биота российских вод японского моря - Materials of Alexey Shipunov
биота российских вод японского моря - Materials of Alexey Shipunov биота российских вод японского моря - Materials of Alexey Shipunov
II. Family AMMOTHEIDAE Dohrn, 1881The family combines rather different forms of sea spiders, having compact orelongated trunk, segmented or unsegmented. Chelifore consists of 1, 2, or 3 segments,poorly developed and usually shorter than proboscis, with feeble or reduced fingers,completely absent in some species. Palp often longer than proboscis, consisting of 6–10 (even 20) segments. Both males and females have 10-segmented oviger, with orwithout terminal claw, with compound spines, placed in rows or irregularly, or withcommon flat spines. Four pairs of legs present. Males have genital pores on 3rd and4th legs, females on all legs.KEY TO THE GENERA OF THE FAMILY AMMOTHEIDAE1(4). Trunk compact, disk-shaped, unsegmented or not completely segmented (insome specimens segmentation may be complete); lateral processes placed incontact or radiate out from trunk like rays.2(3). Chelifore 2-segmented; palp 8–9 segmented; coxa 1 of leg bears one tall dorsodistaltubercle ........................................................................ 1. Achelia (p. 105)3(2). Chelifore usually in shape of rudimentary buds; palp consists of 4–7 segments;coxa 1 of leg lacks dorso-distal tubercle ............................*Tanystylum (p. 114)4(1). Trunk elongated, segmented; lateral processes separated by broad intervals.5(6). Chelifore consists of one segment, very small; segment 7 of oviger curved, withtuft of setae on convex side; segment 8 implanted on side of segment 7 ............................................................................................. 2. Lecythorhynchus (p. 115)6(5). Chelifore consists of 2–3 segments; segment 7 of oviger not curved; segment 8implanted on it distally.7(8). Palp consists of 18–20 segments, distal part of it looking like whip; distal ends of1st legs also whip-shaped ............................................... *Nymphonella (p. 116)8(7). Palp consists of 9–10 segments; all legs identical, without whip-shaped ends.9(10). Anterior part of cephalic segment covers implantations of chelifores hood-like;segment 2 of oviger longest; proboscis oval .................... 3. Cilunculus (p. 117)10(9). Anterior part of cephalic segment does not cover implantations of chelifores;segment 2 of oviger much shorter than segment 4; proboscis with one or twoconstrictions, separating its medial oviform part from proximal and distalconical parts .................................................................. *Ascorhynchus (p. 118)1. Genus Achelia Hodge, 1864Type species: Achelia ehinata Hodge, 1864Trunk short, shield-shaped, unsegmented, partially segmented, or, very rarely,completely segmented. Anterior part of cephalic segment has two lateral angles;lateral processes in contact or separated by small intervals, having different width inmales and females. Ocular tubercle situated on anterior part of cephalic segment; eyeswell-developed. Proboscis heavy, often oval, spindle-shaped, sometimes cylindrical.Abdomen fused with distal segment of trunk, usually long. Chelifore short, consists of105
2 segments; adults have vestigial chelae with reduced fingers or without them. Palpconsists of 6–8 segments, usually longer than proboscis. Oviger consists of 10segments, male oviger longer than female one, bearing few compound spines placedirregularly, or bearing simple spines placed similarly. Legs short, covered withtubercles and spines; femur distally bears conical process, larger in males, havingopening of cement gland on top in males; tarsus small, caliciform; propodus more orless curved; in most species propodal sole provided with large spines proximally andsmall spines medially and distally. Main claw usually large, curved; auxiliary clawsusually smaller than main one. Males have genital pores on special process (genitalspur), situated on coxa 2 of legs 3 and 4.KEY TO THE SPECIES OF THE GENUS ACHELIA1(14). Trunk unsegmented, without articulation lines.2(3). Propodal sole lacks isolated large spines; lateral processes in males long, thin,separated by intervals, expanding distally; no compound spines on oviger .................................................................................................... 