Identification of American Herring Gull in a western European context

Identification of American Herring Gull in a western European context Identification of American Herring Gull in a western European context

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Identification of American Herring Gull in a western European context40 American Herring Gull / Amerikaanse Zilvermeeuw Larus smithsonianus, first-winter, Boston, Massachusetts,USA, January 2001 (Pat Lonergan). Very worn individual, with second-generation scapulars having pattern unlikethat normally exhibited by European Herring Gulls L argentatus.ally be more uniform (and consequently darker looking),thus resembling smithsonianus (plate 21 and 34).Solid darkness on lower hindneck and upper mantleTypically, smithsonianus exhibits a more uniformlybrownish lower hindneck and upper mantle thatmerges with the uniform brownish underparts (plate 28and 36).Greyness of breast-sides and flanks As part of the postjuvenilemoult of body-feathers, many first-wintersmithsonianus acquire plain, contrastingly slate-greycolouredfeathers on the breast-sides and flanks, graduallyextending to the rest of the underparts (plate 27).Possibly because of their usually more mottled underparts,European Herring Gulls undergoing the samemoult show much more subtle contrast between theold (brownish) and new (more greyish) feathers.Pale-headed appearance Many smithsonianus acquirea pale head in late winter as a result of wear (Howell2001) and due to their dark body this feature may drawattention to a smithsonianus among a flock of EuropeanHerring Gulls. The importance of this feature has beenoverstated a little as only a small proportion really ispale-headed (plate 24, 29 and 36) and darker-bodiedEuropean birds are likely to also occasionally lookpale-headed for precisely the same reasons (plate 16and 21). European Herring Gulls from the easternBaltic area are often strikingly pale-headed in winter(Klaus Malling Olsen pers comm).Scapular pattern The range of individual variation infirst-winter scapular-markings exhibited by both smithsonianusand European Herring Gulls, and the degreeof overlap, make it very difficult to identify any particularpatterns that might be considered ‘exclusive’. Thereare, however, certain characteristic patterns in smithsonianusthat are not so usual in their European counterparts.It is important, here, to distinguish betweenoften-retained juvenile scapulars (usually, the rearmostlarger feathers), which are plain, brownish, somewhatworn and with pointed tips, and (freshly) moulted firstwinterfeathers, which have broader, more roundedtips. The most distinctive of these (again, usually seenamong the larger rearmost and lower row(s) of feathers)are rather dark and plain, with or without a diffusedarker centre (plate 38). Due, perhaps, to a tendency inmany smithsonianus for the post-juvenile moult of thescapulars (in which the juvenile scapulars are replacedwith first-winter feathers) to be a rather protracted process,there is often more of a variety of scapular patternsin the one bird than is generally the case inEuropean birds (plate 25 and 31); the explanation forthis is that the appearance of feathers in the same generationcan change depending on the time of year theyare moulted (Howell 2001). In most European HerringGulls, the pattern of the first-winter scapulars tends tobe rather consistent, each individual feather exhibitingmuch the same markings as the next, the overall effect16

