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PDF - Wallace Online

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460 TROPICAL NATUREbotanists, farmers, gardeners, sporting -men, pigeon -fanciers,travellers, and any one who could possiblyafford direct personalinformation on any of the matters he was investigating.Then came his own observation and experiment, to fillup gaps,to settle doubtful points, or to determine questions the importanceof inquiring into which no one had ever suspected ;andlastly, there was the power of arrangement and comparison,the originality and depth of thought, which drew out from thisvast mass of heterogeneous materials conclusions of the highestvalue as bearing on the question of the possible change ofspecies, and the means by which it had been brought about.In order to determine the nature and amount of the variabilityof domestic productions, he prepared skeletons of allthe more important breeds of rabbits, pigeons, fowls, andducks, as well as of the wild races from which they areknown to have been produced, and showed, both by measurementsand by accurate drawings, that not only superficialcharacters, but almost every part of the bony structurevaried to such an amount as usually characterises very distinctspecies or even distinct genera of wild animals. Anotherset of experiments was made by crossing the different breedsof pigeons and fowls which were most completely unlike thewild race, with the result that in many cases the offspringwere more like the wild ancestor than either of the parents.These experiments, supported by a mass of facts observed byother persons, served to establish the principle of the tendencyof crosses to revert to the ancestral form and this;principle enabled him to explain the interesting fact of thefrequent appearance of stripes on mules, and occasionally ondun-coloured horses, on the hypothesis, supported by a massof collateral evidence, that the common ancestor of the horse,ass, and zebra tribe was a partially striped and dun-colouredanimal.A number of very important conclusions were deducedfrom the facts presented by domesticated animals and plants,a few of which may be here referred to. For example, itwas proved that the parts most selected or which had alreadymost varied as the tail in fan-tailed pigeons, which has moretail-feathers than any one of the 8000 different kinds ofliving birds were most subject to further variation; and

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