that protect and incubate their eggs.<br />

5.3 Echis carinatus<br />

Saw scaled viper (E); Wali polanga (S)<br />

Distinct marking on the head <strong>of</strong> Saw scaled Viper, which may look like an arrow, a<br />

foot print <strong>of</strong> a bird or a crucifix.<br />

Simple identification: The <strong>of</strong>f-white distinct marking on the head.<br />

Costal scales are keeled/rough and identical in size except for the pair just above<br />

the eye (supra ocular) and inter-nasals, which are smooth and larger in size. The<br />

subcaudal scales on the underside <strong>of</strong> its tail are single or undivided.<br />

46<br />

Distribution: Found in the arid and the coastal dry zone.<br />

Behavior: It is an aggressive nocturnal species that is rarely diurnal. When<br />

threatened they coil and make a small warning noise by rubbing their body coils<br />

against each other. It is a highly camouflaged species in its natural habitat which<br />

is sand-mixed dry leaf litter. Potentially aggressive and will attack if provoked.<br />

Has limited distribution, hence uncommon but commonly found in its preferred<br />

habitat, the sand dune forests and surrounding home gardens. During the dry<br />

season, they are commonly seen under palmyrah leaves (Borassus flabellifer),<br />

which have fallen close to wells and places where there is water.<br />

Feeds on: Geckoes, lizards, frogs, small rodents etc.<br />

Reproduction: They give birth to live <strong>of</strong>f spring.


5.4 Non venomous species that<br />

mimic the Saw scaled viper<br />

5.4.1 Boiga triginata<br />

Gamma cat snake (E); Garandi mapila, Ran mapila,<br />

Kaeta mapila (S)<br />

Toxicity: Non venomous<br />

Size: 70-100 cm<br />

Colouration: Light brown or pale earthy brown with a series <strong>of</strong> <strong>of</strong>f white ‘V’<br />

shaped markings outlined in dark brown or black running down its spine. On its<br />

head there is a distinct <strong>of</strong>f whitish ‘Y’ shaped marking. Ventral or the underside<br />

is light yellow, with small dark brown spots on either side.<br />

Distinct ‘Y’ shaped marking on the head <strong>of</strong> a Gamma cat snake.<br />

48<br />

Simple identification: The <strong>of</strong>f whitish distinct ‘Y’ shaped marking on head.<br />

Underside scales on tail or subcaudal scales are divided.<br />

Distribution: Found in the arid, dry and intermediate zones.<br />

Behavior: It is a nocturnal species. When threatened they make a warning noise<br />

by patting its tail on to a surface. Non aggressive, and can be commonly found<br />

on trees, piled up goods, and in and around human settlements etc.<br />

Feeds on: geckoes, lizards, skinks, small birds, small sized bird’s eggs, small<br />

rodents etc.<br />

Reproduction: They lay 8-12 eggs per clutch.


6. Cobra<br />

There is only one species <strong>of</strong> cobra (Indian cobra or Spectacled cobra) found<br />

in <strong>Sri</strong> <strong>Lanka</strong>. It is a deadly venomous species, and is the most commonly<br />

known venomous snake, amongst people in the island.<br />

6.1 Naja naja<br />

Indian cobra, Spectacled cobra (E); Naya, Nagaya (S)<br />

Toxicity: <strong>Deadly</strong> venomous<br />

Size: 180 cm<br />

Colouration: The colour on its back varies <strong>from</strong> dark brown to dark reddish<br />

brown or dark grey to grayish black with white or light yellow cross bars. These<br />

cross bars are either incomplete or complete with sets <strong>of</strong> four to six bands<br />

grouped at times. Hood contains a large usually white-coloured spectacle shaped<br />

marking, and the surrounding area is reddish. The ventral or the underside may<br />

vary <strong>from</strong> white to pale yellow to light brown, while some may have a single<br />

colouration, others may have cross bars or blotching.<br />

50<br />

The prominent, large<br />

spectacle shaped marking<br />

on the hood <strong>of</strong> a cobra<br />

which is commonly<br />

white.<br />

Simple identification: It forms a distinctive hood, raising its body and displaying<br />

the prominent spectacle shaped marking on the head. When the hood has not<br />

expanded, the easiest way to identify this snake is by the observing its rostral<br />

scale, the area below the eye which is outlined in black.<br />

The subcaudal scales on the underside <strong>of</strong> its tail are divided.


