PDF - Wallace Online
PDF - Wallace Online PDF - Wallace Online
404 TROPICAL NATUREby butterflies. 1 But bees are always (in the temperate zone)far more abundant than butterflies, and this will be anotherreason why flowers specially adapted to be fertilised by thelatter should be rendered unusually conspicuous. We find,accordingly, the yellow primroses and cowslips of the plainsreplaced by pink and magenta -coloured Alpine species; thestraggling wild pinks of the lowlands by the masses of largeflowers in such mountain species as Dianthus alpinus and D.glacialis; the saxifrages of the high Alps with bunches offlowers a foot long as in Saxifraga longif olia and S. cotyledon,or forming spreading masses of flowers as in S. oppositifolia ;while the soapworts, silenes, and louseworts are equally superiorto the allied speciesof the plains.Why Allied Species of Flowers differ in Size and BeautyAgain, Dr. Miiller has discovered that when there areshowy and inconspicuous species in the same genus of plants,there is often a corresponding difference of structure, thosewith large and showy flowers being quite incapable of selffertilisation,and thus depending for their very existence onthe visits of insects, while the others are able to fertilisethemselves should insects fail to visit them. We haveexamples of this difference in Malva sylvestris, Epilobiumangustifolium, Polygonum bistorta, and Geranium pratensewhich have all large or showy flowers, and must be fertilisedby insects as compared with Malva rotundifolia, Epilobiumparviflorum, Polygonum aviculare, and Geranium pusillum,which have small or inconspicuous flowers, and are so constructedthat if insects should not visit them they are able tofertilise themselves. 2Absence of Colour in Wind-fertilised FlowersAs supplementing these curious facts, showing the relationof colour in flowers to the need of the visits of insects tofertilise them, we have the remarkable, and, on any othertheory, utterly inexplicable circumstance that in all the numerouscases in which plants are fertilised by the agency ofthe wind they never have specially coloured floral envelopes.Such are our pines, oaks, poplars, willows, beeches,1 Nature, vol. xi. pp. 32, 110. Ib., vol. ix. p. 164.
vi COLOURS OF PLANTS 405and hazel, our nettles, grasses, sedges, and many others. Insome of these the male flowers are very conspicuous, as thecatkins of the willows, and these secrete honey and attractnumerous insects at a season when there are few other flowers,and thus secure cross -fertilisation. Sedges and grasses arealso occasionally visited by insects.The same Theory of Colour applicable to Animals and PlantsItmay be thought that this absence of colour where it isnot wanted isopposed to the view maintained in the earlierpart of the preceding chapter, that colour is normal and isconstantly tending to appear in natural objects. It must beremembered, however, that the green colour of foliage, due tochlorophyll, prevails throughout the greater part of the vegetablekingdom, and has, almost certainly, persisted throughlong geological periods. It has thus acquired a fixity ofcharacter which cannot be readily disturbed ; and, as a matterof fact, we find that colour rarely appears in plants except inassociation with a considerable modification of leaf -texture,such as occurs in the petals and coloured sepals of flowers.Wind -fertilised plants never have such specially organisedfloral envelopes, and, in most cases, are entirely without acalyx or corolla. The connection between modification ofleaf-structure and colour is further seen in the greater amountand variety of colour in irregular than in regular flowers.The latter, which are least modified, have generally uniformor but slightly varied colours, while the former, which haveundergone great modification, present an immense range ofcolour and marking, culminating in the spotted and variegatedflowers of such groups as the Scrophularinese andOrchideae. The same laws as to the conditions of a maximumproduction of colour are thus found to obtain both in plantsand animals.Relation of the Colours of Flowers and their GeographicalDistributionThe adaptation of flowers to be fertilised by insectsoften to such an extent that the very existence of the speciesdepends upon it has had an important influence on the distributionof plants and the general aspects of vegetation.
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vi COLOURS OF PLANTS 405and hazel, our nettles, grasses, sedges, and many others. Insome of these the male flowers are very conspicuous, as thecatkins of the willows, and these secrete honey and attractnumerous insects at a season when there are few other flowers,and thus secure cross -fertilisation. Sedges and grasses arealso occasionally visited by insects.The same Theory of Colour applicable to Animals and PlantsItmay be thought that this absence of colour where it isnot wanted isopposed to the view maintained in the earlierpart of the preceding chapter, that colour is normal and isconstantly tending to appear in natural objects. It must beremembered, however, that the green colour of foliage, due tochlorophyll, prevails throughout the greater part of the vegetablekingdom, and has, almost certainly, persisted throughlong geological periods. It has thus acquired a fixity ofcharacter which cannot be readily disturbed ; and, as a matterof fact, we find that colour rarely appears in plants except inassociation with a considerable modification of leaf -texture,such as occurs in the petals and coloured sepals of flowers.Wind -fertilised plants never have such specially organisedfloral envelopes, and, in most cases, are entirely without acalyx or corolla. The connection between modification ofleaf-structure and colour is further seen in the greater amountand variety of colour in irregular than in regular flowers.The latter, which are least modified, have generally uniformor but slightly varied colours, while the former, which haveundergone great modification, present an immense range ofcolour and marking, culminating in the spotted and variegatedflowers of such groups as the Scrophularinese andOrchideae. The same laws as to the conditions of a maximumproduction of colour are thus found to obtain both in plantsand animals.Relation of the Colours of Flowers and their GeographicalDistributionThe adaptation of flowers to be fertilised by insectsoften to such an extent that the very existence of the speciesdepends upon it has had an important influence on the distributionof plants and the general aspects of vegetation.