Identification of American Herring Gull in a western European contextbeing of a series of regular transverse pale and darkbars (plate 35). However, there are many exceptions tothese general tendencies and, at best, certain scapularpatterns should be regarded as offering little more thanmarginal supporting evidence in the identification ofvagrant first-year smithsonianus in Europe.Underwing-coverts The uniformity of the axillaries(especially) and underwing-coverts, and the general‘smokiness’ with a lack of obvious patterning, can bestriking in smithsonianus. In argentatus and argenteus,these areas tend to be paler in tone and more mottledin texture. The underwing-coverts of graellsii, however,can be very like smithsonianus but several other differencesfrom smithsonianus (see above) usually precludeserious confusion.Bill-colour There is a tendency for both smithsonianusand argentatus to develop a pale base to the bill quiteearly in their first winter, with the most extreme birdsapproaching first-year Glaucous Gull in this respect(plate 24 and 36). In argenteus, the contrast in the billpattern tends to be more subdued until later in thewinter.Second-winter (plate 41-58, figure 2)If it often seems as if no two first-winter smithsonianusare quite alike, second-winter birds areeven more variable. Some second-winter individualsare, at first glance, extremely first-winterlike,due to a complete lack of clear grey in thescapulars and extensively brownish underparts(plates 41, 48 and 54). Obviously pale-eyed individualscan be more easily aged but some birdsdo not develop a pale iris until late winter, andeven then it may be difficult to discern in poorlight or at moderate range (plate 43). In addition,most show more intricately and irregularly patternedgreater coverts and tertial-fringes (much assecond-winter European Herring Gulls), a clearlypale basal two-thirds to the bill and, at closerange, the primary-tips are slightly more roundedthan in first-years. Second-winters that differmore obviously from first-winters have at leastsome clear grey scapulars (plate 51, 53 and 55)and, much as in European Herring Gull, there isconsiderable individual variation between thesetwo types.The following characters should help resolvethe identity of second-winter type smithsonianusin Europe:Solid darkness on lower hindneck and upper mantle,and underbody As in first-years, there is a much greatertendency for second-year smithsonianus to havedense brownish colouration on the lower hindneck andupper mantle, extending onto the underparts, thanthere is in European Herring Gulls of the same age(plate 45, 51 and 53). This brownish ‘wash’ is usuallyless intense, more mottled than in first-years but isoften still strong enough to attract attention. SecondwinterEuropean Herring Gulls are generally muchmore sparsely spotted or blotched with grey-brown inthese areas (plate 57).Tertial pattern There is a tendency in second-wintersmithsonianus for the tertials, especially the outer tertials,to average more extensively and solidly darkcentredthan in European Herring Gulls of the sameage, with a corresponding reduction in the extent ofpale at the tips. In European Herring Gulls, the tertialsare often either wholly ‘barred’ or have a small darkcentre and broad pale tip. However, there is variationin both, and considerable overlap, so the tertial patternshould only be used in conjunction with other supportingcharacters. We are not able to explain why, but ourobservations of second-year European Herring Gulls inlate summer and early autumn (plate 58) suggest thatmany at this time of year show darker and more smithsonianus-liketertials than at other times of the year.Undertail-coverts As with younger birds, the pattern ofthe undertail-coverts can be very useful as an aid toidentification. Many second-winter smithsonianus havean almost unchanged (from that of first-years) pattern ofintricate or closely spaced bars, in contrast with thewidely spaced bars and spots or almost unpatternedundertail-coverts of most argentatus/argenteus. Othershave solidly dark-centred feathers (plate 42, 47 and49), a pattern, so far we know, never found in EuropeanHerring Gulls.Pattern of rump and uppertail Often more obviousthan the undertail-coverts (described above), manysecond-winter smithsonianus mirror first-winters inshowing a more heavily patterned rump and uppertailcovertsand a practically all-dark uppertail (figure 2). Asthe season progresses, the rump becomes paler/whiter,the white rump being acquired through moult ratherthan wear; moulting birds may have a patchy mixture of(new) pure-white and patterned brown feathers. SecondwinterEuropean Herring Gulls tend to have less heavilypatterned rump and uppertail-coverts and many are predominantlyor wholly ‘white-rumped’. Paradoxically,they often have a much more extensively dark tail thanfirst-winters and this alone may prompt thoughts ofsmithsonianus. While there is extensive overlap in thetail patterns of second-winter smithsonianus andEuropean Herring Gulls, it seems that even the mostextreme examples of the latter usually show a narrowwedge (widest at the base) or ‘sliver’ of white along theouter edge to the tail (shown by some smithsonianus toobut a definite lack of white edges may be significant).Bill pattern and colour The great majority of secondwintersshow an extensively pale-based bill, the patterntypically resembling that of immature Glaucous Gull(plate 41, 42 and 45). The colour of the bill-base isvariable, sometimes flesh-pink (like most Europeanbirds) but often a more neutral greyish or horn colour,with or without a faintly greenish tinge (plate 47). Thelatter would be unusual in argenteus but is not uncommonin argentatus. Bill colour remains decidedly ‘immature-like’throughout winter and even early spring,being brownish, pinkish or pinkish-white basally.17

