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Contents - LAC Biosafety

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ole in the ecology of tropical forests as pollinators, and parasitoids of injurious<br />

insects. The many species of tropical honeybees alone have provided subsistence<br />

and economic benefits to the tribal, rural and urban societies of the tropics since<br />

ancient times. The role played by parasitoids in the families Ichneumonidae,<br />

Braconidae, Chalcidae, Elasmidae, Eulophidae, Bethylidae, Trichogrammatidae<br />

etc. in keeping the populations of the several tree pests within bounds, by<br />

parasitising their eggs, larvae and pupae, is immeasurable. Although the leafcutting<br />

ants of tropical America are pests in general, large populations of ants<br />

are important predators and scavengers in the tropical forests. In the canopy of<br />

tropical forests in Panama, Erwin (1983b) found that among the 18 orders of<br />

canopy insects present 50.8% of the individuals were hymenopterans, of which<br />

84% were ants. In addition to the leaf-cutting ants, a small number of<br />

hymenopterans such as sawflies, gall wasps and wood wasps are also pests of<br />

trees, although these are more important in temperate than in tropical forests.<br />

In addition, over 100 species of the genus Tetramesa belonging to the<br />

predominantly parasitic family Eurytomidae (subfamily Chalcidoidea) are<br />

phytophagous; T. gigantochloae infests the stem of some bamboos in Malaysia<br />

(Narendran and Kovac, 1995). Among the tropical sawflies (suborder Symphyta),<br />

Shizocera sp. (Argidae) is a defoliator of Manglietia conifera in Vietnam (Tin, 1990)<br />

and several species of Sericoceros (Argidae) feed on the leaves of some trees in<br />

tropical America (Ciesla, 2002).<br />

Order Hemiptera (bugs)<br />

2.2 The diversity of tropical forest insects 43<br />

This order includes bugs that can be distinguished into three main<br />

groups (suborders): Heteroptera or the ‘true bugs’ which includes water skaters,<br />

belostomatids, bed bugs, tingids, lygaeids, pentatomids etc.; Sternorrhyncha<br />

which includes whiteflies, scale insects, aphids and jumping plant lice (psyllids);<br />

and Auchenorrhyncha which includes the leaf hoppers, tree hoppers and<br />

cicadas. In Heteroptera, the forewings are thick and stiff at the basal half and<br />

thin and membraneous at the distal half, and the abdomen has scent or stink<br />

glands. In all bugs, the mouthparts are of a piercing and sucking type. Generally,<br />

the bugs suck the sap of plants, but members of some families such as<br />

Reduviidae and Pentatomidae are predators and suck the fluid of other animals<br />

including insects. The major families of importance to tropical forestry are<br />

Cicadidae, Coccidae, Psyllidae and Tingidae. Cicadas are well-known insects of<br />

the tropical forests and feed on the sap of tender shoots and twigs of trees. They<br />

feed gregariously for long hours and the copious fluid excreta ejected by them<br />

from the tree tops drops on the ground like an incessant spray. The shrill but<br />

loud noise produced by the male cicadas in chorus is characteristic of tropical<br />

forests. The Coccidae include the economically beneficial lac insect on which

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