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Contents - LAC Biosafety

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322 Insect pests in plantations: case studies<br />

by erratic lighter defoliations later in the season. Outbreaks are spectacular<br />

events, creating the impression of severe growth loss, although the trees put<br />

forth a new flush of leaves within weeks. Several attempts have been made in the<br />

past to estimate the loss due to defoliation, based on artificial defoliation<br />

experiments, field observations on the frequency of defoliations etc., which put<br />

the loss figures variously at 6.6–65% of the normal increment (Mackenzie, 1921;<br />

Beeson, 1931a, 1941; Champion, 1934). Nair (1986a) made a critical review of<br />

these early attempts and concluded that the available estimates were not<br />

reliable as they rested on untenable assumptions.<br />

In a detailed experimental study, Nair et al. (1996a) estimated the growth in<br />

volume increment in replicated plots in a young teak plantation at Nilambur, in<br />

Kerala, India. Over a five-year period sets of plots were either exposed to natural<br />

insect defoliation or protected from the insect using insecticide. One set of plots<br />

was fully protected by applying insecticide whenever there was threat of damage<br />

by either H. puera or E. machaeralis, while another set of plots was protected only<br />

against H. puera. Defoliation by the two insects was well separated in time which<br />

facilitated such selective protection. A third set of plots with no protection<br />

served as an untreated control. Differences in volume increment were estimated<br />

by the following method. The experiment was started at the time of routine<br />

4th year mechanical thinning, when as per standard silvicultural practice<br />

alternate rows of trees in a plantation are thinned to facilitate growth.<br />

Measurements made on the thinned trees were used to calculate the initial<br />

volume of the standing experimental trees. There were a total of nine plots, with<br />

100 trees per plot, of which half were felled in the 4th year. The girth under bark<br />

of each felled tree was measured at every 50-cm interval, from which the wood<br />

volume of the tree was calculated. Then the mathematical relationship between<br />

the wood volume, on the one hand, and the girth at breast height and total tree<br />

height, on the other, was determined by fitting the most suitable prediction<br />

equation. This equation was used to arrive at the initial volume of the standing<br />

experimental trees from measurements of their girth at breast height and total<br />

tree height. Similar measurements were carried out at the time of the second<br />

mechanical thinning, when 50% of the remaining trees were felled, to obtain<br />

the volume of the trees at the end of the experimental period.<br />

The study showed that during the experimental period the trees protected<br />

against H. puera put forth a mean annual volume increment of 6.7 m 3 /ha<br />

compared with 3.7 m 3 /ha of unprotected trees, a gain of 3 m 3 of wood/ha per<br />

annum. Thus, in young plantations of teak, loss due to defoliation caused by<br />

H. puera was estimated at 44% of the potential volume increment. They also<br />

projected that protected trees would be ready for harvest at the age of 26 years<br />

instead of the usual 60 years, provided other necessary inputs were given.

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