PDF - Wallace Online
PDF - Wallace Online PDF - Wallace Online
76 NATURAL SELECTION inbrown, so that when on the wing and seen from below theyare undistinguishable. The curious point, however, is thatthe Accipiter has a much wider range than the Harpagus,and in the regions where the insect-eating species is not foundit no longer resembles it, the under wing-coverts varying towhite ;thus indicating that the red-brown colour is kept trueby its being useful to the Accipiter to be mistaken for theinsect-eating species, which birds have learnt not to be afraid of.Mimicry among MammalsAmong the Mammalia the only case which may be truemimicry is that of the insectivorous genus Cladobates, foundin the Malay countries, several species of which very closelyresemble squirrels. The size is about the same, the longbushy tail is carried in the same way, and the colours arevery similar. In this case the use of the resemblance mustbe to enable the Cladobates to approach the insects or smallbirds on which it feeds under the disguise of the harmlessfruit-eating squirrelObjections to Mr. Bates' Theory of MimicryHaving now completed our survey of the most prominentand remarkable cases of mimicry thathave yet been noticed,we must say something of the objections that have been madeto the theory of then- production given by Mr. Bates, andwhich we have endeavoured to illustrate and enforce in thepreceding pages. Three counter explanations have been proposed.Professor Westwood admits the fact of the mimicryand its probable use to the insect, but maintains that eachspecies was created a mimic for the purpose of the protectionthus afforded it. Mr. Andrew Murray, in his paper on the"Disguises of Nature," inclines to the opinion that similarconditions of food and of surrounding circumstances haveacted in some unknown way to produce the resemblances ;and when the subject was discussed before the EntomologicalSociety of London, a third objection was added that heredityor the reversion to ancestral types of form and colorationmight have produced many of the cases of mimicry.Against the special creation of mimicking species there areall the objections and difficulties in the way of special creation
in PROTECTIVE RESEMBLANCES AMONG ANIMALS 77in other cases, with the addition of a few that are peculiar toit. The most obvious is, that we have gradations of mimicryand of protective resemblance a fact which is stronglysuggestive of a natural process having been at work. Anothervery serious objection is, that as mimicry has been shown tobe useful only to those species and groups which are rare andprobably dying out, and. would cease to have any effect shouldthe proportionate abundance of the two species be reversed,it follows that on the special-creation theory the one speciesmust have been created plentiful, the other rare ; and, notwithstandingthe many causes that continually tend to alterthe proportionsof species, these two species must have alwaysbeen specially maintained at their respective proportions, orthe very purpose for which they each received their peculiarcharacteristics would have completely failed. A third difficultyis, that although it is very easy to understand howvariation and the survivalmimicry may be brought about byof the fittest, it seems a very strange thing for a Creator toprotect an animal by making it imitate another, when thevery assumption of a Creator implies his power to create itso as to require no such circuitous protection. These appearto be fatal objections to the application of the special-creationtheory to this particular case.The other two supposed explanations, which may beshortly expressed as the theories of " similar conditions " andof " heredity," agree in making mimicry, where it exists, anadventitious circumstance not necessarily connected with thewell-being of the mimicking species. But several of the moststriking and most constant facts which have been adduceddirectly contradict both these hypotheses. The law thatmimicry is confined to a few groups only is one of these, for" similar conditions " must act more or less on all groups in alimited region, and "heredity" must influence all groupsrelated to each other in an equal degree. Again, the generalfact that those species which mimic others are rare, whilethose which are imitated are abundant, is in no way explainedby either of these theories, any more than is the frequentoccurrence of some papable mode of protection in the imitated"species. Reversion to an ancestral type " no way explainswhy the imitator and the imitated always inhabit the very
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in PROTECTIVE RESEMBLANCES AMONG ANIMALS 77in other cases, with the addition of a few that are peculiar toit. The most obvious is, that we have gradations of mimicryand of protective resemblance a fact which is stronglysuggestive of a natural process having been at work. Anothervery serious objection is, that as mimicry has been shown tobe useful only to those species and groups which are rare andprobably dying out, and. would cease to have any effect shouldthe proportionate abundance of the two species be reversed,it follows that on the special-creation theory the one speciesmust have been created plentiful, the other rare ; and, notwithstandingthe many causes that continually tend to alterthe proportionsof species, these two species must have alwaysbeen specially maintained at their respective proportions, orthe very purpose for which they each received their peculiarcharacteristics would have completely failed. A third difficultyis, that although it is very easy to understand howvariation and the survivalmimicry may be brought about byof the fittest, it seems a very strange thing for a Creator toprotect an animal by making it imitate another, when thevery assumption of a Creator implies his power to create itso as to require no such circuitous protection. These appearto be fatal objections to the application of the special-creationtheory to this particular case.The other two supposed explanations, which may beshortly expressed as the theories of " similar conditions " andof " heredity," agree in making mimicry, where it exists, anadventitious circumstance not necessarily connected with thewell-being of the mimicking species. But several of the moststriking and most constant facts which have been adduceddirectly contradict both these hypotheses. The law thatmimicry is confined to a few groups only is one of these, for" similar conditions " must act more or less on all groups in alimited region, and "heredity" must influence all groupsrelated to each other in an equal degree. Again, the generalfact that those species which mimic others are rare, whilethose which are imitated are abundant, is in no way explainedby either of these theories, any more than is the frequentoccurrence of some papable mode of protection in the imitated"species. Reversion to an ancestral type " no way explainswhy the imitator and the imitated always inhabit the very