PDF - Wallace Online
PDF - Wallace Online PDF - Wallace Online
138 NATURAL SELECTIONgreater danger to which she is exposed and her greater importanceto the existence of the species and this she ; alwaysacquires, in one way or another, through the action of naturalselection.In his Origin of Species, fourth edition, p. 241, Mr. Darwinrecognises the necessity for protection as sometimes being a1cause of the obscure colours of female birds ;but he does notseem to consider it so very important an agent in modifyingcolour as I am disposed to do. In the same paragraph (p.240) he alludes to the fact of female birds and butterfliesbeing sometimes very plain, sometimes as gay as the males ;but, apparently, considers this mainly due to peculiar laws ofinheritance, which sometimes continue acquired colour in theline of one sex only, sometimes in both. Without denyingthe action of such a law (which Mr. Darwin informs me hehas facts to support), I impute the difference, in the greatmajority of cases, to the greater or less need of protection inthe female sex in these groups of animals.This need was seen to exist a century ago by the Hon.Daines Barrington, who, in the article already quoted (see p.104), after alluding to the fact that singing birds are all small,and suggesting (but I think erroneously) that this may havearisen from the difficulty larger birds would have in concealingthemselves if they called the attention of their enemies byloud notes, goes on thus :" I should rather conceive it is forthe same reason no hen bird sings, because this talent wouldbe still more dangerous during incubation, which may possiblyalso account for the inferiority in point of plumage." This is acurious anticipation of the main idea on which this essay isfounded. It has been unnoticed for near a century, and myattention was only recently called to itby Mr. Darwin himself.ConclusionTo some persons it will perhaps appear that the causes towhich I impute so much of the external aspect of nature aretoo simple, too insignificant, and too unimportant for such amighty Ayork.But I would ask them to consider that thegreat object of all the peculiarities of animal structure is topreserve the life of the individual, and to maintain the exist-1This passage is omitted in the sixth edition.
vi A THEORY OF BIRDS' NESTS 139ence of the species. Colour has hitherto been too often lookedupon as something adventitious and superficial, somethinggiven to an animal not to be useful to itself, but solely togratify man or even superior beings to add to the beautyand ideal harmony of nature. If this were the case, then, itis evident that the colours of organised beings would be anexception to most other natural phenomena. They would notbe the product of general laws, or determined by ever-changingexternal conditions and we must;give up all inquiry intotheir origin and causes, since (by the hypothesis) they aredependent on a Will whose motives must ever be unknown tous. But, strange to say, no sooner do we begin to examineand classify the colours of natural objects, than we find thatthey are intimately related to a variety of other phenomena,and are, like them, strictly subordinated to general laws. Ihave here attempted to elucidate some of these laws in thecase of birds, and have shown how the mode of nidificationhas affected the colouring of the female sex in this group. Ihave before shown to how great an extent, and in how manyways, the need of protection has determined the colours ofinsects, and of some groups of reptiles and mammalia, and Iwould now call particular attention to the fact that the gaytints of flowers, so long supposed to be a convincing proof thatcolour has been bestowed for other purposes than the good ofits possessor, have been shown by Mr. Darwin to follow thesame great law of utility. Flowers do not often need protection,but very often require the aid of insects to fertilisethem, and maintain their reproductive powers in the greatestvigour. Their gay colours attract insects, as do also theirsweet odours and honeyed secretions ;and that this is themain function of colour in flowers is shown by the strikingfact that those flowers which can be perfectlyfertilised bythe wind, and do not need the aid of insects, rarely or neverhave gaily-coloured flowers.This wide extension of the general principle of utility tothe colours of such varied groups, both in the animal andvegetable kingdoms, compels us to acknowledge that the"reign of law" has been fairly traced into this stronghold ofthe advocates of specialcreation. And to those who opposethe explanation now given of the various facts bearing upon
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138 NATURAL SELECTIONgreater danger to which she is exposed and her greater importanceto the existence of the species and this she ; alwaysacquires, in one way or another, through the action of naturalselection.In his Origin of Species, fourth edition, p. 241, Mr. Darwinrecognises the necessity for protection as sometimes being a1cause of the obscure colours of female birds ;but he does notseem to consider it so very important an agent in modifyingcolour as I am disposed to do. In the same paragraph (p.240) he alludes to the fact of female birds and butterfliesbeing sometimes very plain, sometimes as gay as the males ;but, apparently, considers this mainly due to peculiar laws ofinheritance, which sometimes continue acquired colour in theline of one sex only, sometimes in both. Without denyingthe action of such a law (which Mr. Darwin informs me hehas facts to support), I impute the difference, in the greatmajority of cases, to the greater or less need of protection inthe female sex in these groups of animals.This need was seen to exist a century ago by the Hon.Daines Barrington, who, in the article already quoted (see p.104), after alluding to the fact that singing birds are all small,and suggesting (but I think erroneously) that this may havearisen from the difficulty larger birds would have in concealingthemselves if they called the attention of their enemies byloud notes, goes on thus :" I should rather conceive it is forthe same reason no hen bird sings, because this talent wouldbe still more dangerous during incubation, which may possiblyalso account for the inferiority in point of plumage." This is acurious anticipation of the main idea on which this essay isfounded. It has been unnoticed for near a century, and myattention was only recently called to itby Mr. Darwin himself.ConclusionTo some persons it will perhaps appear that the causes towhich I impute so much of the external aspect of nature aretoo simple, too insignificant, and too unimportant for such amighty Ayork.But I would ask them to consider that thegreat object of all the peculiarities of animal structure is topreserve the life of the individual, and to maintain the exist-1This passage is omitted in the sixth edition.