<strong>Identification</strong> <strong>of</strong> <strong>American</strong> <strong>Herr<strong>in</strong>g</strong> <strong>Gull</strong> <strong>in</strong> a <strong>western</strong> <strong>European</strong> <strong>context</strong>40 <strong>American</strong> <strong>Herr<strong>in</strong>g</strong> <strong>Gull</strong> / Amerikaanse Zilvermeeuw Larus smithsonianus, first-w<strong>in</strong>ter, Boston, Massachusetts,USA, January 2001 (Pat Lonergan). Very worn <strong>in</strong>dividual, with second-generation scapulars hav<strong>in</strong>g pattern unlikethat normally exhibited by <strong>European</strong> <strong>Herr<strong>in</strong>g</strong> <strong>Gull</strong>s L argentatus.ally be more uniform (and consequently darker look<strong>in</strong>g),thus resembl<strong>in</strong>g smithsonianus (plate 21 and 34).Solid darkness on lower h<strong>in</strong>dneck and upper mantleTypically, smithsonianus exhibits a more uniformlybrownish lower h<strong>in</strong>dneck and upper mantle thatmerges with the uniform brownish underparts (plate 28and 36).Greyness <strong>of</strong> breast-sides and flanks As part <strong>of</strong> the postjuvenilemoult <strong>of</strong> body-feathers, many first-w<strong>in</strong>tersmithsonianus acquire pla<strong>in</strong>, contrast<strong>in</strong>gly slate-greycolouredfeathers on the breast-sides and flanks, graduallyextend<strong>in</strong>g to the rest <strong>of</strong> the underparts (plate 27).Possibly because <strong>of</strong> their usually more mottled underparts,<strong>European</strong> <strong>Herr<strong>in</strong>g</strong> <strong>Gull</strong>s undergo<strong>in</strong>g the samemoult show much more subtle contrast between theold (brownish) and new (more greyish) feathers.Pale-headed appearance Many smithsonianus acquirea pale head <strong>in</strong> late w<strong>in</strong>ter as a result <strong>of</strong> wear (Howell2001) and due to their dark body this feature may drawattention to a smithsonianus among a flock <strong>of</strong> <strong>European</strong><strong>Herr<strong>in</strong>g</strong> <strong>Gull</strong>s. The importance <strong>of</strong> this feature has beenoverstated a little as only a small proportion really ispale-headed (plate 24, 29 and 36) and darker-bodied<strong>European</strong> birds are likely to also occasionally lookpale-headed for precisely the same reasons (plate 16and 21). <strong>European</strong> <strong>Herr<strong>in</strong>g</strong> <strong>Gull</strong>s from the easternBaltic area are <strong>of</strong>ten strik<strong>in</strong>gly pale-headed <strong>in</strong> w<strong>in</strong>ter(Klaus Mall<strong>in</strong>g Olsen pers comm).Scapular pattern The range <strong>of</strong> <strong>in</strong>dividual variation <strong>in</strong>first-w<strong>in</strong>ter scapular-mark<strong>in</strong>gs exhibited by both smithsonianusand <strong>European</strong> <strong>Herr<strong>in</strong>g</strong> <strong>Gull</strong>s, and the degree<strong>of</strong> overlap, make it very difficult to identify any particularpatterns that might be considered ‘exclusive’. Thereare, however, certa<strong>in</strong> characteristic patterns <strong>in</strong> smithsonianusthat are not so usual <strong>in</strong> their <strong>European</strong> counterparts.It is important, here, to dist<strong>in</strong>guish between<strong>of</strong>ten-reta<strong>in</strong>ed juvenile scapulars (usually, the rearmostlarger feathers), which are pla<strong>in</strong>, brownish, somewhatworn and with po<strong>in</strong>ted tips, and (freshly) moulted firstw<strong>in</strong>terfeathers, which have broader, more roundedtips. The most dist<strong>in</strong>ctive <strong>of</strong> these (aga<strong>in</strong>, usually seenamong the larger rearmost and lower row(s) <strong>of</strong> feathers)are rather dark and pla<strong>in</strong>, with or without a diffusedarker centre (plate 38). Due, perhaps, to a tendency <strong>in</strong>many smithsonianus for the post-juvenile moult <strong>of</strong> thescapulars (<strong>in</strong> which the juvenile scapulars are replacedwith first-w<strong>in</strong>ter feathers) to be a rather protracted process,there is <strong>of</strong>ten more <strong>of</strong> a variety <strong>of</strong> scapular patterns<strong>in</strong> the one bird than is generally the case <strong>in</strong><strong>European</strong> birds (plate 25 and 31); the explanation forthis is that the appearance <strong>of</strong> feathers <strong>in</strong> the same generationcan change depend<strong>in</strong>g on the time <strong>of</strong> year theyare moulted (Howell 2001). In most <strong>European</strong> <strong>Herr<strong>in</strong>g</strong><strong>Gull</strong>s, the pattern <strong>of</strong> the first-w<strong>in</strong>ter scapulars tends tobe rather consistent, each <strong>in</strong>dividual feather exhibit<strong>in</strong>gmuch the same mark<strong>in</strong>gs as the next, the overall effect16

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