Colorado Statewide Forest Resource Assessment

Colorado Statewide Forest Resource Assessment Colorado Statewide Forest Resource Assessment

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Mixed Conifer 14The mixed conifer forest type occurs at approximately 6,900 to 10,500 feetin elevation, nestled between lower-elevation forests such as ponderosapine and higher-elevation subalpine forests such as spruce-fi r. Colorado has1.8 million acres of mixed-conifer forest, which is primarily managed by theUSFS (873,280 acres) and private landowners (602,356 acres). As its namesuggests, the mixed-conifer type includes a diverse range of species. Thedistribution and structure of mixed-conifer forests are strongly infl uenced bytemperature and moisture gradients, in addition to soil types and fi re. White fi roften dominates as the climax species on moist sites and in the southern partof the state, while ponderosa pine, Douglas-fi r or Rocky Mountain juniper tendto be the climax species on warmer and drier sites. Engelmann spruce, bluespruce, subalpine fi r, bristlecone pine and limber pine also may be present inthe mix.Fire regimes in mixed-conifer forests vary by site, but generally are of lowfrequency and mixed severity, ranging from 20-year cycles to 150 years ormore depending on moisture and elevation. 15 As a result of fi re suppression,many mixed-conifer forests currently are denser and contain more deadfuel than they did historically. These heavy accumulations greatly increasethe chances for high-intensity, stand-replacing crown fi res. Prior to humansettlement, these stand-replacing fi res would only occur every 200 to 500years. Mixed-conifer forests are important for watershed protection andrecreational and scenic opportunities. Mixed-conifer forests provide manywood products manufactured by local Colorado businesses. Compared toother forest types, the value of mixed-conifer forests for wildlife is not wellunderstood. However, many neotropical migratory songbirds breed in mixedconiferforests and elk use them extensively during calving season and insummer. Flammulated owls and northern goshawks are among the sensitivespecies that use mixed-conifer stands, especially those that include bluespruce. Twenty Species of Greatest Conservation Need use mixed-coniferforests as a primary habitat (CDOW 2006).Ponderosa PinePonderosa pine forests, woodlands and savannas occupy approximately 2million acres in Colorado or 8 percent of the state’s forested land. Althoughponderosa pine is most common between 6,000 and 9,000 feet, it begins toappear on the landscape around 5,000 feet where prairies and shrublandstransition into open ponderosa pine forests. 16 Private landowners control 49percent of this forest type, while the USFS manages 34 percent. Becauseit is more accessible than other species, and largely in private ownership,ponderosa pine has been an important source of lumber and other woodproducts throughout Colorado’s history. These forests currently are asignifi cant resource for recreation and water supply. They also are thebackdrop for the majority of the state’s wildland-urban interface communities.14This section is adapted from information available from LANDFIRE (www.landfire.gov) and NatureServe (www.natureserve.org/explorer).15For more specific information about this variation in fire regimes, please see the LANDFIRE biophysical settingdescriptions for mixed-conifer at www.landfire.gov or the USFS fire effects information system at www.fs.fed.us/database/feis/.16For more information on Colorado’s ponderosa pine forests, see the CSFS 2004 Report on the Health of Colorado’sForests at http://csfs.colostate.edu/pages/pub-csfs2.html.10

Ponderosa pine forests are the only forest type designated as a high-priorityhabitat in Colorado’s State Wildlife Action Plan. Twenty-eight of Colorado’sSpecies of Greatest Conservation Need rely on ponderosa pine forests astheir primary habitat (CDOW 2006). One of the most signifi cant of thesespecies is the Pawnee montane skipper. This butterfl y is federally listed asThreatened and is restricted to the Upper South Platte watershed in Coloradowhere it inhabits ponderosa pine forests in which blue grama grass andprairie gayfeather are found in the understory. Other species include theMexican spotted owl, northern goshawk, Townsend’s big-eared bat and theAmerican three-toed woodpecker. Primary threats to ponderosa pine forestsinclude urban development, noxious weeds and the long-term impactsof fi re exclusion. Ponderosa pine forests evolved with natural cycles offrequent, low-intensity fi re that allowed mature pine trees to survive, but killedcompeting conifer seedlings. Several decades of near total fi re exclusion haveresulted in signifi cantly increased tree density in many of these forests, aswell as encroachment from Douglas-fi r. Such conditions increase the risk oflarge, high-intensity crown fi res in ponderosa pine forests and have increasedtheir susceptibility to native insects and diseases. The Colorado BiodiversityScorecard ranks ponderosa pine forests as one of the state’s weaklyconserved forest types (CNHP and TNC 2008).Piñon-JuniperPiñon-juniper woodlands are widespread in the lower elevations ranging from4,900 to 8,000 feet on Colorado’s West Slope and exist in limited distributionin south-central Colorado and on the Eastern Plains. The most commonspecies within this type are Colorado piñon pine and Utah and one-seedjuniper, although Rocky Mountain juniper may codominate or replaceone-seed at higher elevations. Piñon-juniper woodlands account for just over5 million acres or approximately 21 percent of the state’s forested lands. Fullyhalf of Colorado’s piñon-juniper landscapes are managed by the Bureau ofLand Management (BLM) with another 32 percent in private ownership. Piñonjuniperstands vary considerably in appearance and composition, dependingon soil depth, elevation and geographic locations. While piñon and junipermay co-occur in some landscapes, either species may be dominant on itsown in others. Some believe that the ecological dynamics of these woodlandshave changed since European settlement, resulting in increased density andencroachment on grasslands, but limited research is available to verify theseconcerns (Romme and others 2008). The research that does exist suggeststhat piñon-juniper forests likely were diverse historically, as they are today.Although not a traditional timber species, piñon-juniper forests have longbeen important to local communities for such products as fuelwood, fenceposts, pine nuts, forage for livestock and watershed protection. More recently,communities and businesses have begun to turn to piñon-juniper forests asa source of biomass for fuel and energy. Piñon-juniper forests also providehabitat for many of Colorado’s rarest plants along with the gray vireo, one ofthe state’s rare birds. Piñon-juniper forests are signifi cant habitat for reptiles,in general, and for lizards in particular. They are extensively used as winteringgrounds by elk on the Western Slope and as year-round cover by elk on theEastern Plains (e.g., Purgatoire Canyon). A notable 32 of the state’s Speciesof Greatest Conservation Need rely on piñon-juniper forests as a primaryhabitat (CDOW 2006).11

