Ecology and Development Series No. 10, 2003 - ZEF
Ecology and Development Series No. 10, 2003 - ZEF Ecology and Development Series No. 10, 2003 - ZEF
Impacts of human use on the forest vegetation5.4.2 Population structure of treesSome tree species (Type I population structure group) are less affected by disturbance dueto management of the forests for coffee production. Species with such population structureare considered as being in good regeneration condition (Demel 1997). It appears that thechange in vegetation structure and the light conditions under the managed forests do notaffect the ability of their seeds to germinate. These are large canopy trees, which are alsoimportant as shade trees in the forests managed for coffee production. Hence, they are lesstargeted during the thinning of trees and clearing of understorey vegetation, unless there isa higher density of their stems than required for shade. The tree species in Type II appear tobe pioneer species, whose seeds require disturbance for germination. The number of maturestems of Cordia africana, which belongs to this group, decreased in the managed forestcategories compared to the undisturbed forests, and finally disappeared from the oldestmanaged forest type (SEMIFOR-PLAN) (Table 5.7). This can be due to two possiblereasons: (1) broad-leaved deciduous tree species like Cordia are considered as unsuitablefor shade and are gradually replaced by legumes or other broad-leaved evergreen treespecies, and (2) Cordia is highly hunted for its quality timber.The tree species in the Type III appear to be shade-tolerant, and cannot regeneratein gaps created under disturbed forest conditions. The trees in this group are also notrepresented even in the higher height classes of the vegetation of the managed forestcategories, probably because they are not useful trees for shade in coffee production.5.4.3 The forest structureThere is a clear difference in the structure of the forest vegetation between the undisturbedforests and the forests managed for coffee production. It appears that forests are convertedfrom complex and continuous-structured vegetation in undisturbed forests to two-layeredvegetation in the managed forests. The two distinguishable layers in managed forests are:the canopy layer of shade trees and the coffee stand layer. The high degree of decline in theabundances of plants in the lower height classes in the managed forest (Table 5.10), therelatively lower decline in abundances of trees in the higher height class and canopy cover(Table 5.7 and 5.10), and the very high increase in the proportion of matured coffee trees93
Impacts of human use on the forest vegetation(Table 5.8) in the managed forests demonstrate this fact. Beside the high degree of changein abundances of lower height classes of the forest trees in the managed forests, manyspecies of climbers that make the structure more continuous and complex have eithertotally disappeared or are represented only by seedlings (Appendix 7). On the contrary,Parthasarathy (1999) observed higher liana diversity in a frequently disturbed forest than inan undisturbed natural forest. The situation observed in Yayu forest is due to the highintensity of management mainly targeting at understorey vegetation and climbers to avoidcompetition with coffee and for ease of access during harvesting.The lowest average canopy cover in the managed forest is 51% (SEMIFOR-PLAN) compared to the highest of 86% in undisturbed forest (NATUREFOR) (Table 5.3).The change in canopy cover is small, even though the decline in the abundance of stems indifferent height classes is as high as four- to ten-fold in the managed forests as compared tothat of undisturbed forests. In spite of enormous changes in vegetation structure, highcanopy cover is maintained in the managed forest by keeping those trees with large andspreading canopies. Mature coffee plants fully occupy the lower stratum, since clearing thecompeting vegetation increases secondary branch development in the coffee trees (Tewolde1978).The decline in the basal areas of the two older managed forests (SEMIFOR-OLDand SEMIFOR-PLAN) is very high (Table 5.11). However, the basal area of theSEMIFOR-NEW remained quite similar to the undisturbed forests, even though there is asignificant decline in the number of trees in higher height classes (Table 5.10; Table 5.11).This is mainly because the shade trees retained in the forest are big canopy trees with largediameters in the recently opened-up forest.Even though the abundance of coffee trees in all forests is relatively similar (Table5.8), the basal area increases greatly in the managed forests (from 1.4 m 2 ha -1 in NATFORto 7.8 m 2 ha -1 in SEMIFOR-PLAN), indicating that the population has converted frombeing dominated by seedlings in the undisturbed forests to being dominated by mature treesin the managed forest. Abayneh (1998) reported a higher number of coffee seedlings inforest gaps than in closed canopy forest. The lower proportion of seedlings in the managedforests at Yayu can be either for biological or human-induced reasons, or both, i.e., due to:94
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- Page 136 and 137: ReferencesBatisse M. 1986 Developin
- Page 138 and 139: ReferencesDavis A.P. and Rokotonaso
- Page 140 and 141: ReferencesESRI. 1996. ArcView GIS:
- Page 142 and 143: ReferencesIUCN 1992. Protected Area
- Page 144 and 145: ReferencesMesfin Ameha and Bayetta
- Page 146 and 147: ReferencesSmith R.F. 1985. A histor
- Page 148 and 149: ReferencesVan Jaarsveld A.S., Freit
Impacts of human use on the forest vegetation5.4.2 Population structure of treesSome tree species (Type I population structure group) are less affected by disturbance dueto management of the forests for coffee production. Species with such population structureare considered as being in good regeneration condition (Demel 1997). It appears that thechange in vegetation structure <strong>and</strong> the light conditions under the managed forests do notaffect the ability of their seeds to germinate. These are large canopy trees, which are alsoimportant as shade trees in the forests managed for coffee production. Hence, they are lesstargeted during the thinning of trees <strong>and</strong> clearing of understorey vegetation, unless there isa higher density of their stems than required for shade. The tree species in Type II appear tobe pioneer species, whose seeds require disturbance for germination. The number of maturestems of Cordia africana, which belongs to this group, decreased in the managed forestcategories compared to the undisturbed forests, <strong>and</strong> finally disappeared from the oldestmanaged forest type (SEMIFOR-PLAN) (Table 5.7). This can be due to two possiblereasons: (1) broad-leaved deciduous tree species like Cordia are considered as unsuitablefor shade <strong>and</strong> are gradually replaced by legumes or other broad-leaved evergreen treespecies, <strong>and</strong> (2) Cordia is highly hunted for its quality timber.The tree species in the Type III appear to be shade-tolerant, <strong>and</strong> cannot regeneratein gaps created under disturbed forest conditions. The trees in this group are also notrepresented even in the higher height classes of the vegetation of the managed forestcategories, probably because they are not useful trees for shade in coffee production.5.4.3 The forest structureThere is a clear difference in the structure of the forest vegetation between the undisturbedforests <strong>and</strong> the forests managed for coffee production. It appears that forests are convertedfrom complex <strong>and</strong> continuous-structured vegetation in undisturbed forests to two-layeredvegetation in the managed forests. The two distinguishable layers in managed forests are:the canopy layer of shade trees <strong>and</strong> the coffee st<strong>and</strong> layer. The high degree of decline in theabundances of plants in the lower height classes in the managed forest (Table 5.<strong>10</strong>), therelatively lower decline in abundances of trees in the higher height class <strong>and</strong> canopy cover(Table 5.7 <strong>and</strong> 5.<strong>10</strong>), <strong>and</strong> the very high increase in the proportion of matured coffee trees93