Schriften zu Genetischen Ressourcen - Genres
Schriften zu Genetischen Ressourcen - Genres Schriften zu Genetischen Ressourcen - Genres
L. FRESE are reasons to assume that the section Procumbentes does not at all belong to the genus Beta but to a separate genus (WILLIAMS et al. 1976, JUNG et al. 1993). Nevertheless, major users of germplasm holdings are satisfied with the taxonomic system presented in Table 1 as it transfers all the information they need. Inventory and collecting The wild species of the genus Beta are native to Europe and adjacent areas. The section Beta is mainly distributed along the shores of the Mediterranean basin and along the Atlantic coast from the Canary Islands as the most southern outpost to the South of Sweden. Section Corollinae frequently occurs in Turkey and the adjacent Caucasus region with outposts in Daghestan and the Talysch mountain (Iran/ Azerbaijan). The only species of section Nanae is endemic in Greece where it grows in mountainous areas at elevations higher that 1800 m. Section Procumbentes has its major distribution area on the Canary Islands but can also be found in Southeast Spain and along the coast of Morocco where it is rare. Even wider distribution areas have cultivated forms. Where soil, water and temperature allow sugar beet is grown in almost every country of the northern hemisphere as well as in Chile. A similar large distribution area is known for the Leaf Beet Group which is adapted to a very broad range of growing conditions in Europe, Northern Africa, the Americas and in particular countries of the Asian region. In 1979 the first Beta germplasm collecting mission was funded by IPGRI and a number of additional missions followed with the objective to sample wild beets and landraces in the Mediterranean area. Between 1980 and 1990, large geographic gaps were closed by IPGRI and USDA/ARS funded missions (South Italy, Southern part of France, West Atlantic coast, British Islands, Ireland and Denmark) (DONEY et al. 1995) and the collecting data were entered into national databases. On the initiative of the ECP/GR programme, a European inventory of Beta collections was established in 1987, and in view of the effective collaboration achieved with the NPGS (USA) it was recommended to assume an international role. The International Database for Beta (FRESE and HINTUM 1989) contains information provided by 28 germplasm holdings in 24 countries. This central crop database currently stores passport data on 2,324 sugar beet, 863 fodder beet, 892 garden beet, 631 leaf beet and 4,022 wild beet accessions. The geographic coordinates of collecting sites were used for plotting distribution maps to visualise smaller geographic gaps in the world holding. Subsequently the German-Dutch Cooperation on Beta Genetic Resources (Iberian peninsula, East Caucasus region), the Turkish genebank (explorations within the country) and the Polish Gene Bank (landraces in Southeast Poland and adjacent areas) (DALKE et al. 2000) organised collecting trips to purposefully complete the world holding. Today, for wild beets only smaller geographic gaps need to be closed in Europe such as for B. vulgaris subsp. maritima in Northwest Spain. In countries like 173
Sugar beets and related wild species – from collecting to utilisation China, India as well as Central and West Asian countries interesting distinct landraces may still exist but these countries are difficult to access either because of their sheer size (China - leaf beet landraces) and/or political reasons (India, Pakistan, Afghanistan). The main promoters of Beta collecting missions (IPGRI, USDA/ARS, German-Dutch cooperation, Turkish genebank) began to change the objectives of collecting trips from capturing useful genetic variations to monitoring and in situ management of Beta genetic resources. There are indications that species like B. macrocarpa (Portugal and Spain), B. macrorhiza (East Caucasus), B. lomatogona and B. vulgaris subsp. (Azerbaijan, inland wild type) are threatened by genetic erosion. To prevent discontinuation of the evolutionary processes in these species, the WBN recommended assessing the threat of genetic erosion case wise and to promote and establish in situ management programmes for the extant populations in the relevant areas. In particular, wild species of the sections Nanae and Corollinae are not adapted to the climatic conditions of Northwest German lowland. They are subjected to a very strong selection pressure towards adaptation to “genebank seed production procedures”. Loss of genetic variation in ex situ collections as well as a lack of precise information on the magnitude of