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354 TROPICAL NATUREMr. Darwin's Descent of Man, chapters x. to xviii., and tochapters iii. iv. and vii. of the first portion of the presentvolume.Normal ColoursThe fourth group of normally coloured animals includesall species which are brilliantly or conspicuously coloured inboth sexes, and for whose particular colours we can assign nofunction or use. 1 It comprises an immense number of showybirds, such as kingfishers, barbets, toucans, lories, tits, andstarlings ; among insects most of the largest and handsomestbutterflies, innumerable bright-coloured beetles, locusts, dragonflies,and hymenoptera ;a few mammalia, as the zebras ;agreat number of marine fishes ;thousands of striped andspotted caterpillars ;and abundance of mollusca, star-fish, andother marine animals. Among these we have included somewhich, like the gaudy caterpillars, have warning colours but;as that theory does not explain the particular colours or thevaried patterns with which they are adorned, it is best toinclude them also in this class. It is a suggestive fact thatall the brightly-coloured birds mentioned above build in holesor form covered nests, so that the females do not need thatprotection during the breeding season which I believe to beone of the chief causes of the dull colour of female birds whentheir partners are gaily coloured. This subject is fully arguedin chapter vi. of the present volume.Leaving the colours of flowers to be discussed in anotherchapter, we will now consider how the general facts of colourhere sketched out can be explained. We have first to inquirewhat is colour, and how it isproduced j secondly, what isknown of the causes of change of colour ; and, lastly,whattheory best accords with the whole assemblage of facts.The Nature of ColourThe sensation of colour is caused by vibrations or undulationsof the ethereal medium of different lengths and velocities.The whole body of vibrations caused by the sun is termed* Distinctness of marking for purposes of recognition is probably the usein all cases. See p. 367, and Darwinism, p. 217.

v COLOURS OF ANIMALS 355radiation, or, more commonly, rays ;and consists of sets ofwaves which vary considerably in their dimensions and rateof recurrence, but of which the middle portion only is capableof exciting in us sensations of light and colour. Beginningwith the largest waves, which recur at the longest intervals,we have first those which produce heat-sensations only ;asthey get smaller and recur quicker, we perceive a dull redcolour ;and as the waves increase in rapidity and diminish insize, we get successively sensations of orange, yellow, green,blue, indigo, and violet, all fading imperceptibly into eachother. Then come more invisible rays, of shorter wavelengthand quicker recurrence, which produce, solely orchiefly, chemical effects. The red rays, which first becomevisible, have been ascertained to recur at the rate of 458millions of millions times in a second, the length of each wave-being ?6 ^ 60 of an inch ;while the violet rays, which lastremain visible, recur 727 millions of millions times per second,and have a wave-length of a ^ lfl of an inch. Although thewaves recur at different rates, they are all propagated throughthe ether with the same velocity (192,000 miles per second) ;just as different musical sounds, which are produced bywaves of air of different lengths and rates of recurrence, travelat the same speed, so that a tune played several hundredyards off reaches the ear in correct time. There are, therefore,an almost infinite number of different colour-producingundulations, and these may be combined in an almost infinitevariety of ways, so as to excite in us the sensation of all thevaried colours and tints we are capable of perceiving. Whenall the different kinds of rays reach us in the proportion inwhich they exist in the light of the sun, they produce thesensation of white.If the rays which excite the sensation ofany one colour are prevented from reaching us, the remainingrays in combination produce a sensation of colour often veryfar removed from white. Thus green rays being abstractedleave purple light; blue, orange-red light; violet, yellowishgreenlight, and so on. These pairs are termed complementarycolours. And if portions of differently colouredlights are abstracted in various degrees, we have produced allthose infinite gradations of colours, and all those varied tintsand hues which are of such use to us in distinguishing

354 TROPICAL NATUREMr. Darwin's Descent of Man, chapters x. to xviii., and tochapters iii. iv. and vii. of the first portion of the presentvolume.Normal ColoursThe fourth group of normally coloured animals includesall species which are brilliantly or conspicuously coloured inboth sexes, and for whose particular colours we can assign nofunction or use. 1 It comprises an immense number of showybirds, such as kingfishers, barbets, toucans, lories, tits, andstarlings ; among insects most of the largest and handsomestbutterflies, innumerable bright-coloured beetles, locusts, dragonflies,and hymenoptera ;a few mammalia, as the zebras ;agreat number of marine fishes ;thousands of striped andspotted caterpillars ;and abundance of mollusca, star-fish, andother marine animals. Among these we have included somewhich, like the gaudy caterpillars, have warning colours but;as that theory does not explain the particular colours or thevaried patterns with which they are adorned, it is best toinclude them also in this class. It is a suggestive fact thatall the brightly-coloured birds mentioned above build in holesor form covered nests, so that the females do not need thatprotection during the breeding season which I believe to beone of the chief causes of the dull colour of female birds whentheir partners are gaily coloured. This subject is fully arguedin chapter vi. of the present volume.Leaving the colours of flowers to be discussed in anotherchapter, we will now consider how the general facts of colourhere sketched out can be explained. We have first to inquirewhat is colour, and how it isproduced j secondly, what isknown of the causes of change of colour ; and, lastly,whattheory best accords with the whole assemblage of facts.The Nature of ColourThe sensation of colour is caused by vibrations or undulationsof the ethereal medium of different lengths and velocities.The whole body of vibrations caused by the sun is termed* Distinctness of marking for purposes of recognition is probably the usein all cases. See p. 367, and Darwinism, p. 217.

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