A C. Canadian Journal of Earth Sciences, 2008,45(2): 213-229Hundreds of shell borings of different originand displaying variable patterns were found instrophomenide, pentameride, and orthidebrachiopods of five Paleozoic localities in northernand eastern Canada. The borings were analyzedusing simple statistics as well as clusterand nonmetric multidimensional scaling analyses.At the Ordovician Anticosti Island locality,all borings are parasitic or post-mortem in origin,while at the Wenlock-Ludlow Baillie-HamiltonIsland, almost all borings are predatory. At theremaining three localities, borings represent amix of predatory, parasitic, and post-mortemdomichnial borings in all three brachiopod taxa,the proportions of which were controlled largelyby brachiopod shell morphology and paleoecology.For the strophomenides, predatory boringscan be segregated from parasitic and postmortemdomichnial using simple and multivariatestatistical analyses. Sowerbyella-type strophomenideshave a higher proportion of predatoryborings at the Lochkovian localities than atthe Ordovician localities, while the reverse istrue for the Strophomena-type strophomenides.In pentamerides and orthides, very few predatoryborings are identified; most borings were emplacedby parasitic or post-mortem domichnialborers. In pentamerides, this is due to the internalstructure of the shells, which elevated themuscles of the organism above the shell floor,rendering them inaccessible to boring predators.In orthides, more deliberate defense mechanismssuch as toxins or external ornamentation mayhave deterred predation. This study indicates thatthroughout the Paleozoic interactions betweenborers and brachiopods were complex but can beelucidated using a large sample size and statisticalanalyses.2008040035美 国 辛 辛 那 提 地 区 上 奥 陶 统 泥 质 和 介 壳 层 发育 中 的 沉 积 供 给 与 风 暴 漂 选 = Sediment supplyversus storm winnowing in the developmentof muddy and shelly interbeds from the UpperOrdovician of the Cincinnati region, USA. ( 英文 ). Dattilo B F; Brett C E; Tsujita C J; FairhurstR. Canadian Journal of Earth Sciences, 2008,45(2): 243-265Shell-bed development can be a product ofcomplex sedimentological and biological factors.The Upper Ordovician sediments near Cincinnati,Ohio constitute a succession of thinly interbeddedshelly carbonates and mudrocks. Despiteyears of study, the development of Cincinnatianshell beds and metre-scale cycles has, until recently,been attributed solely to storm reworking.This “storm-winnowing model” treats shells aspassive sedimentary clasts, ignoring other factorsof shell-bed development. A recently proposedalternative is Brett and Algeo's idea thatthese shell beds grew during long periods ofnormally low sedimentation, while most mudaccumulated during brief periods of high sedimentation.Under this “episodic starvationmodel,” any storms would winnow pre-existingmuds and shell beds alike. We tested both modelsin the Edenian-Maysvillian (early to midKatian) strata of the Cincinnati region by compilingobservations on their petrologic, taphonomic,and paleoecologic characteristics.The storm-winnowing model does not explainseveral observed features that the episodic starvationmodel does, including (i)storm-relatedsedimentary structures in mudrocks and limestones;(ii) lack of a sufficiently fossiliferousprecursor deposit to winnow; (iii)deep-waterfaunas in grainstones; (iv)mixed taphonomicconditions of shell-bed fossils; (v) ubiquitousdiscontinuity surfaces; (vi)carbonate concretionhorizons; (vii)unwinnowed shell beds; and(viii)micrite in packstones. Episodic starvation isa superior explanation because it explains all ofthese features and allows for the complex interplayof other environmental and biological factorsthat contribute to shell-bed growth. It mayalso be applicable to other deposits, previouslyinterpreted as tempestites.2008040036埃 迪 卡 拉 纪 海 洋 和 动 物 的 协 同 演 化 = On thecoevolution of Ediacaran oceans and animals.( 英 文 ). Shen Y A; Zhang T G; Hoffman P F.proceedings of the National Academy of Sciences,2008, 105(21): 7376-7381 3 图 版 .Fe speciation and S-isotope of pyrite datafrom the terminal Proterozoic Sheepbed Formationin Canada and Doushantuo Formation inChina reveal that ocean deep waters were anoxicafter the global glaciations (snowball Earth) ending635 million years ago, but that marine sulfateconcentrations and inferentially atmosphericoxygen levels were higher than before the glaciations.This supports a long-postulated link betweenoxygen levels and the emergence ofeumetazoa. Subsequent ventilation of the deepocean, inferred from shifts in Fe speciation inNewfoundland (previously published data) andwestern Canada (this report), paved the way forEdiacaran macrobiota to colonize the deep seafloors.2008040037动 物 的 黎 明 : 新 元 古 代 - 寒 武 纪 动 物 化 石 记 录的 演 化 和 古 生 态 模 式 = The dawn of animallife: Evolutionary and palaeoecological patternsin the Neoproterozoic-Cambrian animal fossilrecord. ( 英 文 ). Dornbos S Q; Xiao Shuhai. Pa-12