1. A. borealis (p. 107)3(2). Propodal sole bears large spines.4(5). Dorsal side of trunk has two tubercles, placed one after another, with spines ontops; in adults abdomen base rises over trunk surface, making it saddle-shaped................................................................................. 2. A. bituberculata (p. 108)5(4). Dorsal side of trunk smooth and even, without any tubercles.6(7). Proboscis twice shorter than trunk, almost cylindrical, with rounded tip....................................................................................... *A. brevirostris (p. 108)7(6). Proboscis robust, almost equal to trunk in length, oval, spindle-shaped, orpyriform.8(9). Ocular tubercle cylindrical, tall, with conical top bearing small eyes; chelifores,palps and legs bear long spines, covered with short thin setae .......................................................................................................................... *A. latifrons (p. 109)9(8). Ocular tubercle low, conical; eyes situated in its middle part; all appendages bearlong spines without setae.10(11). Two small tubercles with apical spines on frontal margin of trunk and at distalangles of lateral processes; proboscis oval; chelifore longer than half ofproboscis; scape bears medial tubercle with spine on top ...................................................................................................................... 3. A. kurilensis (p. 109)11(10). Frontal margin of trunk smooth; each lateral process in adults bears one smalltubercle without spines (juveniles may lack such tubercles, or may have 2tubercles with apical spines on each lateral process); proboscis has anothershape; chelifore much shorter than half of proboscis.12(13). Proboscis spindle-shaped; propodus strong, noticeably curved; main clawheavy, reaching past middle of propodus; auxiliary claws slightly longer thanhalf of main claw ....................................................... 4. A. alaskensis (p. 110)13(12). Proboscis pyriform, consisting of conical basal part and large oviform part;propodus relatively thin, curved; main claw curved, reaching middle ofpropodus; auxiliary claws hardly reach middle of main claw ................................................................................................................ 5. A. gracilipes (p. 111)106
- Page 57 and 58: 2. Neomysis awatschensis (Brandt, 1
- Page 59 and 60: 6. Genus Boreoacanthomysis Fukuoka
- Page 61 and 62: 8. Genus Exacanthomysis Holmquist,
- Page 63 and 64: exopod with one simple lateral seta
- Page 65 and 66: Mauchline, J. 1980. The biology of
- Page 67 and 68: Plate II. Archaeomysis grebnitzkii
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- Page 71 and 72: Plate VI. Xenacanthomysis pseudomac
- Page 73 and 74: Plate VIII. Disacanthomysis dybowsk
- Page 75 and 76: Plate X. Neomysis awatschensis: 1 -
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- Page 79 and 80: Plate XIV. Exacanthomysis stelleri:
- Page 81 and 82: Plate XVI. Paracanthomysis shikhota
- Page 83 and 84: (pl. III, figs. 2, 7-9), do not fee
- Page 85 and 86: 1. Thysanoessa inermis (Kroyer, 184
- Page 87 and 88: 35°45′N) of Honshu Island. In th
- Page 89 and 90: Hansen, H.J. 1911. The genera and s
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- Page 93 and 94: SUBPHYLUM CHELICERATA HEYMONS, 1901
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- Page 97 and 98: 14(15). Compound spines with 1 pair
- Page 99 and 100: Geographical distribution. N. brevi
- Page 101 and 102: *Nymphon longitarse brevicollis Los
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- Page 105: 8. Nymphon uniunguiculatum Losina-L
- Page 109 and 110: 2. Achelia bituberculata Hedgpeth,
- Page 111 and 112: setae. Oviger relatively short; ovi
- Page 113 and 114: Description. Utinomi’s collection
- Page 115 and 116: Geographical distribution. A. echin
- Page 117 and 118: 1. Lecythorhynchus marginatus Cole,
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- Page 125 and 126: *Genus Phoxichilidium Milne-Edwards
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- Page 133 and 134: Kim, I.H. & J.S. Hong. 1986. Korean
- Page 135 and 136: Turpaeva, E.P. 2004b. Shallow-water