Ponderosa pine forests are the only forest type designated as a high-priorityhabitat in <strong>Colorado</strong>’s State Wildlife Action Plan. Twenty-eight of <strong>Colorado</strong>’sSpecies of Greatest Conservation Need rely on ponderosa pine forests astheir primary habitat (CDOW 2006). One of the most signifi cant of thesespecies is the Pawnee montane skipper. This butterfl y is federally listed asThreatened and is restricted to the Upper South Platte watershed in <strong>Colorado</strong>where it inhabits ponderosa pine forests in which blue grama grass andprairie gayfeather are found in the understory. Other species include theMexican spotted owl, northern goshawk, Townsend’s big-eared bat and theAmerican three-toed woodpecker. Primary threats to ponderosa pine forestsinclude urban development, noxious weeds and the long-term impactsof fi re exclusion. Ponderosa pine forests evolved with natural cycles offrequent, low-intensity fi re that allowed mature pine trees to survive, but killedcompeting conifer seedlings. Several decades of near total fi re exclusion haveresulted in signifi cantly increased tree density in many of these forests, aswell as encroachment from Douglas-fi r. Such conditions increase the risk oflarge, high-intensity crown fi res in ponderosa pine forests and have increasedtheir susceptibility to native insects and diseases. The <strong>Colorado</strong> BiodiversityScorecard ranks ponderosa pine forests as one of the state’s weaklyconserved forest types (CNHP and TNC 2008).Piñon-JuniperPiñon-juniper woodlands are widespread in the lower elevations ranging from4,900 to 8,000 feet on <strong>Colorado</strong>’s West Slope and exist in limited distributionin south-central <strong>Colorado</strong> and on the Eastern Plains. The most commonspecies within this type are <strong>Colorado</strong> piñon pine and Utah and one-seedjuniper, although Rocky Mountain juniper may codominate or replaceone-seed at higher elevations. Piñon-juniper woodlands account for just over5 million acres or approximately 21 percent of the state’s forested lands. Fullyhalf of <strong>Colorado</strong>’s piñon-juniper landscapes are managed by the Bureau ofLand Management (BLM) with another 32 percent in private ownership. Piñonjuniperstands vary considerably in appearance and composition, dependingon soil depth, elevation and geographic locations. While piñon and junipermay co-occur in some landscapes, either species may be dominant on itsown in others. Some believe that the ecological dynamics of these woodlandshave changed since European settlement, resulting in increased density andencroachment on grasslands, but limited research is available to verify theseconcerns (Romme and others 2008). The research that does exist suggeststhat piñon-juniper forests likely were diverse historically, as they are today.Although not a traditional timber species, piñon-juniper forests have longbeen important to local communities for such products as fuelwood, fenceposts, pine nuts, forage for livestock and watershed protection. More recently,communities and businesses have begun to turn to piñon-juniper forests asa source of biomass for fuel and energy. Piñon-juniper forests also providehabitat for many of <strong>Colorado</strong>’s rarest plants along with the gray vireo, one ofthe state’s rare birds. Piñon-juniper forests are signifi cant habitat for reptiles,in general, and for lizards in particular. They are extensively used as winteringgrounds by elk on the Western Slope and as year-round cover by elk on theEastern Plains (e.g., Purgatoire Canyon). A notable 32 of the state’s Speciesof Greatest Conservation Need rely on piñon-juniper forests as a primaryhabitat (CDOW 2006).11

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