PDF - Wallace Online
PDF - Wallace Online PDF - Wallace Online
30 NATURAL SELECTIONThe Partial Reversion of Domesticated Varieties explainedLet us now turn to domesticated animals, and inquire howvarieties produced among them are affected by the principleshere enunciated. The essential difference in the condition ofwild and domestic animals is this, that among the former,their well - being and very existence depend upon the fullexercise and healthy condition of all their senses and physicalpowers, whereas, among the latter, these are only partiallyexercised, and in some cases are absolutely unused. A wildanimal has to search, and often to labour, for every mouthfulof food to exercise sight, hearing, and smell in seeking it,and in avoiding dangers, in procuring shelter from the inclemencyof the seasons, and in providing for the subsistenceand safety of its offspring. There is no muscle of itsbodythat is not called into daily and hourly activity ;there is nosense or faculty that is not strengthened by continual exercise.The domestic animal, on the other hand, has food providedfor is it, sheltered, and often confined, to guard it againstthe vicissitudes of the seasons, is carefully secured from theattacks of its natural enemies, and seldom even rears itsyoungwithout human assistance. Half of its senses and facultiesbecome quite useless, and the other half are but occasionallycalled into feeble exercise, while even its muscular system isonly irregularly brought into action.Now when a variety of such an animal occurs havingincreased power or capacity in any organ or sense, suchincrease is totally useless, is never called into action, and mayeven exist without the animal ever becoming aware of it. Inthe wild animal, on the contrary, all its faculties and powersbeing brought into full action for the necessities of existence,any increase becomes immediately available, is strengthenedby exercise, and must even slightly modify the food, the habits,and the whole economy of the race. It creates as it were anew animal, one of superior powers, and which will necessarilyincrease in numbers and outlive those which are inferior to it.Again, in the domesticated animal all variations have anequal chance of continuance and those which would decidedlyrender a wild animal unable to compete with its;fellows and continue its existence are no disadvantage what-
ii ON THE TENDENCY OF VARIETIES, ETC. 31ever in a state of domesticity. Our quickly fattening pigs,short -legged sheep, pouter pigeons, and poodle dogs couldnever have come into existence in a state of nature, becausethe very first steps towards such inferior forms would haveled to the rapid extinction of the race ; still less could theynow exist in competition with their wild allies. The greatspeed but slight endurance of the racehorse, the unwieldlystrength of the ploughman's team, would both be useless ina state of nature. If turned wild on the pampas, suchanimals would probably soon become extinct, or underfavourable circumstances might each gradually lose thoseextreme qualities which would never be called into action,and in a few generations revert to a common type, whichmust be that in which the various powers and faculties are soproportioned to each other as to be best adapted to procurefood and secure safety, that in which, by the full exercise ofevery part of its organisation, the animal can alone continueto live. Domestic varieties, when turned wild, must returnto something near the type of the original wild stock, orbecome altogether extinct}-We see, then, that no inferences as to the permanence ofvarieties in a state of nature can be deduced from the observationsof those occurring among domestic animals. Thetwo are so much opposed to each other in every circumstanceof their existence, that what applies to the one is almost surenot to apply to the other. Domestic animals are abnormal,irregular,artificial ; they are subject to variations whichnever occur, and never can occur, in a state of nature : theirvery existence depends altogether on human care so far aremany of them removed from that just proportion of faculties,that true balance of organisation, by means of which alone ananimal left to its own resources can preserve its existence andcontinue its race.Lamarck's Hypothesis very different from that now advancedThe hypothesis of Lamarck that progressive changes inspecies have been produced by the attempts of animals to1That is, they will vary, and the variations which tend to adapt them tothe wild state, and therefore approximate them to wild animals, will be preserved.Those individuals which do not vary sufficiently will perish.
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30 NATURAL SELECTIONThe Partial Reversion of Domesticated Varieties explainedLet us now turn to domesticated animals, and inquire howvarieties produced among them are affected by the principleshere enunciated. The essential difference in the condition ofwild and domestic animals is this, that among the former,their well - being and very existence depend upon the fullexercise and healthy condition of all their senses and physicalpowers, whereas, among the latter, these are only partiallyexercised, and in some cases are absolutely unused. A wildanimal has to search, and often to labour, for every mouthfulof food to exercise sight, hearing, and smell in seeking it,and in avoiding dangers, in procuring shelter from the inclemencyof the seasons, and in providing for the subsistenceand safety of its offspring. There is no muscle of itsbodythat is not called into daily and hourly activity ;there is nosense or faculty that is not strengthened by continual exercise.The domestic animal, on the other hand, has food providedfor is it, sheltered, and often confined, to guard it againstthe vicissitudes of the seasons, is carefully secured from theattacks of its natural enemies, and seldom even rears itsyoungwithout human assistance. Half of its senses and facultiesbecome quite useless, and the other half are but occasionallycalled into feeble exercise, while even its muscular system isonly irregularly brought into action.Now when a variety of such an animal occurs havingincreased power or capacity in any organ or sense, suchincrease is totally useless, is never called into action, and mayeven exist without the animal ever becoming aware of it. Inthe wild animal, on the contrary, all its faculties and powersbeing brought into full action for the necessities of existence,any increase becomes immediately available, is strengthenedby exercise, and must even slightly modify the food, the habits,and the whole economy of the race. It creates as it were anew animal, one of superior powers, and which will necessarilyincrease in numbers and outlive those which are inferior to it.Again, in the domesticated animal all variations have anequal chance of continuance and those which would decidedlyrender a wild animal unable to compete with its;fellows and continue its existence are no disadvantage what-