laeogeography Palaeoclimatology Palaeoecology,2008, 258(3): 135-1372008040038早 寒 武 世 帽 天 山 页 岩 生 物 群 的 群 落 古 生 态学 : 鳃 曳 动 物 门 蠕 虫 的 生 态 优 势 = Communitypalaeoecology of the early Cambrian MaotianshanShale biota: Ecological dominance ofpriapulid worms. ( 英 文 ). Dornbos S Q; ChenJunyuan. Palaeogeography PalaeoclimatologyPalaeoecology, 2008, 258(3): 200-212The early Cambrian soft-bodied MaotianshanShale (Chengjiang) biota of Yunnan Province,China provides a critical glimpse of animal lifeduring the heart of the Cambrian radiation. TheShankou biota is the focus of this current work,and 9963 specimens from it have been examinedand tallied. This collection of fossils represents atime-averaged assemblage of unknown durationfrom a 10-meter thick sequence. The predominantlybenthic community recovered from thissection is presumably autochthonous. Thiscommunity was buried through several hundredmillimeter to centimeter thick burial events.Only specimens interpreted as buried alive,based on either soft-part preservation or fullyarticulated skeletons, were counted. Algae weretoo fragmented to be counted. To avoid includingsynonymous species, diversity was evaluatedat the genus level. A total of 57 genera from 9phyla, encompassing 14 ecological groups, werefound in this assemblage. The results reveal thatthe three most abundant genera comprise 43.2%of all specimens: the tubiform priapulid wormParaselkirkia (16.0%), the diminutive priapulidworm Sicyophorus (14.3%), and the brachiopodHeliomedusa (12.9%). No other genera totalmore than 9% of specimens. At the phylum level,there is an interesting dichotomy between taxonomicdiversity and ecological dominance. Thearthropods are the most diverse phylum (37% ofgenera) and rank second in relative abundance ofspecimens (26.3%). The priapulids, however,with only 17.5% of genera is the most abundantgroup in the assemblage (43.2%). All otherphyla, excluding brachiopods (19.6%), representonly 10.9% of the assemblage. Because of theirsheer numerical abundance, this study indicatesthat priapulid worms may have exerted moreinfluence on energy flow and community structurethan other phyla in this particular trophicweb. This result contrasts strongly with bothtraditional views of Maotianshan Shale biotapalaeoecology, which often claimed arthropoddominance based solely on taxonomic diversity,and the palaeoecology of modern priapulids,which are relegated to marginal marine settings.This result also demonstrates the importance ofcollecting quantitative relative abundance datawhen performing palaeoecological investigations.2008040039中 国 南 部 贵 州 中 寒 武 世 凯 里 生 物 群 高 茎 棘 皮动 物 的 栖 息 策 略 = Settling strategy of stalkedechinoderms from the Kaili Biota (middle Cambrian),Guizhou Province, South China. ( 英 文 ).Lin J; Ausich W; Zhao Y. Palaeogeography PalaeoclimatologyPalaeoecology, 2008, 258(3):213-221A total of 124 articulated echinoderms, includingSinoeocrinus lui (n = 30), “S. globus”(n = 92), one indeterminate gogiid, and one indeterminateedrioasteroid, from the Kaili Formation(Middle Cambrian), eastern Guizhou, Chinawas examined. Among them, 73% of gogiidechinoderms are preserved attached to skeletalsubstrates. Suitable skeletal substrates for gogiidholdfasts in the Kaili Biota are organophosphaticbrachiopods; large trilobite fragments, includingcranidia, free cheeks, thoracic segments, andpygidia; hyoliths; and Scenella shells. The highfrequency of “S. globus” attached to organophosphaticbrachiopods is due to the high richnessof organophosphatic brachiopods ratherthan a host-specific association in the Kaili Biota.However, the possible discrimination againstattachment to small trilobites, such as Pagetia,requires further investigation. Based on 11 handsamples, echinoderm population density (EPD)varies from 0.3 to 4.5 echinoderms per 50 grindpoints (0.06 to 0.96 echinoderms per cm 2 ), dependingon the percent coverage of skeletal debris.Crowding effect is evident when EPD isapproximately 0.32 echinoderms per cm 2 and theskeletal coverage is less than 20%.2008040040布 尔 吉 斯 页 岩 中 大 型 叶 脚 类 动 物 群 落 的 古 生态 学 = Paleoecology of the Greater PhyllopodBed community, Burgess Shale. ( 英 文 ). Caron J;Jackson D A. Palaeogeography PalaeoclimatologyPalaeoecology, 2008, 258(3): 222-256To better understand temporal variations inspecies diversity and composition, ecologicalattributes, and environmental influences for theMiddle Cambrian Burgess Shale community, westudied 50,900 fossil specimens belonging to158 genera (mostly monospecific and nonbiomineralized)representing 17 major taxonomicgroups and 17 ecological categories. Fossilswere collected in situ from within 26 massivesiliciclastic mudstone beds of the GreaterPhyllopod Bed (Walcott Quarry — Fossil Ridge).Previous taphonomic studies have demonstratedthat each bed represents a single obrution eventcapturing a predominantly benthic communityrepresented by census- and time-averaged assemblages,preserved within habitat. The GreaterPhyllopod Bed (GPB) corresponds to an esti-13
- Page 2 and 3: 第 4 卷 第 4 期古 生 物 学
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Pitted surfaces associated with wal
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Saanich than exists today sedimento