Distribution: < 2000 m asl.<br />

Behavior: It is a diurnal species that is rarely nocturnal (but large sized cobras<br />

may be encountered at night very rarely). Commonly found in and around<br />

human settlements, paddy fields, under piled-up goods, piled-up coconut<br />

husks etc. A potentially non-aggressive species, but if threatened or provoked<br />

will expand its hood and warns by its fake attacks.<br />

Feeds on: Small mammals, frogs, lizards, monitors, other snakes, birds and<br />

their eggs etc. Juveniles may consume skinks, geckoes etc.<br />

Reproduction: They lay 20-40 eggs per clutch.<br />

Remarks: Juvenile cobras could be confused with juvenile rat snakes.<br />

Literature cited<br />

1. Smith, E., Manamendra-Arachchi, K. and Somaweera, R. (2008). A new<br />

species <strong>of</strong> coralsnake <strong>of</strong> the genus Calliophis (Squamata: Elapidae) <strong>from</strong><br />

the Central Province <strong>of</strong> <strong>Sri</strong> <strong>Lanka</strong>. Zootaxa. 1847:19–33.<br />

2. Wickramasinghe, L.J.M., Vidanapathirana, D.R., Wickramasinghe,<br />

N. and Ranwella, P.N. (2009). A New Species <strong>of</strong> Rhinophis Hemprich,<br />

1820 (Reptilia: Serpentes: Uropeltidae) <strong>from</strong> Rakwana massif, <strong>Sri</strong> <strong>Lanka</strong>.<br />

Zootaxa. 2044: 1–22.<br />

52<br />

People <strong>of</strong>ten get confused between the more widespread and harmless Rat snakes<br />

with cobras. The rat snake plays a very important role as a pest controller.<br />

A close-up view <strong>of</strong> the head <strong>of</strong> a Rat Snake showing black lines outlined on scales<br />

bordering the lip, which is the simplest feature that can be used for identification<br />

<strong>of</strong> rat snakes.<br />

3. Maduwage K., Silva A., Manamendra-Arachchi K. and Pethiyagoda R.<br />

(2009). A taxonomic revision <strong>of</strong> the South Asian hump-nosed pit vipers<br />

(Squamata: Viperidae: Hypnale). Zootaxa. 2232:1–28.<br />

4. Gower, D.J. and Maduwage, K. (2011). Two new species <strong>of</strong> Rhinophis<br />

Hemprich (Serpentes: Uropeltidae) <strong>from</strong> <strong>Sri</strong> <strong>Lanka</strong>. Zootaxa. 2881:<br />

51–68.<br />

5. Wickramasinghe, L.J.M., Conservation Status <strong>of</strong> the Reptile Fauna <strong>of</strong> <strong>Sri</strong><br />

<strong>Lanka</strong>. The 2012 Red List <strong>of</strong> Threatened Fauna and Flora <strong>of</strong> <strong>Sri</strong> <strong>Lanka</strong>. (Un<br />

der review)<br />

6. de Silva, P.H.D.H. (1980). Snake Fauna <strong>of</strong> <strong>Sri</strong> <strong>Lanka</strong> – with special<br />

reference to skull, dentition and venom in snakes. National Museums <strong>of</strong> <strong>Sri</strong><br />

<strong>Lanka</strong>. Colombo. xi + 472 pp.<br />

7. Das, I. and de Silva, A. (2005) A photographic guide to snakes and other<br />

reptiles <strong>of</strong> <strong>Sri</strong> <strong>Lanka</strong>. New Holland publishers (UK). 144 pp.<br />

8. Somaweera, R. (2006) <strong>Sri</strong> <strong>Lanka</strong>we Sarpayin [in Sinhalese; “<strong>Snakes</strong> <strong>of</strong> <strong>Sri</strong><br />

