PDF - Wallace Online
PDF - Wallace Online PDF - Wallace Online
148 NATURAL SELECTION vnmoths ;and this would be a necessary result from the fact thatnature ever fluctuates about a mean, or that in every generationthere would be flowers with longer and shorter nectaries,and moths with longer and shorter probosces than the average.No doubt there are a hundred causes that might have checkedthis process before it had reached the point of developmentat which we find it. If, for instance, the variation in thequantity of nectar had been at any stage greater than thevariation in the length of the nectary, then smaller mothscould have reached it and have effected the fertilisation. Orif the growth of the probosces of the moths had from othercauses increased quicker than that of the nectary, or if theincreased length of proboscis had been injurious to them inany way, or if the species of moth with the longest proboscishad become much diminished by some enemy or other unfavourableconditions, then, in any of these cases, the shorternectaried flowers, which would have attracted and could havebeen fertilised by the smaller kinds of moths, would have hadthe advantage. And checks of a similar nature to these nodoubt have acted in other parts of the world, and have preventedsuch an extraordinary development of nectary as hasbeen produced by favourable conditions in Madagascar only,and in one single species of orchid. I may here mention thatsome of the large Sphinx moths of the tropics have proboscesnearly as long as the nectary of Angrsecum sesquipedale. Ihave carefully measured the proboscis of a specimen of Macrosilacluentius from South America, in the collection of the BritishMuseum, and find it to be nine inches and a quarter long!One from tropical Africa (Macrosila morganii) is seven inchesand a half. A species having a proboscis two or three incheslonger could reach the nectar in the largest flowers of Angraecumsesquipedale, whose nectaries vary in length from tento fourteen inches. That such a moth exists in Madagascarmay be safely predicted ;and naturalists who visit that islandshould search for it with as much confidence as astronomerssearched for the planet Neptune, and I venture to predictthey will be equally successful !Now, instead of this beautiful self-acting adjustment, theopposing theory is, that the Creator of the universe, by adirect act of His will, so disposed the natural forces influencing
vii CREATION BY LAW 149the growth of this one species of plant as to cause its nectaryto increase to this enormous length ;and at the same time,by an equally special act, determined the flow of nourishmentin the organisation of the moth, so as to cause its proboscis toincrease in exactly the same proportion, having previously soconstructed the Angrsecum that it could only be maintainedin existence by the agency of this moth. But what proof isgiven or suggested that this was the mode by which the adjustmenttook place None ? whatever, except a feeling thatthere is an adjustment of a delicate kind, and an inability tosee how known causes could have produced such an adjustment.I believe I have shown, however, that such anadjustment is not only possible but inevitable, unless at somepoint or other we deny the action of those simple laws whichwe have already admitted to be but the expressions of existingfacts.Adaptation brought about by General LawsIt is difficult to find anything like parallel cases in inorganicnature, but that of a river may perhaps illustrate the subjectin some degree. Let us suppose a person totally ignorant ofmodern geology to study carefully a great river system. Hefinds in its lower part a deep broad channel filled to thebrim, flowing slowly through a flat country and carrying outto the sea a quantity of fine sediment. Higher upit branchesinto a number of smaller channels, flowing alternately throughflat valleys and between high banks sometimes he finds a;deep rocky bed with perpendicular walls, carrying the waterthrough a chain of hills where the stream is narrow he finds; it deep, where wide shallow. Farther up still, he comes to amountainous region, with hundreds of streams and rivulets,each with its tributary rills and gullies, collecting the waterfrom every square mile of surface, and every channel adaptedto the water that it has to carry.He finds that the bed ofevery branch and stream and rivulet has a steeper andsteeper slope as it approaches its sources, and is thus enabledto carry off the water from heavy rains, and to bear awaythe stones and pebbles and gravelthat would otherwise blockup its course. In every part of this system he would seeexact adaptationof means to an end. He would say thatthis system of channels must have been itdesigned,answers
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vii CREATION BY LAW 149the growth of this one species of plant as to cause its nectaryto increase to this enormous length ;and at the same time,by an equally special act, determined the flow of nourishmentin the organisation of the moth, so as to cause its proboscis toincrease in exactly the same proportion, having previously soconstructed the Angrsecum that it could only be maintainedin existence by the agency of this moth. But what proof isgiven or suggested that this was the mode by which the adjustmenttook place None ? whatever, except a feeling thatthere is an adjustment of a delicate kind, and an inability tosee how known causes could have produced such an adjustment.I believe I have shown, however, that such anadjustment is not only possible but inevitable, unless at somepoint or other we deny the action of those simple laws whichwe have already admitted to be but the expressions of existingfacts.Adaptation brought about by General LawsIt is difficult to find anything like parallel cases in inorganicnature, but that of a river may perhaps illustrate the subjectin some degree. Let us suppose a person totally ignorant ofmodern geology to study carefully a great river system. Hefinds in its lower part a deep broad channel filled to thebrim, flowing slowly through a flat country and carrying outto the sea a quantity of fine sediment. Higher upit branchesinto a number of smaller channels, flowing alternately throughflat valleys and between high banks sometimes he finds a;deep rocky bed with perpendicular walls, carrying the waterthrough a chain of hills where the stream is narrow he finds; it deep, where wide shallow. Farther up still, he comes to amountainous region, with hundreds of streams and rivulets,each with its tributary rills and gullies, collecting the waterfrom every square mile of surface, and every channel adaptedto the water that it has to carry.He finds that the bed ofevery branch and stream and rivulet has a steeper andsteeper slope as it approaches its sources, and is thus enabledto carry off the water from heavy rains, and to bear awaythe stones and pebbles and gravelthat would otherwise blockup its course. In every part of this system he would seeexact adaptationof means to an end. He would say thatthis system of channels must have been itdesigned,answers