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2008040208大 堡 礁 区 域 绿
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was pulsed, with major declines usu
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mycelium, was observed where reticu
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suggest that reticulopodia are self
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to OIS 3, compared to the more stab
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Patterson R T. Journal of Foraminif
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known xenophyophore species grouped
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aminiferal Research, 2001, 31(4): 2
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arctic environment over this critic
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Semeniuk T A. Journal of Foraminife
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sible to make production calculatio
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arcellacean strains from within the
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annual flux rates is calculated and
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corded in the Colli Berici (type lo
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most complete OAE 2 sections. While
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oceanic plate stratigraphy from gra
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A series of layers rich in organic
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lowermost Danian planktonic foramin
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characters shows that only one geno
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able for regional palaco-environmen
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margin of the southwest Caribbean.
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Brachyelasma, Amplexobrachyelasma,
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litic biozones lower than their FAD
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The middle Permian Cryptospirifer f
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石 科 = Ammonoid faunas from the
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Buchiid bivalves are geographically
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velopment of both the sedimentary a
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China. ( 英 文 ). Huang Diying; N
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德 国 南 部 上 侏 罗 统 (Ox
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shelf basin off the northern coast
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like base referred to Primocandelab
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鱼 类2008040413大 西 洋 鳕
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long as the humerus plus radius, ve
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members in both size and abundance
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ple's Republic of China. ( 英 文
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K-P transition in Brazil. ( 英 文
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wide tooth crowns, a sub-rectangula
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2008040459罗 马 尼 亚 特 兰
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logical descriptions of all availab
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东 副 特 提 斯 晚 萨 马 期
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S; Vaiani S C. Rivista Italiana di
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adiocarbon dating the arrival of th
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able problems, particularly differe
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done with glyphs on paper. In this
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obtained through extensive field-ba
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noli G; Maletz J; Stouge S. Canadia
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法 国 蒙 塔 格 尼 努 尔 中
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presence of well-rounded, coarse-sa
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缅 甸 北 部 Hukawng 峡 谷 琥
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graphy and Geological Correlation,
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plex,northwestern Iran. ( 英 文 )
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松 辽 盆 地 晚 白 垩 世 青
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ous nannoplankton) points of view.O
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the"Yanshan Movement".During the Mi
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geochemical approach discussed in t
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sediments from the northern shelf o
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意 大 利 北 亚 平 宁 Lamone
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nano record further into the early
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lankovitch-scale frequency. Analyse