<strong>Lanka</strong>”]. WHT Publications, Colombo, <strong>Sri</strong> <strong>Lanka</strong> x + 297 pp.<br />

9. Rooijen, J.V. and Vogel, G. (2008). An investigation into the taxonomy<br />

<strong>of</strong> Dendrelaphis tristis (Daudin, 1803): revalidation <strong>of</strong> Dipsas schokari<br />

(Kuhl, 1820) (Serpentes, Colubridae). Contributions to Zoology. 77(1):<br />

29–39.<br />

10. Vogel, G. and Rooijen, J.V. (2011). A new species <strong>of</strong> Dendrelaphis (Ser<br />

pentes: Colubridae) <strong>from</strong> the Western Ghats – India. Taprobanica.<br />

3(2):77–85.


11. De Silva, A., Wijekoon, A.S.B., Jayasena, L., Abeysekera, C.K., Bao,<br />

C.-D., Hutton, R.A. and Warrel, D.A. (1994). Haemostatic dysfunction<br />

and acute renal failure following envenoming by Merrem’s hump-nosed<br />

viper (Hypnale hypnale) in <strong>Sri</strong> <strong>Lanka</strong>: first authenticated case. Transactions<br />

<strong>of</strong> the Royal Society <strong>of</strong> tropical Medicine and Hygiene. 88: 209–12.<br />

12. Premawardena, A..P., Gunathilake, S.B. and de Silva, H.J. (1996).<br />

Haemostatic dysfunction following Hypnale hypnale bites. X1V<br />

International congress for Tropical Medicine and Malaria,Nagasaki, Japan.<br />

17– 22.<br />

13. Sellahewa, K. (1997). Lessons <strong>from</strong> four studies on the management <strong>of</strong><br />

snake bite in <strong>Sri</strong> <strong>Lanka</strong>. Ceylon Medical Journal. 42: 8–15.<br />

14. Kularatne, S.A.M. and Ratnatunge, N. (1999). Severe systemic effects<br />

<strong>of</strong> Merrem’s hump nosed viper bite. The Ceylon Medical Journal. 44(4):<br />

169–170.<br />

15. Ariaratnam C.A., Thuraisingam, V., Kularatne, S.A.M., Sheriff, M.H.R.,<br />

Theakston, R.D.G., de Silva, A. and Warrell, D.A. (2008). Frequent and<br />

potentially fatal envenoming by hump-nosed pit vipers (Hypnale hypnale<br />

and H. nepa) in <strong>Sri</strong> <strong>Lanka</strong>: lack <strong>of</strong> effective antivenom. The Royal Society<br />

<strong>of</strong> Tropical Medicine and Hygiene. 102: 1120–1126.<br />

16. de Silva H.J., Fonseka M.M.D., Gunatilake S.B., Kularatne S.A.M. and<br />

Sellahewa K.H. (2002). Anti-venom for snakebite in <strong>Sri</strong> <strong>Lanka</strong>. The<br />

Ceylon Medical Journal. 47(2): 43–45.<br />

17. Whitehall, J.S., Yarlini., Arunthathy., Varan., Kaanthan., Isaivanan. and<br />

Vanprasath. (2007). Snake bites in north east <strong>Sri</strong> <strong>Lanka</strong>. Rural and<br />

Remote Health. 7: 751 1–6.<br />

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18. de Silva, A. and Ranasinghe, L. (1983). Epidemiology <strong>of</strong> snakebite in <strong>Sri</strong><br />

<strong>Lanka</strong>. Ceylon Medical Journal. 28:144–54.<br />

19. Seneviratne, U. and Dissanayake, S. (2002). Neurological Manifestations<br />

<strong>of</strong> Snake Bite in <strong>Sri</strong> <strong>Lanka</strong>. Journal <strong>of</strong> Postgraduate Medicine. 48: 275–279.<br />

20. Rajapakse, L., (2012) ‘Alarming increase in road accident, deaths,<br />

Island, 9 April (Online). Available at: http://www.island.lk/index.<br />

php?page_cat=article-details&page=article-details&code_title=49371<br />

(Accessed: 6 December 2012).

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