- Page 137 and 138: Plate II. Nymphon grossipes (from S
- Page 139 and 140: Plate IV. Nymphon longitarse longit
- Page 141 and 142: Plate VI. Nymphon japonicum (from N
- Page 143 and 144: Plate VIII. Nymphon stocki (from Ut
- Page 145 and 146: Plate X. Achelia kurilensis (from L
- Page 147 and 148: Plate XII. Achelia gracilipes (from
- Page 149 and 150: Plate XIV. Tanystylum scrutator (fr
- Page 151 and 152: Plate XVI. Nymphonella tapetis (fro
- Page 153 and 154: Plate XVIII. Decachela discata (fro
- Page 155 and 156: Plate XX. Phoxichilidium ungellatum
2 segments; adults have vestigial chelae with reduced fingers or without them. Palpconsists <strong>of</strong> 6–8 segments, usually longer than proboscis. Oviger consists <strong>of</strong> 10segments, male oviger longer than female one, bearing few compound spines placedirregularly, or bearing simple spines placed similarly. Legs short, covered withtubercles and spines; femur distally bears conical process, larger in males, havingopening <strong>of</strong> cement gland on top in males; tarsus small, caliciform; propodus more orless curved; in most species propodal sole provided with large spines proximally andsmall spines medially and distally. Main claw usually large, curved; auxiliary clawsusually smaller than main one. Males have genital pores on special process (genitalspur), situated on coxa 2 <strong>of</strong> legs 3 and 4.KEY TO THE SPECIES OF THE GENUS ACHELIA1(14). Trunk unsegmented, without articulation lines.2(3). Propodal sole lacks isolated large spines; lateral processes in males long, thin,separated by intervals, expanding distally; no compound spines on oviger .................................................................................................... 1. A. borealis (p. 107)3(2). Propodal sole bears large spines.4(5). Dorsal side <strong>of</strong> trunk has two tubercles, placed one after another, with spines ontops; in adults abdomen base rises over trunk surface, making it saddle-shaped................................................................................. 2. A. bituberculata (p. 108)5(4). Dorsal side <strong>of</strong> trunk smooth and even, without any tubercles.6(7). Proboscis twice shorter than trunk, almost cylindrical, with rounded tip....................................................................................... *A. brevirostris (p. 108)7(6). Proboscis robust, almost equal to trunk in length, oval, spindle-shaped, orpyriform.8(9). Ocular tubercle cylindrical, tall, with conical top bearing small eyes; chelifores,palps and legs bear long spines, covered with short thin setae .......................................................................................................................... *A. latifrons (p. 109)9(8). Ocular tubercle low, conical; eyes situated in its middle part; all appendages bearlong spines without setae.10(11). Two small tubercles with apical spines on frontal margin <strong>of</strong> trunk and at distalangles <strong>of</strong> lateral processes; proboscis oval; chelifore longer than half <strong>of</strong>proboscis; scape bears medial tubercle with spine on top ...................................................................................................................... 3. A. kurilensis (p. 109)11(10). Frontal margin <strong>of</strong> trunk smooth; each lateral process in adults bears one smalltubercle without spines (juveniles may lack such tubercles, or may have 2tubercles with apical spines on each lateral process); proboscis has anothershape; chelifore much shorter than half <strong>of</strong> proboscis.12(13). Proboscis spindle-shaped; propodus strong, noticeably curved; main clawheavy, reaching past middle <strong>of</strong> propodus; auxiliary claws slightly longer thanhalf <strong>of</strong> main claw ....................................................... 4. A. alaskensis (p. 110)13(12). Proboscis pyriform, consisting <strong>of</strong> conical basal part and large oviform part;propodus relatively thin, curved; main claw curved, reaching middle <strong>of</strong>propodus; auxiliary claws hardly reach middle <strong>of</strong> main claw ................................................................................................................ 5. A. gracilipes (p. 111)106