genetic erosion within natural growing sites underlines the urgent need for in situ monitoring projects. Maintenance Beta germplasm collections consist of species and populations with a remarkable diversity of breeding systems and survival strategies (annuality, seed dormancy, seed hardiness, frost resistance, variation for day length reaction and cold requirement). Parts of the section Beta germplasm does not at all require vernalisation and starts bolting only 40 days after sowing like B. vulgaris subsp. adanensis while accessions from Ireland bolt incompletely even after a prolonged vernalisation period. The timely induction of bolting and flowering is only one of the technical management aspects ex situ managers of Beta collections have to consider. The WBN recommended compiling a seed production manual that is available at the BAZ Gene Bank since 1996. The manual describes experiences made at the location of the BAZ Gene Bank and is valid for this location, only. Some of the experiences are summarised in Table 2. At the BAZ Gene Bank young plants are produced routinely during autumn, winter and early spring depending on the type of germplasm and vernalised artificially if required. The information given below refers to that cultivation method. 174
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L. FRESE<br />
are reasons to assume that the section Procumbentes does not at all belong to the<br />
genus Beta but to a separate genus (WILLIAMS et al. 1976, JUNG et al. 1993). Nevertheless,<br />
major users of germplasm holdings are satisfied with the taxonomic system<br />
presented in Table 1 as it transfers all the information they need.<br />
Inventory and collecting<br />
The wild species of the genus Beta are native to Europe and adjacent areas. The<br />
section Beta is mainly distributed along the shores of the Mediterranean basin and<br />
along the Atlantic coast from the Canary Islands as the most southern outpost to the<br />
South of Sweden. Section Corollinae frequently occurs in Turkey and the adjacent<br />
Caucasus region with outposts in Daghestan and the Talysch mountain (Iran/ Azerbaijan).<br />
The only species of section Nanae is endemic in Greece where it grows in<br />
mountainous areas at elevations higher that 1800 m. Section Procumbentes has its<br />
major distribution area on the Canary Islands but can also be found in Southeast<br />
Spain and along the coast of Morocco where it is rare. Even wider distribution areas<br />
have cultivated forms. Where soil, water and temperature allow sugar beet is grown<br />
in almost every country of the northern hemisphere as well as in Chile. A similar large<br />
distribution area is known for the Leaf Beet Group which is adapted to a very broad<br />
range of growing conditions in Europe, Northern Africa, the Americas and in particular<br />
countries of the Asian region.<br />
In 1979 the first Beta germplasm collecting mission was funded by IPGRI and a<br />
number of additional missions followed with the objective to sample wild beets and<br />
landraces in the Mediterranean area. Between 1980 and 1990, large geographic<br />
gaps were closed by IPGRI and USDA/ARS funded missions (South Italy, Southern<br />
part of France, West Atlantic coast, British Islands, Ireland and Denmark) (DONEY et<br />
al. 1995) and the collecting data were entered into national databases. On the initiative<br />
of the ECP/GR programme, a European inventory of Beta collections was established<br />
in 1987, and in view of the effective collaboration achieved with the NPGS<br />
(USA) it was recommended to assume an international role. The International Database<br />
for Beta (FRESE and HINTUM 1989) contains information provided by 28 germplasm<br />
holdings in 24 countries. This central crop database currently stores passport<br />
data on 2,324 sugar beet, 863 fodder beet, 892 garden beet, 631 leaf beet and 4,022<br />
wild beet accessions. The geographic coordinates of collecting sites were used for<br />
plotting distribution maps to visualise smaller geographic gaps in the world holding.<br />
Subsequently the German-Dutch Cooperation on Beta Genetic Resources (Iberian<br />
peninsula, East Caucasus region), the Turkish genebank (explorations within the<br />
country) and the Polish Gene Bank (landraces in Southeast Poland and adjacent areas)<br />
(DALKE et al. 2000) organised collecting trips to purposefully complete the world<br />
holding. Today, for wild beets only smaller geographic gaps need to be closed in<br />
Europe such as for B. vulgaris subsp. maritima in Northwest Spain. In countries like<br />
173